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SMITHSONIAN INSTITUTION 
UNITED STATES NATIONAL MUSEUM 
Bulletin 63 


A MONOGRAPHIC REVISION OF THE COLE- 

OPTERA BELONGING TO THE TENEBRI- 
ONIDE TRIBE ELEODIINI INHABITING 
THE UNITED STATES, LOWER CALI- 


FORNIA, AND ADJACENT ISLANDS 


BY 


FRANK E. BLAISDELL, SR. 


——— 


Of San Francisco, California 


WASHINGTON 
GOVERNMENT PRINTING OFFICE 
1909 


co ee 
“a Cant CLOCK BRA 


SMITHSONIAN INSTITUTION 
‘UNITED STATES NATIONAL MUSEUM 
Bulletin 63 


A MONOGRAPHIC REVISION OF THE COLE- 
OPTERA BELONGING TO THE TENEBRI- 
ONIDE TRIBE ELEODIINI INHABITING 
THE UNITED STATES, LOWER CALI- 
FORNIA, AND ADJACENT ISLANDS 


BY 


FRANK E. BLAISDELL, Sr. 


Of San Francisco, California 


WASHINGTON 
GOVERNMENT PRINTING OFFICE 
1909 


BULLETIN OF THE UNITED STATES NATIONAL MI 
IssuEp JuNE 24, 1909. - | ) 


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ADVERTISEMENT. 


The seientific publications of the National Museum consist of two 
series—the Bulletin and the Proceedings. 

The Bulletin, publication of which was begun in 1875, is a series 
of more or less extensive works intended to illustrate the collections 
of the U. S. National Museum and, with the exception noted below, 
is issued separately. These bulletins are monographic in scope 
and are devoted principally to the discussion of large zoological and 
botanical groups, faunas and floras, bibliographies of eminent natural- 
ists, reports of expeditions, etc. They are usually of octavo size, 
although a quarto form, known as the Special Bulletin, has been 
adopted in a few instances in which a larger page was deemed indis- 
pensable. 

This work forms No. 63 of the Bulletin series. 

_ Since 1902 the volumes of the series known as ‘‘Contributions from 
~ the National Herbarium,” and containing papers relating to the botan- 
ical collections of the Museum, have been published as bulletins. 

. The Proceedings, the first volume of which was issued in 1878, are 
- intended as a medium of publication of brief original papers based 
~» on the collections of the National Museum, and setting forth newly 
acquired facts in biology, anthropology, and geology derived there- 
~ from, or containing descriptions of new forms and revisions of limited 
“» groups. A volume is issued annually, or oftener, for distribution 
e, to libraries and scientific establishments, and in view of the impor- 
tance of the more prompt dissemination of new facts a limited edition 
of each paper is printed in pamphlet form in advance. 
RicHARD RATHBUN, 
Assistant Secretary, Smithsonian Institution, 
In Charge of the United States National Museum. 
Wasuineton, U.S. A., May 15, 1909. 


It 


6 My.42 g HF Wickam 


| 195286 


PREFACE, 


In 1901 when work was first seriously begun on the Eleodiini, 
a bibliography was prepared. The great conflagration in San 
Francisco that followed the earthquake of 1906 destroyed the 
California Academy of Sciences with its library, so that when the 
present monograph was about completed there were no library 
facilities at hand, nor was it possible for me to review the literature 
elsewhere.. The manuscript was therefore submitted for publication 
without a final review. Accordingly Mr. E. A. Schwarz, the eminent 
coleopterist at the U.S. National Museum, most kindly undertook 
the task for me, and to him I extend my most sincere thanks for 
the correction of the bibliography, for many suggestions, and for 
rearranging certain parts of the work. 

In the text I have applied the term ‘‘heterotype’’ to the individuals 
that form the extremes of a specific or a varietal series. The hetero- 
types therefore, represent the extreme modifications in size, form, 
seulpturing, and color. The individuals connecting the extremes of a 
series constitute the intermediates or ‘‘mesotypes.’’ Frequently 
individuals of one species are observed to simulate those of another 
species in form and sculpturing; an individual of this character has 
been called an ‘‘amphitype.” A unique, when serving as a type, is 
spoken of as a ‘‘monotype.’”’ When more than one or a series of 
individuals serve as a basis for a specific or varietal description, they 
are referred to as ‘‘polytypes;’’ if a male and female serve they are 
“*sexitypes;” otherwise they are ‘‘co-types.” 

Furthermore, in order to systematically deal with the many minor 
degrees of divergencies exhibited by individuals that are presumably 
the progeny of parents specifically or racially identical, I have deemed 
it conservative and scientific to recognize incipient races and incipient 
subraces. In accordance with that view I have used the name forma 
to make it possible to relatively define aggregations of individuals 
possessing some particular or salient characteristic; it is believed that 
these divergent characters have arisen through local or general cli- 
matic or environmental conditions acting upon the progeny of parents 
specifically or racially identical. 


NL PREFACE. 


In other words, the several formx enumerated under Fleodes den- 
tipes may arise from eggs deposited by a single typical female, under 
the influence of climatic, geographical, or environmental conditions 
capable of producing sich, divergencies. 

A Latin name has been used to express the salient characteristic of 
any given forma, and these characteristics may be similar for each 
species or variety. A species or variety may have smooth and rough, 
short and elongate, smali and large, caudate or ecaudate forms, 
consquently a repetition of Latin names is called for. 

These terms are absolutely synonymous with the specific or varietal 
terms and must not be perpetuated as distinct grades, but simply used 
as an aid in recording data, and as a compromise between unscientific 
lumping and splitting, or the difference between 47 and 400 species. 

Logically the present treatment ought not to seriously collide with 
the rules of the International Code governing nomenclature. 

Frank E. BLarspEtt, Sr. 


TABLE OF CONTENTS, kt 


Page 
REN. Ss dy en ne l 
ee NNREMRS no os PP Sa en Ses ew ar bie Poet ONS on oie ee tut ad = Wk k + 
EE Ra ae a a rs Ay eee a a te re 6 
OS Se ee 2 Pe ee nee See ney ie eee ee 8 
I Sls ee 2s Sate, Sid Ac inside mle nm sae Picye'S nine ~'4 9% coe 27 
ewmomn key to ponera of Eleodiini:.............--...-- 205-2 eee eee se eee 29 
TE DS | a es ET OMELET ES OT OS EP ee ee 31 
Analytical key to subgenera of Eleodes........................-.------- 33 
Synopsis of general subgeneric characters...................-----.------- 35 
NEE ECU HEMONCIM CR og Sean vot, SOs Sa e ok nap ie evie we oo tno 36 
noth ---Carbonaria Section. 2. 5155 5.02 Fp. be Som ote ea ae: 41 
RS 8 0 Te, ee ee See ee 41 
ee Pema Res ORMPUMI REDON gn a alg Sinn ke < 20s eS Sek ene 45 
= Eleodes.earbonaria var. soror-¥. 2.22... --- 225--250 te. sae 50 
Son RL Ne) ae ek ae a Oe a a 53 
Oo a a a a 57 
en Cee Goniee QOMOle Ia Mes a centr eh Es ee 58 
{ Eleodes obsoleta var. porcata.-.......-.-.---.-----.------ 63 
a TSS pe Oe RT eee a ee a ee = 67 
I RRs GUM Rots Se on Ria e eo gn aie mam ile dol 72 
_, Eleodes omissa var. pygmeea. ¥.-.- 77 

y, Eleodes omissa var. peninsularis™ ..............-.---------- 7 
eensa ti — Quedricoliis ceCHOn7. 22.5. oc kev sich -+ . Sees 81 
a Se RACE CIRCIIOCON IE ML 3232 Fei att ns 20 a De aE cnn 81 
3 Eleodes quadricollis var. anthracina.¥.................-.-. 87 
~ Eleodes quadricollis var. lustrans. 2v.- . 89 
Oe OT aoa et as eG 2 es oo a ee 91 
RETa ICICLE TTHOTAIIN Seas nf oe ives oe ee 95 
Sate RU MRIS EI DS. Onde Fa OE etc rote in = KG 100 
romp fT ricostata nechion. 20.55. 2.522 nn cece s+. ce-- 104 
te Pie NNR AONE I gna wis So win dig apis an cwae es 104 
eRe RINUNRNS RMANTOIENE I Pek re Ne ale ey Aw a dhe So 109 
4» Eleodes pedinoides var. neomexicana “............./....-. 113 
rE NN idee are Nes ne nica poh wits Me bam 114 
Analytical key to species of Litheleodes ...................-.-....--- 116 
Pa rreernIOO ear inthe enc St ot oo Rg oo 2. Stole eee 116 
Of OE 5 i ee ee ee ee ns eee 121 
2+ Eleodes extricata var. arizonensis....- 125 
ret ei Mi INES NRE RG eee 5 kk. cua dyipeews adds ese 127 
Z AMMO ree re waa tccies-sce---< 188 
»Eleodes letcheri var. vandykei.¥.........................-. 136 
RUNNINE. 1 PRCRONGOIIOR, Corea oe - ba ee wns cnn cw cn |) (=e ee 138 


VIE TABLE OF CONTENTS. 


Subgenus Tricheleodes—Continued. Page. 

Analytical key to BpOcins's ot. Pricheleodes< >. 2.25254. 23 Se 138 

| Bleodes hinsutac’. 2-22... 320225-¢..- s2e0see se 139 

B. Mleodes pitodae Yrs oS. 0.6 So oa as Sa eee 142 

Subgenus Pseudeleodee- 22220. 2.53 cos ve Sees ee 146 

y9. #4 Hleodes ‘prations -~ 2: 2.0. 2502. '20u ee 147 

Bitasane Promiuse <i... -.-. 0s -saee eis 22S eae oe 152 

ete key to species of Promus......... a ee 156 

me Se Bleodes insiiazgia*~ So lotey Sa ee Ne 156 

4 rT Biléodes subnitensm. 220-2 2,-5.3 oes hee a oe 161 

, \y Edeodes poryineY 2.4 028 a a Se eet, 2 an 20d 164 

4°O 4) Hleodes siriolateaM:.20502 See co ae Aes . Se 168 

2 oy Eleodes fusiformis.”: -..:..-. 32.22). 55. Pio. sees Se 172 
QS WiPleoder opataces. i266 begs. ct ees ee eee . ce) ae 

Subgenus Heteropromius: <<. 22.25 22c225.252002-26, 52-2 eee 179 

#6 $6Eleodes veterator.Y.-2...202-20222 005 22-22-22 ee 

Subpenud Wleodes. ... 202 este a eten miss ue fee eo 183 

Analytical key to groups of subgenus Eleodes Uoutooe-- pode oe 184 

Group Obscura: .. 222.5062: oes: 2 ae: 2S ee 184 

Analytical key to species of Obscura group........-. i3¢22 See 185 

2’7 9" Bleodes obpeteans.c22 235-0920. 12 een oe 187 

3 Eleodes obscura var. dispersa.“.....-...-.---..-2-----+e-- 190, 191. 

% Eleodes obscura (typical haa Set A eee 190, 193 

,o Eleodes obscura var. sylcipennis. Bp eee sere 190, 194 

26 Eleodes arata. .¥0« 4 00¢etem MU ee 194 

as | Eleodes acuta: ¥:.2.422-.0.5. 4 224-2 ee 195 

R Fleodes:suturalis. ~~. 2590.02 els 2-3 oe ne eee 199 

Eleodes suturalis (typical variety) “2... 2.2 2-2 See 202 

Eleodes suturalis var. texana..27-.--.-.-+-..2.. 0... aso 202 

Group Grandicollig. ..32 22222. wa ae oe oe 205 

3) 6% Eleodes prandicollis..“-.: 22-2055. . 2.305.204. 0-2) 

Group Dentipes, section Az: 215254052500 242-) 75a de 211 

Analytical key to species of Dentipes group. Seki es 212 

p) leodes sponse. i.e 0. 2o3. ase ee eee ee 213 

® Hleodes hispilabris-©. 05 ..¢.2205 5 oS Re ee 217 

epHleodes caudiiera 10-2 055... Bis2 oi in 5s. oe 225 

Wf Eleodes longipiloss.¥-.-<... 222.7222. -2. OS 2 a 230 

Group Dentipes, section B.2.:. 2-222... 203 2222: == nso 234 

Analytical key to Bpoces of Dentipes group. 82¢¢.8:............ 234 

¢ We Rleodes pracilise 2 el eS eee 237 

wy Eleodes gracilis var. distans-7.. 2.20 2. 22. ee a ee 242 

3) $e Eleodes subcylindrica™:. .-22. 00.42.2435 .2 1. 246 

af. @9 Eleodes subpimguis-<-5 = 4.4. 2s 3 2 <. 247 

3 s Eleodes ears ee ES PE PT REE EE ETE 251 

+6 ff Mileoden Orinbiae occ 2. secs ei on ee ct ow oes 259 

» Bleodes‘armata var. Impotens.<: :. .:-.<.- --2 62.5.2 ceeeeee 263 

¥/. 7 Fleodes moibttarigl 2. o5 2... so ee ee oe 267 

9 Eleodes militaris var. femorata™..-....-..... 2. .2cne ee 272 

vt é Eleodes acwticauda. f0 2... ac. gl. 22 ee 

« Eleodes acuticauda var. laticollis.¥ oY... de onc ee 280 

vs <7 tuleodes cachacholtel i... -....--.-...scen ee eee ee 285 

£3 Eleodes eschscholtzi var. lucee.“......... 2.2... .en<+-s--< 4s 289 

gu -¢¥ Bileodes tenuiped- <9. i522... ae. ae as oe see 294 


up 4 Eleodes wigkhatil. 0.2 2.ict gken ~ sos kee. eee ae ee 297 


TABLE OF CONTENTS. IX 


Group Dentipes, section B—Continued. Page. 
eter te WOTTON Moe Soren ee wwe ns eC uvevcleveeeces 300 
67 Bleodes ventricosa var. falli.¥-...... 2.225006. 050 0. eee e ce eo 305 
Subgenus Blapylis.............. pe SN CE ee ee 306 
Synopers Of genital characters... .... wii es. 0 deg... 1... . 2. ee 310 
Analytical key to species of subgenus Blapylis....................... 311 
eA neta CAIGEM, Ce ci, oa Rae eat Ca ci toldale des -scnsece 313 
ET aernee MOMMA ocd, yc Sone oe ke ds US ome ne ee een 317 
ef 7a Wleodes lecontei.v...........- 2-520 cete eee eee eee eee ig 321 
aay rumah LenenmeniaN eset Lo. Sr Rs oes ee oe avert Soe 
»-Eleodes tenebrosa var. nana. .......... 22.22 cece eee ea wees 328 
ene Oem PURER, Fos 5c eh oe Saas opt bel ok das oe 329 
sv 4 Eleodes consobrina.%............ i habe bead Sete nde tics 3 = So > 0 333 
emneay SP OCRCD BCADTADOMBIE Wo 8 aac oo. ois slag eee as 338 
nee ever Dlanchardncs ye. 52-6 SUN Sede. ia eu ces = oo SV 339 
an ee een CUEIT ST cco saves oI Se paws Sifive Seawehh cece ed see 343 
y% 7¢ Hleodes neotome.¥.............. Seba lal he pe OER A 347 
CPE EN EMR ni a ala ih iu Soden sink 8 we Pea weiss pays SS 350 
SEP EC RUA VAEVIGOUIS Mo, ano ota se Peds Welt 2k) Sree. tes 354 
g( Eleodes parvicollis var. planata.“~......... ee a LE: 359 


&@. Eleodes parvicollis var. producta’.........:................ 362 
$3 Eleodes parvicollis var. constricta.v......................... 365 
i nies nliiod MODDING. Ne ee io ceed Snes Kaw satan an rewe. — BOB 


Semen ert CIAWICOMAIA, YO. 0 coo oe oh is wa une Ae be bee fees ou 7 
Pee eevee GCARRORR. Mess 1.2. lease eel. SLL ee aS 375 
aE BeOS. COLCA ce S58 a tiae anomie Leet Ri eset oes. 2 ae - 379 
¢f Eleodes cordata var. rotundipennis’....................... 384 
43 #4 Eleodes pimelioides..'............ EAR Bes wee Pelt ae Seo 384 
Yo Eleodes pimelioides var. brunnipes.¥.....................-- 386 
Se URE AMO ES Wee Sask ee pe nk Sk ld ew page 388 
0 nd SSUES See ee a ae eae er. ha 391 
Analytical key to species of subgenus Metablapylis.................. 393 
Penrmemnemnr rs wimeritia. Sno. he Sue. a ee ge one 393 
92 Eleodes Miptrasa at. “Pemonee neo vole oc aes ee eee cs Sa 398 
Be Satender dieing. ¥~.- tones. ole. Lo aries oo eS See 398 
9;-Eleodes dissimilis var. nevadensis.“.................-----. 402 
42 7(Eleodes schwarzii.¥...................- cS eee Sawa 406 
ePEHILY CHMMNOOGOD aya ain cease As rake oban sw edc esa ak db fed ow 2 409 
Analytical key to species of Steneleodes......................-.-.-. 411 
CN VE EE ee ee ae ane 412 
; gf Eleodes gigantea var. gentilis.v_....-..-................22-- 416 
q Eleodes gigantea var. estriata.Y.................---222---.--- 421 
a pOOGOd IORPICOMIB. Wan sees tpt <cwaaer<ssy sas... 425 
ee i TR Cy eee ea ee 429 
SERIE STOUT SS 33 cB el ie RY Sodan a ol tbc ae. 434 
Analytical key to groups of Discogenia....................-.!...... 434 
Analytical key to species of Discogenia, Group A................ Z2gee, -AS6 
Analytical key to species of Discogenia, Group B........--. Pace: 435 
Fl: Pe ee MINI cs 2 on dias = in a pe ee ole aku s coe OSes css 436 
es Oh ON NEE eo Fe a non ki ack od odd any ao 440 
B/E DCGUEE SHMUMOIINN nc coca nel neneweteen cast ocns,css--. 445 
en eee) eh a ee a 450 
ROO A Me Bn Sta nb een ois ose nS un ---sn-e 450 
PMID TMM PUR ON an er clin Nae eoecs oie ss scsucee 450 


x TABLE OF CONTENTS. 


Unrecognized species—Continued. Page. 
Genus Embaphion. 22: 222265 dacs owes cok d gee ane ea oe 450 
Analytical key to species of genus Embaphion................™-......-.- 453 

‘ Embaphion depressum. - .2...2-2 20-5... 0.2. oe 453 

+ Embaphionelonsatim =). 2.: 9.2: ba eer 454 

3 Embaphion' glabrum. :..c-.4.2--22- 225-3 ee eee 457 

% Embaphign contractum-.....-: 2.22. .2s-8 <a eee 460 

~ Ean baphion planunt 2225 3-.... les. 3. Sch Bie dkes eee ce 464 

. Embaphion contusum..:...2....2 4.42402.) J5ee ee 467 

? Embaphion contusum var. laminatum..............-------- 472 

? Exsbaphion muricatumc: ..: -:-2_25.< 4.e- eng 2) 2 473 

Gents HileodmMorper. . 2-262. gap 2 cp oes Be ie do leg a 477 

. Bleodimorpha bolean...... cs awa, ec 2 fae 479 

Genus Iropioderta. ct 22s aon 22k Ss. sa eee ee ee ee 483 
Analytical key to species of genus Trogloderus..............-..-.--------- 486 
/Trogloderus’costatus=se-. < 25222 ae S52 ee ee 486 

s Trocloderus tuberculatus:: < 2.5.0. 2-50 sees 5-8 ee | ee 490 

Brecies hokrecamniged ol 2.1.56 2 Se sewn ew so os we a 493 
Bleoger wititing: > 4 iccate so. ee ee bee oe ios <9. 493 

Facodes nitidus. .2 24 2.2 2'-. 52. 20s 28% ... 1 eee 493 

Bleodes, reflexicolliss: 224.225 ks. i225. oo Se 494 

arly stapes of the Mleodiini-- 3... 320200 A} see ses bt ee 495 
IRAE IADS: 22 22a th mn ete enews estes fase ieee oo ee 502 
Appendix (Qiwaternary species)’ .+.. - 22 0. 2. + en benoe~ gee 505 
Eleodes:elongata.-... 523-55. -¢.2\umoence foe eee 506 

Hleodes; bebrit. 2s. 2... isa26) Ss Je 506 

Hisodes intermedia $71.28. 23352. 8he52 oud’. a 2 507 

Beplanation of plates... 2-252 -202 -22sqien- - tet ete ss ee 509 


Pex)... 2 Ss ys Bt tae RO See As ee 521 


LIST OF ILLUSTRATIONS. 


TEXT FIGURES. 


Genealogical diagram of the subgenus Melaneleodes.............-....----.---- 
Genealogical diagram of the subgenus Promus...............--..------------ 
Genealogical diagram of the Obscura group, subgenus Eleodes. (ae ee, See 
Genealogical diagram of the Dentipes group, section A, subgenus Eleodes.... - 
Genealogical diagram of the Dentipes group, section B, subgenus Eleodes.. - - 
Genealogical diagram of the subgenus Blapylis...........-.-.-...----------- 
Genealogical diagram of the subgenus Steneleodes.................----------- 
Genealogical diagram of the genus Embaphion...............--..-..-.----- 


PLATES. 


Page. 
39 
156 
186 
212 
236 
309 
411 
452 


Facing page. 


mmnnetarcharactersof the Mleodiini=.-- 22. .-).222. - 2222s ees ee tb oe 
marnitan characters of the Mleodini..:-... 2.55... 22. 220 J. i Sees ne 
Bua Guarichers. of the mlecdtini_-... - 55 22 oe 
wererta) characters of the. Mleodiini. 2. -.... 2.2.22 - sto. sce. oe eee eee 
Pmimuanaracters Of the MiCOGMINI.. --.. 22s... je 22 ndbue sc cae sen ccce de eee 
Comparative and genital characters of the Eleodiini and related tribes... -- 
. Topography of the genital segments of the Eleodiini....................- 
RIM SE AONSTRINS 8 OP Ro on cae Sawn ones ee 
. Homology of the genital segments of the Eleodiini....................--. 
SERA PHOOGLENT. 22 |e ee Ln. tee ern ae eo ee 
DMR UNCP COUINS. (2 Pte Me ek eS ek occt dee 
SRIOELE TE DOG UIT y 25 Yas ees 8 er sk ee ES ee es 
EE ET) ee 


XI 


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SODBNAMR WH 


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524 


A MONOGRAPHIC REVISION OF THE COLEOPTERA 
BELONGING TO THE TENEBRIONIDE TRIBE 
ELEODIINT TNHABITING THE UNITED 
STATES, LOWER CALIFORNTA, AND 
ADJACENT ISLANDS. 


By Frank E, Buatspevy, Sr., 


Of San Francisco, California. 


INTRODUCTION. 


Since the publication of the Revision of the Tenebrionide * no 
attempt has been made to treat the Eleodiini in a revisional way. 
Material has been constantly accumulating in collections, until it has 
become a very perplexing problem to refer it in all instances to species 
already named, or to correlate the numerous variations so evident in 
many species. 

At first I had intended to simply deal with the species of the 
genus Hleodes, but little by little I was induced to include all of the 
genera constituting the tribe as limited by previous authorities. 

My method has been radically different from any heretofore pur- 
sued in the United States and it has been a laborious one. In the 
following pages I have set forth the results of my investigations 
‘and deductions which have been carried on during past five years. 
About one thousand specimens have been dissected during this time 
and over five thousand studied superficially. 

I became convinced from the results previously arrived at by other 
writers on the family that all methods resorted to had been inade- 
quate for the proper understanding of species that are so polymor- 
phous and present such a community of habitus. 

I decided to carry on a study along the lines of primary sexual 
characters, and if possible to investigate the mouth-parts and then 
to correlate the data obtained with characters already known. 


@Amer. Phil. Soc., XIV. 
597T80—Bull. 68—09——1 


Z BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


The first part of my task was perfectly feasible; but not so with 
the mouth-parts, on account of the necessity of m#ttilating the speci- 
mens. 

The first species were named by Thomas Say in the year 1823. He 
placed them in the genus Blaps. 

Eschscholtz a few years later® erected the genus /Jeodes, deserib- 
ing several species. 

Mannerheim still later” described a few species, redescribing some 
of Eschscholtz’s species with references to the original descriptions. 

Then followed the period between the years 1858 and 1865, the 
most active in the making known of new species, during which Dr. 
John L. LeConte described more species than any previous author. 
Later a number of his species were suppressed by Dr. George Horn, 
who from 1874 to 1891 occasionally described new species. 

Many of the species suppressed by Doctor Horn have been restored 
to specific standing in the present paper. Doctor Horn was too 
conservative and his knowledge of the species was based upon char- 
acters that were too superficial. 

Thomas L. Casey described Eleodes arcuatus in 1884; nine others 
in 1890,° and three more in the sixth volume of the annals of the 
same society. 

In 1902 I described ELleodes interrupta from an anomalous speci- 
men of /. omissa; in 1895 EF. impotens, a race of armata, and £. 
confinis, the latter no doubt being an incipient race of 2. dentipes. 

LeConte’s £. seriata proved to be identical with Goryi, described 
by Solier from Mexico. 

Lacordaire 4 explains the generic term Yysta as follows: 

Le menton * * * est composé de trois parties; une mediane de forme 
variable; et deux latérales en forme d’ailes, placées sur un plan plus interne, 
tranchantes latéralement, et d’autant plus larges que la premiére est moins 
développée. C’est sur la forme de cet organe qui Eschscholtz s’était basé pour 
diviser le genre en deux. I] donnait le nem de Yysta aux espéces qui ont la 
partie médiane trés-développée et arrondie en avant, et réservait celui d’Bleodes 
A celles of. son bord antérieur est rétréci et plus ou moins triangulaire, mais il y 
a entre ces deux formes des passages qui leur enlévent Ja faible valeur qu’elles 
pourraient avoir. 

In a footnote he says: 

Eschscholtz en indique quatre espéces [gravida, angulata, rotundicollis, sul- 
cata] [belonging to Yysta] du Mexique, mais il y en a beaucoup d’autres, la 
tricostata par exemple, * * * Chez la plupart de ces espéces, les ailes lat- 
érales du menton étant petites, paraissent au premier coup-d’eil, ne plus exister, 
mais, avec un peu d’attention on les découvre sans peine. 


@ Zool. Atlas, 1829-1833, I-V. 
> Bull. Moscow, 1843. 

¢ Annals N. Y. Acad. Sci., V. 
4 Genera, p. 149. 


REVISION OF ELEODITNI—BLAISDELL. 3 


Kirby ¢ describes Pimelia alternata, which is now a synonym of 
Eleodes tricostata Say. After the description he says: “ Mr. Say 
says of his Pimelia (Edrotes) rotunda that it was the first of that 
genus found on the New Continent; that above described (a/ternata) 
furnishes therefore a valuable addition to the American insect fauna.” 

A brief reference to European literature shows that formerly the 
name Pimelia had a wide application, sufficient to include even such 
remotely related forms as /'drotes and Lleodes. 

Pimelia is a Fabrician genus now including a considerable number 
of European species about the Mediterranean, also Northern Afric: 
and Western Asia. 

It may be interesting to the student to compare Eschscholtz’s ? 
synoptic characters of the four genera, Blaps, L'leodes, Nystus, and 
Nycterinus, which are as follows: 

Blaps.—Tibiwe antic apice bispinosee. Antenne articulo tertio sequenti 
duplo longiore, ultimo acuminato, tribus anticedentibus subglobosis. Labium 
apice latius, planum. Elytra carina laterali. Mas segmento basali abdominis 
scopifero. 

BHleodes.—Antennze articulo ultimo transverso, duobus antecedentibus, rotun- 
dis compressis, tertio sequenti duplo longiore. Labium apice angustius triangu- 
lare. Clypeus indistinctus, antice leviter emarginatus. 

XYystus.—Antenne articulo ultimo transverso, duobus antecedentibus, rotundis 
compressis, tertio sequenti duplo longiore. Labium transversum, medio dila- 
tatum, apice rotundatum. Clypeus indistincetus, emarginatus. 

Nycterinus.—Antenne articulo ultimo transverso, tribus antecedentibus com- 
pressis, tertio sequenti parum longiore. Labium apice latius. 


In the synopsis 7’agona precedes Blaps, and Prosodes between 
Blaps and Eleodes, Nycterinus between Vystus and Amphidora. The 
characters used are very variable. I have not had an opportunity to 
study Vycterinus. I have a species from Northern Chile that I 
believe belongs to, or quite close to, this genus. 

Gemminger and Harold ° give two species of Zagona as inhabiting 
Russia Meridionale and Turcomania; Prosodes, Oriental Europe and 
Western Asia; Vycterinus, twelve species, mostly Chilean, and Blaps 
as very numerous in Europe, Asia, and Africa. 

I hope this work may be the forerunner of others upon the Tene- 
brionide, and have therefore taken the opportunity to place before the 
American student many suggestions that would be out of place in an 
ordinary revision. I sincerely trust that my colleagues will continue 
to aid me in the future as they have done in the past. 


*Fanua Boreali-Americana, IV, 1837, p. 232. 
» Zool. Atlas, III, 1829. 
¢Catalogus Col., VII. 


4 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


ACKNOWLEDGMENTS. 


It is a great pleasure to acknowledge the generous loan and gift of 
material, as follows: 

The United States National Museum in Washington made the loan 
of 1,914 specimens, which were transmitted to me through the kind 
intercession of Dr. L. O. Howard. This material was of invaluable 
aid and of great interest, as it contained material from the collections 
of the following weil-known entomologists: H. G. Hubbard, E. A. 
Schwarz, C. V. Riley, H. Soltau, E. J. Oslar, D. W. Coquillett, H. K- 
Morrison, M. L. Linell, J. B. Smith, J. D. Mitchell, H. S. Barber, 
C. F. Baker, and A. 8S. Fuller. 

Mr. Charles Fuchs and Dr. E. C. Van Dyke put all of their material 
at my disposal. 

T am especially indebted to my friend, Henry C. Fall, for the com- 
parison of material with authentic types in the collection of the Mu- 
seum of Comparative Zoology in Cambridge, Massachusetts, and for 
the loan of species not well represented from other sources. 

To Mr. Frederick Blanchard, of Tyngsboro, Massachusetts, I 
extend my sincere thanks for his painstaking care in the comparison 
of many consignments of specimens from my collection with the Le- 
Conte types in the collection of the Museum of Comparative Zoology 
in Cambridge. His valuable notes and short descriptions aided me 
greatly, and without which I could not have written so authorita- 
tively. At his request Mr. Samuel Henshaw, curator at the museum 
in Cambridge, very kindly transmitted to me one of LeConte’s speci- 
mens of Hleodes gentilis. 

At my request Mr. Charles Liebeck, of Philadelphia, looked up the 
literature on Blaps and made notes on specimens in the collection of 
the Academy of Science in that city. 

Prof. H. F. Wickham, at the sacrifice of his own limited time, most 
kindly looked up literature upon the tribe now under consideration, 
and those references that could not be obtained upon this coast he 
copied and forwarded to me, namely, the original descriptions of 
Mannerheim. I must also thank him for the loan of material, espe- 
cially for specimens taken in Texas, Nevada, and New Mexico. 

IT am indebted to Mr. Ralph Hopping for the examination of the 
material in his collection, especially that from Tulare County, Cali- 
fornia. I had the privilege of studying a collection taken at Fort 
Tejon by Messrs. Hopping and Fuchs. 

Prof. J. J. Rivers presented me with a series of EJeodes taken at 
Ocean eee Los Angeles County. 

Dr. C. F. Clark most kindly contributed a collection of Eleodes 
taken in ae County, California. 


REVISION OF ELEODITINI—BLAISDELL. 5 


Mr. Warren Knaus very kindly aided me by the loan and gift of 
most interesting material collected in Kansas, New Mexico, and 
Colorado. 

The late Beverly Letcher permitted me to study a series of speci- 
mens that had been collected along the Colorado River in Arizona. 

Lieut. Col. Thos. L. Casey donated several of his species and 
other specimens. LVeodes rile yi was of particular interest. 

The California Academy of Sciences, of San Francisco, placed its 
material at my service. The collection was of more than or dinary 
interest on account of material from Lower California. 

Mr. Fordyce Grinnell gave me a series of specimens taken in Kern 
County, California. 

I received from Mr. Edward Ehrhorn a small collection from the 
mountains at Monte Bello, near Mountain View, Santa Clara County, 
California. 

My friend and colleague, Dr. William Lee Moore, and his son 
Edgar, collected a large series at Verdi, Nevada, and sent the same 
to me. 

My mother, Mrs. Anna G. Blaisdell, gathered an immense series 
at San Diego, California, and transmitted most of them alive for 
special studies. 

Mr. Albert Koebele very generously gave me a series of specimens 
taken about Nogales, Arizona, and also permitted me to study those 
in his collection. 

Mr. F. W. Nunenmacher also supplied me bountifully with speci- 
mens from about Nogales. 

Dr. A. Fényes very kindly sent me material for examination and 
study. ; 

Prof. F. H. Snow, of the University of Kansas, permitted me 
to study and identify numerous specimens collected by himself in 
Arizona and elsewhere. 

Dr. R. H. Wolcott, of the University of Nebraska, transmitted to 
me the entire collection of /7eodes of that institution for study, and 
also a collection of Mexican species. 

I most sincerely thank Mr. Charles O. Waterhouse, of the British 
Museum, Cromwell Road, London, for a series of Blaps, which he 
very kindly loaned in response to my request. 

Miss Julia Wright, of Palo Alto, California, allowed me to study 
the native species in her collection, also a collection of Blaps obtained 
by her while abroad. 

I also looked over the collection in the entomological depart- 
ment of Stanford Univ ersity, through the courtesy of Prof. Vernon 
Kellogg. 

It is my duty and pleasure to thank all who have so promptly 
aided me, and especially my intimate associates, Dr. Edwin C. Van 


6 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Dyke, Charles Fuchs, and the late Beverly Letcher for their en- 
couragement and advice. Without this material 4nd moral aid the 
present work could not possibly have been as complete as it is. 


METHODS OF STUDY. 


All comparative measurements have been made from outlines 
drawn by aid of a binocular microscope with either a 4, 1, 2, or 3 
inch objective and a camera lucida. The legs were measured directly 
with a finely divided metric ruler, under a hand lense. 

Fresh specimens were hardened in alcohol or a two per cent solu- 
tion of formalin; dried specimens were quickly relaxed in hot water. 

To determine the relative length of the antenne—the parts being 
relaxed—the head is either retracted or extended so that the apical 
margin of the pronotum corresponds to the posterior margin of the 
tempora. (See line C, Plate 8, fig. 1.) The antenna is then 
thrown back over the dise of the pronotum so that it falls a little 
within the basal angles. If it does not reach beyond the apical three- 
fourths of the pronotal disc, it is said to be short; if it reaches to the 
basal margin it is moderate in length; if beyond the basal margin it 
is Jong. Qualifying terms are used to express nearness to these 
points. 

This method is arbitrary, but sufficiently correct for all practical 
purposes, if care be taken to have the head in the proper position. 

In determining the relative width and length of the head a tracing 
is made, care being taken to have the surface plane parallel to the 
focal plane of the microscope. The drawing is then treated as in 
fig. 1, Plate 8. A line is drawn transversely, just touching the pos- 
terior margins of the eyes and called the post-marginal ocular line, 
fig. 1b, Plate 8. A second line is drawn from a point on the post- 
marginal line midway between the eyes to the middle of the anterior 
margin of the epistoma, and termed the interlocular line (fig. la, 
Plate 8). Line } practically divides the head into equal halves, 
near enough for our purpose. If the distance across the widest or 
most prominent part of the eyes is twice the length of line a, the head 
is said to be moderate in width; if less, the head is Jong, and if 
greater it is wide or broad. The proper qualifying terms being used 
to express nearness to or extremes of these relative dimensions. For 
example, the head may be said to be longer than wide, somewhat 
longer, or distinctly longer than wide. The head is subquadrate, 
and if line a be one-half the total length, and if the width be twice 
line a, the head is as wide as long. The labrum can not be included 
in the measurement as it is not fixed. 

For the removal of the genital segments the parts must be soft 
and flexible. In cabinet specimens hot water swells the soft parts, 


REVISION OF ELEODITNI—BLAISDELL. 7 


so that they approximate their fresh condition. The specimen, when 
ready, is grasped between the thumb and index finger of the left 
hand, dorsum to the palm, then with a lance-pointed dissecting 
needle depress the last two ventral segments, pass the needle internal 
to the sternite of the seventh segment and forward along the inner 
surface of the sixth and fifth abdominal segments, cutting first to the 
right, then to the left, detaching the integumentary attachment along 
the pleural region; when this is done, with a curved or hooked pointed 
needle pull out the viscera. Place them on the stage of a dissecting 
microscope and carefully remove the ensheathing sclerites and mem- 
branes from the genital organs; the latter should now be washed in 
60 per cent alcohol, then placed into another bath of 60 per cent 
alcohol to which a little glycerine has been added to prevent too 
rapid drying while being studied. Remove and wipe with a camel’s 
hair brush, and mount on a white bristol-board slide for the micro- 
scope, a minute drop of glue being used for the purpose. 

Study the dorsal surface first and be sure to orient the specimen 
so that the surface is parallel to the focal plane; when finished wet 
slightly and remove from the slide, wash in water to remove the glue, 
throw into the alcohol and glycerine, remove and wipe and mount as 
before, but in-a reversed position, and study the ventro-lateral sur- 
faces. When this is done remove as before, wash in alcohol, and 
mount on a small piece of cardboard and place on the pin beneath 
the insect for future study. 

Parts to observe about the female organs: 

Dorsal surface—General form, pubescence, relation of the valves 
to each other, apices, cercopoda, dorsal plates, and superior pudendal 
membrane. 

Ventrolateral surfaces—General surface contour, pubescence, basal 
prominences, submarginal grooves, position of the fosse and cer- 
copoda, genital fissure, and inferior pudendal membrane. 

Parts to observe about the male organs: 

Dorsal surface —General surface contour and form of the edeago- 
phore and its basale and apicale, presence or absence of a membranous 
area or groove. 

Ventral surface—General form and condition of the membrane, 
ale; clave, form and relation to the edeagus; degree of inflexion of 
the sides of the basale. In the male the sternite of the seventh seg- 
ment must be removed separately and studied in conjunction with 
the edeagophore. It is of value in classification. Prepare and 
mount as the other organs. 

In order to study the relation of the sclerites of the retractile seg- 
ments and reproductive organs, clear with liquor potassi. 

The study of the tarsi should be most carefully done. I have used 
a bull’s-eye condenser for concentrating the light, and while holding 


8 BULLETIN 63, UNITED STATES NATIONAL MUSEUM, 


the specimen in the light in the most convenient position I have 
examined the tarsi with a hand lens, for the small"species using the 
compound microscope. The first three joints of the anterior and 
those of the middle tarsi should be most carefully examined under 
different angles of illumination. Many specimens have the hairs or 
spinues rubbed off; if possible use perfect specimens. 

A study of the mouth-parts, while exceedingly desirable for the 
purpose of correlation with the genital parts, is not possible without 
dissection and mutilation of the specimens. In many _ instances, 
nevertheless, this has been done, especially when the material war- 
ranted it. 

Caution.—The student must bear in mind not to accept measure- 
ments representing length, breadth, and thickness in a literal sense, 
but only as indicative of relative proportions. 


GENERAL EXTERNAL ANATOMY. 


The following results have been obtained chiefly from a study of 
Eleodes dentipes. 

Form variable. Integuments usually strongly chitinized. 

Head (Plate 8, fig. 1) quadrate, nearly encircled by a rather broad 
impression, which is strongly marked behind the gular peduncle 
and buccal fissures, less so behind the tempora, sometimes feebly 
indicated across the dorsum. By this impression the epicranial wall 
is divided into two regions, the occipital and fronto-buccal. The 
epicranial sutures are mostly obsolete. 

The occipital region is globular and condylar, and the surface more 
or less finely granulato-punctate; the foramen usually vertically 
oval, superior margin bisinuate, sinuations separated by a median 
subtriangular process, inferior margin broadly sinuate and straight 
between the gular sutures. The superior and visible portion of this 
region forms the vertex. 

Fronto-buccal region moderately depressed, surface lines converg- 
ing to meet about a length of the region in advance. Frons more or ~ 
less transverse, never strongly convex; limited laterally by the eyes 
and the more or less prominent and arcuate preocular frontal ridges, 
which thicken and descend in front of the eyes to their inferior 
fourth, articulating with the gene below. The inflexed and externo- 
anterior surface of the frontal ridges are excavated, forming the 
antennal fosse, the anterior margins of which bear a narrow oblong- 
oval mandibular condyle, just behind the lateral frontal suture. The 
anterior frontal margin is deeply emarginate, middle three-fifths of 
the emargination straight, sides prominent anteriorly forming the 
frontal angles, the internal margins of which are straight and an- 
teriorly divergent, forming an obtuse angle with the middle of the 


REVISION OF ELEODITNI—BLAISDELL. 9 


emargination, the latter receiving the base and obliquely truncate 
basal angles of the epistoma, forming the more or less evident frontal 
suture, the latter sometimes apparently sinuate at middle. 

Epistoma transverse, surface more or less plane to feebly convex; 
base embraced by the anterior margin of the frons (see Plate 8, fig. 
1); sides straight and convergent anteriorly, continuing the sides 
of the frons and forming therewith the fronto-epistomal border; 
anterior margin more or less sinuate, angles rather narrowly rounded. 

Tempora small, and not in the least prominent, sloping from the 
posterior margins of the eyes, and feebly convex, passing evenly into 
the frons above and the gene beneath and limited posteriorly by the 
transverse impression. 

Gene not prominent laterally, surfaces feebly convex, limited inter- 
nally by the gular sutures. Each anterior margin is irregular, pre- 
senting for examination two angles—a superior or frontal, and an 
inferior or mental; three emarginations—a superior, mandibular, 
and buccal; two processes—the mandibular and buccal. (Plate 8, 
fig. 15.) 

The superior or frontal angle joins the inferior margin~ of the 
antennal fossa in front of the inferior fourth of the eye, and separated 
from the mandibular process which lies below it by a very short and 
somewhat oblique, shallow apical or superior emargination, the 
ventral margin of which is more or less beveled; the mandibular 
process is short and obtuse, and when the superior emargination is 
obsolete, scarcely recognizable as a distinct part near the superior 
angle; the mandibular emargination is a small but well marked 
reéntrant margin between the mandibular process above and buceal 
process below, its curvature is somewhat oblique superiorly where it 
forms the ventral edge of the mandibular process; the buccal process 
is anteriorly prominent, subacute, but not produced, forming the 
abrupt and external boundary of the deep and evenly rounded buccal 
emargination, which terminates internally at the mental angle of the 
gular peduncle. That part of the anterior margin bounded above 
by the apical angle and the buceal process below is the mandibular 
portion, while that part between the buccal process above or exter- 
nally and the mental angle of the gular peduncle ventrally and 
internally is the buccal portion. The second joint of the maxillary 
palpus rests in the mandibular emargination when extended against 
the side of the head. The mandibular portion is opposed to the base 
of the mandible, and the buccal portion at the internal margin of the 
emargination gives attachment to the cardo of the maxilla and forms 
the posterior boundary of the buccal fissure. 

The buccal process on its internal surface has a small glenoid 
eavity by which it articulates with the small globular and somewhat 
prominent condyle of the mandible. 


10 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


The buccal process is moderately produced in /Lleodes blanchardi 
and strongly so in Eleodimorpha bolcan. (See Plate 8, fig. 24.) 

Eyes narrowly subreniform, transverse, and slightly oblique, su- 
perior margins rounded, the inferior slightly subtruncate against the 
gene ventro-anteriorly, rounded somewhat behind this point and 
continuously so with the arcuate posterior margin; the anterior 
margin is broadly and evenly emarginate, adapted to the deflexed 
frontal ridge, and briefly so to the superior genial angle. Surface 
finely faceted and not setigerous. 

Gular region, besides being punctato-granulate, is finely and ir- 
regularly or transversely regulose. The gular sutures are widely 
separated inferiorly at the sides of the occipital foramen, and some- 
what arcuately converging anteriorly to become nearly contiguous at 
a point, whose distance behind the base of the mentum is equal to the 
width of the gular peduncle, and continuing forward for a short 
distance to become divergent, passing to the mental angles of the 
peduncle, bounding laterally the small triangular submentum, whose 
base is usually a little longer than one of its sides. 

The transverse cephalic groove is strongly impressed across the 
apex of the submentum, becoming less so outwardly just behind the 
posterior boundary of the buccal fissures. 

Gular peduncle short and slightly deflexed, formed by the sub- 
mentum, whose base forms a free margin, to which is attached the 
mentum. 

Gula.—Between the gular sutures posteriorly is the subtriangular 
gula, whose base forms the straight inferior boundary of the occipi- 
tal foramen. 

Labrum more or less transverse, sinuate at middle, sinus rounded, 
lobes rounded and continuously so with the sides; surface feebly 
convex, punctate, sete longer latero-anteriorly, and decurved; the 
anterior margin each side of the central sinuation, just beneath 
clothed with dense transverse tufts of golden pubescence (/abral 
tufts), which are directed downward, inward, and slightly forward. 

Mentum variable, internally giving attachment to the labium; 
apex of middle lobe free. (See descriptions of species. ) 

Antenne variable in length. The outer three or four joints are 
always more or less feebly compressed and slightly dilated; second 
joint always short, frequently wider than long, and subannular; the 
third always long, from three to five times as long as the second and 
usually about as long as the fourth and fifth taken together; the 
fourth and fifth may be equal in length, or the former a little the 
longer; fifth, sixth, and seventh usually about equal in length and size 
and obconical in form: eighth more or less triangular; eighth, ninth 
and tenth usually more or less transverse; eleventh oval and ob- 
liquely truncate at tip. The first joint is moderately arcuate, and 


REVISION OF ELEODITNI—BLAISDELL. 11 


distinctly constricted beyond the condyle. Joints eight to eleven, 
inclusive, more or less densely and finely pubescent, with tactile 
sete. All the joints are irregularly punctate, and from each pune- 
ture arises a seta; sete longer on the apical moiety of each. 

Right mandible (Eleodes dentipes, Plate 8, fig. 12) short, robust, 
triangular, a little decurved at tip, the latter feebly emarginate. 

Superior surface feebly convex, divided by a feeble arcuate ridge 
into an apical or incisive surface occupying about the anterior third, 
and a basal or molar surface occupying the basal two-thirds. In- 
cisive surface with a series of short parallel ridges near and at right 
angles to the internal margin. 

External surface with outline quite strongly arcuate, most so in 
apical half, which becomes apparently transverse; divided by a 
longitudinal ridge-like prominence into two parts, a superior and 
an inferior, both feebly concave and finely seabro-punctate. Sur- 
face at apex not concave. 

Ventral surface (Plate 8, fig. 10) deeply and arcuately excavated; 
lacinia prominent, rounded and free anteriorly, finely pubescent. 

Internal surface (Plate 8, fig. 2) deeply concave within the incisive 
third; lacinia prominent before the well-developed and transversely 
oval molar, behind which is a membranous lobe: the latter is pubes- 
cent, and the hairs longer than on the lacinia. Molar with a fine 
elevated margin, the inclosed surface area feebly and irregularly 
excavated. 

Left mandible (. dentipes) more strongly decurved apically and 
more deeply excavated within; incisor margin slightly emarginate; 
external surface at tip more strongly beveled from the emarginate 
edge. As a whole it is less strongly arcuate and a little longer than 
the right. 

Mawilla (EF. dentipes, Plate 8, fig. 11).—The palpi are compara- 
tively large and four-jointed. The basal joint small, quite strongly 
and outwardly bent at middle; second joint elongate, obconical and 
nearly twice as long as the third, the latter also obconical and slightly 
arcuate at base; fourth broadly triangular, not equilateral, the inner 
margin the shortest, the external and apical subequal. 

Lacinia ending in a chitinous claw-like point; external surface con- 
eave with margins arcuately dilated at middle, adapted to the in- 
ternal surface of the galea. The internal surface just basad to the 
claw-like extremity is densely set with rather long incurving cilia- 
like sete, elsewhere the surface is shining and very glabrous. 

Galea of one joint, subtriangular (variable in the different species) : 
external surface evenly convex and glabrous, limited apically by a 
dense and evenly transverse row of rather short spinules, immediately 
beyond which the apex is densely set with rather long, incurving 
spinule-like cilia. 


12 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Cardo subparabolic in outline. Conical eminence at base of palpus 
quite evident. Palpus and palpiger punctate and ®etose; cardo more 
finely so. 

Labium (E. dentipes, Plate 8, figs. 3 and 15).—Transverse, body 
or palpiger subquadrate, with what are apparently the chitinized 
paraglosse expanded laterally and connate with the sides above the 
insertion of the palpi. 

The ventral surface of the palpiger is transversely prominent ven- 
trally between the palpi, and gradually declivous posteriorly to base ; 
rounded laterally basad to the palpi, apicad passing into the concave 
and expanded paraglosse, which are directed outward and forward, 
and rounded at apex. The basal joints of the palpi glide over the 
concave surface of the paraglosse. 

The apical margin of the palpiger is not noticeably membranous, 
but rapidly and obliquely beveled from the prominent interpalpal 
surface, at its oral margin giving attachment each side of a brief 
central interval, to a thick transverse tuft of rather stout but soft 
golden cilia, which curve upward and forward. The corresponding 
oral margins of the paraglosse are also frimbriate with similar cilia, 
which increase somewhat in length from within outward, longest at 
apex and gradually diminishing to some extent externally, as they are 
traced backward and inward along the floor of the mouth. The 
cilia on the paraglosse are directed inward and forward as shown 
in fig. 3, Plate 8. 

Labial palpi (Plate 8, fig. 3) rather small and three-jointed, in- 
serted into the sides of the palpiger ventrad to the paraglose. Joints 
one and two subequal, the first narrower and obconical, the second 
slightly stouter and of similar form, the third a little longer, scarcely 
triangular (subpyriform), feebly flattened and constricted at base 
within. (Compare figs. 3, 5, and 6, Plate 8.) 

Ligula (FE. dentipes, Plate 8, fig. 13).—The internal surface of the 
labium is membranous and marked by a median groove that ends 
posteriorly at an oval but small eminence; anteriorly just within the 
apical margin on each side of the median groove is a rounded convex- 
ity; they he within a triangular area bounded anteriorly by the cilia 
of the apical margin, and laterally by two converging rows of cilia, 
which pass inward and backward from the apices of the paraglosse 
to terminate at the oval eminence. Externally to the rows of cilia 
the surface declines to the external margins of the paraglosse, and 
scattered over which are a number of rather short hairs. Upon the 
oval membranous eminence at the apex of the triangle is a small ar- 
cuate row of very small sete; these are at the proximal end. 

Prothorax.—Inlet bounded above by the apical margin of the 
pronotum, below by the apical margin of the prosternum. Laterally 
within the apical angles is a small triangular area, which may be 


REVISION OF ELEODIINI—BLAISDELL. 13 


termed the apical triangle. It is bounded above by the apical pro- 
notal margin, within by the margin of the inlet, and below by a 
feebly marked ridge that passes downward from the apex. The out- 
let is circular and its plane is quite strongly oblique, while that of the 
inlet is quite vertical. The dorsum is nearly twice as long as the 
sternum. Ventrally along the margin of the outlet the surface is 
rather widely impressed from side to side and behind the acetabula. 
The impression is interrupted at the middle by the prosternal mucro. 

Propleure (prothoracic episterna and epimera) with parts not de- 
fined from each other nor from the inflexed sides of the pronotum ; 
limited internally before the acetabula by the prosternal suture; pos- 
teriorly no sutures are visible. Surface more or less prominent from 
the external margin of each acetabulum. 

Prosternum with the coxe more or less protuberant ventrally. 
Anterior margin broadly sinuate in circular arc, beaded, and usually 
narrowly deflexed. Prosternal sutures distinct before the acetabula, 
straight, impressed and subfoveate in basal half. Each suture ex- 
tends from the external acetabular margin to lateral margin of the 
prothoracic inlet, where it bounds the inner side of the apical triangle. 

Mesothorax.—Condyle annular. The dorsal part of the condyle 
is formed by the transverse and somewhat triangular scutellum, the 
apex entering between the elytral bases to appear visible as a small, 
glabrous, and impunctate triangular plate, marked off from the 
scutellar body by a strong surface deflexion, which is in line with a 
similar deflexion across each elytral base, the condylar portion be- 
ing on a lower plane than that which enters the elytral disk. The 
condylar portion is minutely punctato-scabrous and finely pubescent, 
its basal margin is feebly and broadly emarginate, with the angles 
projecting a little beyond the episternal border, and at which point 
the two plates are not closely articulated, being separated laterally 
from the episterna by the articular processes of the elytra. 

Mesosternum.—It presents for examination two parts—a condylar 
and an acetabular portion. 

The condylar portion is directed horizontally forward, and arcuate 
from side to side, forming apparently, when viewed from below in 
reference to the greatest transverse diameter of the inlet, the middle 
two-fourths ventrally of the condylar wall. The anterior or free 
margin is beaded and feebly arcuate anteriorly. From the middle of 
the margin there extends backward a short glabrous carina, which 
is slightly dilated where it is continuous with the marginal bead; 
surface punctato-scabrous and finely pubescent. 

The acetabular portion is more or less declivous posteriorly from 
the horizontal condylar part, the middle third posteriorly entering 
between the mesacetabula forming the intercoxal salient of the me- 
sosternum, where it may become briefly horizontal, and its apical 


14 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


margin truncate, feebly arcuate or sinuate even in individuals of the 
same species, widely separating the mesocoxe amd terminating at 
the transverse mid-acetabular line; each lateral third of the posterior 
border of the mesosternum is arcuately reéntrant, forming the anterior 
moiety of each mesacetabular cavity, and terminating in the externo- 
posterior angle, which is rather broad and transversely truneate, 
forming the anterior boundary of the mesacetabular recess (see Ace- 
tabula) ; the surface is more or less concave at middle between the 
convex and prominent acetabular margins. 

The sides of the mesosternum are limited by the mesosternal sutures ; 
these are straight and extend obliquely forward and inward from 
the external angle of the mesacetabular recess, to the condylar mar- 
gin, at a point whose distance from the mid-line is about equal to the 
length of the condylar portion in the mid-line before the prominent 
intercoxal part. The sutures join the mesosternum and episterna 
only. 

Mesosternal episterna.—Each episternum is triangular and with its 
surface on the same general plane; anteriorly forming the condylar 
margin between that formed by the mesosternum below and the meso- 
notum above; internally uniting with the mesosternum; apical angle 
received between the epimeron and mesosternum at the angle of the 
mesacetabular recess; externally and dorsally connate with the epi- 
pleural margin and base of the elytron external to its articular process. 

The surface is divided into two areas by a gentle arcuate line or 
ridge which extends transversely from about the middle of the meso- 
sternal suture to the inferior humeral angle. This line may be called 
the episternal line. 

The area anterior to the episternal line is the condylar portion; its 
surface is finely punctulo-scabrous and pubescent, and subdivided 
into two unequal portions by a submarginal line, which, beginning at 
the antero-internal angle runs outward nearly parallel to the mar- 
ginal bead, thence gently diverging, to arcuately pass to the middle 
of the superior margin of the condylar area. 

Mesosternal epimera.—FEach epimeron is rhomboidal, with its in- 
ferior border bounding the mesacetabular recess externally; searcely 
twice as wide as long, obliquely placed between the mesosternal epi- 
sternum in front and the metasternal episternum behind, and if its 
posterior margin be prolonged as a line it will touch the internal angle 
of the metacoxa of the same side. 

The superior margins of the mesosternal episterna and epimera are 
inflexed beneath the inferior epipleural margins of the elytra, at the 
line of inflexion where they pass under, the surface is grooved and 
beveled to receive and hold the elytral margins; the inflexed portions 
are continuous with less strongly chitinized dorsal and undifferen- 
tiated sclerites, which with similarly inflexed parts of the metasternal 


REVISION OF ELEODITINI—BLAISDELL., 15 


episterna, form between them the boundaries of a rather large fossa, 
in the deep and somewhat anterior wall of which is situated the meso- 
thoracic stigma. This stigmatic fossa is situated immediately within 
the mesosternal epimera under cover of the elytra. 

Metasternum short, the median part or body is subquadrate, some- 
what equal in length to the abdominal intercoxal process, anteriorly 
entering between the mesacetabula quite equally with the mesosternal 
salient and forming with it the intersternal suture; the latter is either 
straight or arcuate, according to whether the intercoxal processes are 
truncate or more or less rounded—a variable condition even within 
specific limits; posteriorly the border is transverse, not in the least 
entering between the metacetabula. 

The sides of the body are produced laterally, much narrowed be- 
tween the mesacetabula in front and the metacetabula behind to 
expand somewhat triangularly more externally against the episterna, 
as the ale. The anterior borders of the ale, laterad to the intercoxal 
process, are arcuately reéntrant to form the posterior moieties of the 
mesacetabula; the posterior borders of the ale are less strongly and 
arcuately reéntrant to form the anterior boundaries of the metrace- 
tabula. The basal borders of the ale are very feebly reéntrant and 
rounded to fit the feebly arcuate edges of the episterna, forming the 
metasternal sutures, each extending from the external angle of the 
mesacetabular recess in front to the external angle of the metacetab- 
ulum behind. The anterior basal angle of the metasternal ala is 
shut off from the externo-posterior angle of the mesosternum by the 
mesosternal epimeron at the external angle of the mesacetabular 
recess; the posterior basal angle of the ala is prevented from meeting 
the first abdominal segment by the interposed episternum at the ex- 
ternal metacetabular angle. 

Metasternal episternum.—Elongately subrhomboidal, slightly wid- 
ening at middle, thence feebly narrowing to the posterior border, 
about three times longer than wide and about three and a half times 
longer than the mesosternal epimeron; base obliquely truncate and 
connate with the posterior edge of the mesosternal epimeron, forming 
the mesepimeral suture; internal border feebly arcuate and connate 
with the base of the metasternal ala; the external margin is straight 
and adapted to the inferior epipleural margin. The posterior edge 
internally is somewhat inflexed at the external angle of the metace- 
tabulum, externally giving attachment to the small subacutely digiti- 
form metasternal epimeron, which is feebly and inwardly arcuate, 
resting in a small depression on the surface of the first abdominal 
segment at the externo-anterior angle, and bounding the metacetab- 
ulum at its external angle. 

The metathoracic stigma is situated immediately opposite to the 
posterior extremity of the metasternal episternum and under cover 


16 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


of the elytra; unlike the mesothoracic stigma it is surrounded by the 
soft cuticle; it is large and deeply impressed andethe rima stigmatis 
is antero-posteriorly directed; that of the mesothoracic stigma is di- 
rected antero-internally. 

The dorsum of the mesothorax is more or less chitinized laterally, 
less so at middle behind the scutellum. Invisible without removal 
of the elytra. 

The dorsum of the metathorax is membranous except laterally 
where the episternum extends inward for a short distance beneath 
the epipleura. 

The proacetabula are quite circular, except at the internal margin 
where the intercoxal prosternal process dilates slightly and impinges 
upon them. 

The mesacetabula are quite circular if the recess for the trochan- 
tine be not included, with that it is slightly transverse. The anterior 
margin of the acetabular recess is more or less angulate from the 
truncate externo-posterior angle of the mesosternum; the posterior 
margin is continuously arcuate with that of the acetabulum proper. 

The metacetabula are slightly more transverse, but slightly and 
obtusely angulate anteriorly where the posterior angles of the meta- 
sternal ale form their boundary. 

The procove are globose. 

The mesocove are really slightly transverse, but subdivided by an 
antero-posterior impression into the rounded coxa proper and the 
outer and smaller division,—the trochantine which occupies the ace- 
tabular recess. The coxa proper is quite broadly and outwardly im- 
pressed from the articular cavity to the trochantine. 

The metacovw are more transverse, ovate and not divided by an 
antero-posterior impression; the surface of each is broadly and 
strongly grooved external to the articular cavity. 

The trochanters are small and ovate, scarcely forming a half of the 
width of the basal extremity of a femur, and but a little longer than 
wide. 

The crural characters are given under the different specifie head- 
ings. For a study of the relative lengths of the different segments 
see diagram, fig. 4, Plate 8. 

The grooves on the inferior surfaces of the femora I have denomi- 
nated as the tibial grooves; because they are adapted to and receive 
the tibiz in extreme flexion. 

Likewise the grooves observed on the external surfaces of the 
tibiw I call the tarsal grooves. These tarsal grooves are not always 
present in the different species; when present they extend from the 
more or less open articular cavities on the tibial apices, to become 
evanescent at varying distances from the same. These tarsal grooves 
receive the tarsi in hyperextension. 


REVISION OF ELEODIINI—BLAISDELL. 17 


The tarsi are grooved beneath by the plantar grooves. 

Each tarsal joint is more or less obconical and grooved beneath. 
The sides of the groove are set with a row of marginal spinules, the 
apical margin also bears a row of apical spinules. At the apex where 
the apical meet the marginal row, the spinules are usually longer and 
more conspicuous as acute tufts; these are the apico-marginal tufts. 
(See Plate 8, fig. 220.) 

The first two joints of the anterior tarsi are frequently thickened 
and somewhat produced at apex beneath (Plate 8, fig. 92), obliterat- 
ing the groove. When the apex is thickened beneath it nearly always 
bears a tuft of yellowish pubescence, especially in the males of many 
species, and sometimes in the female a tuft of spinules. 

In the males of the subgenus Blapylis it is normal to observe on 
the first and second joints of the mesotarsi similar tufts on the tips 
beneath. This is also observed in some species of Promus. 

The ultimate joint of each tarsus at apex has the margin above 
feebly bilobed; beneath the margin is produced into an interungual 
process, which varies considerably in shape but is probably of no 
value in the separation of species. (See Plate 6, fig. 18.) 

The ungues are slender and acute, moderately arcuate, and slightly 
subangulate at base beneath. (Plate 6, fig. 19.) 

Abdomen.—Ventrally there are only five visible segments, while 
dorsally there are six. 

Morphologically the first visible ventral segment is the second; 
the study of the retractile segments requires that this be recognized 
as such. 

The first segment dorsally in the mid-line is as long as the second; 
laterally it gradually narrows so that it points at the bases of the 
metasternal epimera. 

Ventrally the first segment is greatly reduced, and although un- 
differentiated it forms a part of the metacetabular walls, reaching 
the sides of the abdomen to appear as a small and short sclerite, con- 
nate with the externo-anterior angle of the second segment (first 
visible), dorsad to the base of the metasternal epimeron, and at the 
point where the evident dorsal membranous first segment reaches the 
abdominal margin. 

There are six pairs of abdominal stigmata, and in the male are 
located on each side as follows: 

The first stigma is situated about 4 mm. above the posterior margin 
of the metathoracic stigma. 

The second is about 0.8 mm. above the lateral edge of the second 
ventral (first visible) segment and on a line drawn transversely 
across the posterior margins of the metacetabula. 

59780—Bull. 683—09 2 


18 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


The third is 0.6 mm. above the basal angle of the third ventral 
(second visible) segment. . 

The fourth immediately above and close to the inner membranous 
edge and basal angle of fourth ventral (third visible) segment. 

The 7fth is in the same position relative to the fifth ventral (fourth 
visible) segment, and only partly visible without traction on the 
sixth tergite or a voluntary extension of the same by the insect. 

The séxth is in the same position relative to the sixth ventral (fifth 
visible) segment as the fourth, but entirely invisible without trac- 
tion on the sixth tergite. It is distinctly smaller than the other 
stigmata and is overlapped by the lateral fold or edge of the sixth 
tergite and also by the apical fold of the fifth. At this point the 

The tergite of the sixth segment (Plate 7, figs. 27 and 4p) is 
the other segments. 

The first three pairs of stigmata are the largest, the fourth and 
fifth are slightly smaller. 

The tergite of the sixth segment (Plate 7, figs. 2p and 4p) is 
moderately chitinized and flexible, evenly and arcuately rounded at 
apex, the margin set with short sete and sparsely so over the dorsal 
surface apically. 

The sternite is strongly and firmly chitinized and constitutes the 
so-called fifth visible ventral segment. 

Retractile segments.—F our in number and homologous to those of 
Hydrophilus piceus as figured by Packard.t 

Seventh seqment.—The tergite and sternite are chitinized and very 
flexible. The tergite is short, broadly, and not strongly arcuate at 
apex, most strongly chitinized laterally and quite membranous at 
middle; margin frimbriate with short sete and the dorsal surface 
sparsely pubescent. (Plate 7, figs. 27 and 47.) 

The lateral angles at base articulate with the lateral and basal 
angles of the sternite and in both sclerites these angles are more 
strongly chitinous than the other parts of the same sclerite. This 
articulation is stronger in the male. 

The sternite is different in the sexes. (See figs. 2 and 5, Plate 2, 
and figs. le and 30, Plate 7.) In the male it is bilobed and described 
in detail for each species in the body of this work; in the female 
it is broadly arcuate at apex, in the male deeply emarginate between 
the lobes. 

In both sexes each lateral moiety is made up of a triangular 
chitinous plate, which at the external and internal basal angles are 
more or less produced. In the male the internal and more or less 
anteriorly produced angles are separate and not connate—so as to 
permit of the abduction and adduction of the lobes which no doubt 


“ Text-Book Ent., p. 181. 


REVISION OF ELEODIINI—BLAISDELL. 19 


act as guides to and steady the edeagophore under certain condi- 
tions; in the female the interval between these triangular sclerites 
is continuously membranous and feebly chitinized, so that movement 
between the lateral moieties is impossible, and the internal basal 
processes are connate. (Plate 7, fig. 30’.) The external basal proc- 
esses articulate with the tergite of the same segment as described 
above. 

The eighth segment is membranous in both sexes. 

The tenth segment is aborted in the female as usual, the tergite 
fusing with the ninth tergite to constitute the supra-anal plate. 
(Plate 7, fig. 4g.) In the male the tergite of the tenth segment 
forms the supra-anal plate; the subanal or sternal wall is mem- 
branous in both sexes, possibly somewhat chitinous in the male. 

The supra-anal plate is more strongly chitinous laterally and more 
membranous at middle (Plate 7, figs. 27 and 47) and is in relation 
laterally with the paranal plates or sclerites. 

In the male these paranal sclerites are much smaller and less tri- 
angular than in the female, and articulate with a chitinous rod 
on each side; these rods converge anteriorly so as to form a Y-shaped 
support and a mechanism for the extension and retraction of the 
edeagophore ; they are wholly internal and give attachment to muscles 
and ligaments. 

The paranal plates in the female are irregularly subquadrate, 
occupying the lateral wall below the supra-anal plate. Each paranal 
sclerite may be said to present for examination four borders: a 
superior, posterior, inferior, and an anterior; four angles: the antero- 
inferior and postero-inferior being the most important. 

The superior border is connected to the sides of the supra-anal 
plate by membrane, apparently forming an articulation at its antero- 
superior angle with the antero-external angle of the anal plate; the 
posterior border is connected by articulating membrane to the basal 
margin of the ventral plate or valve; the inferior border gives attach- 
ment to the membrane of the ventral wall, and the anterior border 
giving attachment to the membrane of the eighth segment. 

The antero-ventral angle is short and apparently gives attach- 
ment to muscles and ligaments which steady the plate against the 
valve. 

The postero-ventral angle is produced into the process of the 
paranal plate to articulate with the strongly chitinized basal margin 
of the ventro-lateral plate of the valve, of the same side and upon 
which the valve plays during abduction and adduction. (Plate 3. 
fig. 4a.) 

The above description of the retractile segments has been founded 
upon those of Lleodes cordata, as illustrated in Plate 7. The seg- 


20 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


ments were studied both as opaque objects and cleared with: liquor 
potassi. . 

The sclerites of these segments vary a great deal in the different 
tribes and even in the genera of the Tenebrionide and are of taxo- 
nomic value. 

The development of these segments depends a great deal upon the 
habits of the insect and the manner of oviposition required by the 
species. In Microschatia inwqualis they are developed to a wonder- 
ful degree and are capable of extrusion to the distance of nearly an 
inch. 

The different species of the present tribe exhibit considerable 
variation in this respect and which might be of aid in the classificea- 
tion of species; but on account of the great amount of work and 
material required and the time already devoted to this paper, I have 
not been able to study them nor deemed it wise to further delay the 
publication of work already done and therefore leave that investiga- 
tion for a future thesis. 

The male intromittent apparatus.—This consists of a carrier, the 
Edeagophore, and the intromittent organ proper, the /deagus, and 
its clave. 

Edeagophore, usually more or less flax-seed-shaped, somewhat 
arched, and consisting of two segments or lobes: a basal segment or 
body, the basale; an apical segment or apicale. 

The apparatus has been described in detail for each species. 

The basale is variable in length, half-cylindrical in form, chitinous 
above and laterally, membranous beneath, decurved at base into a 
process which gives attachment to muscles and ligaments by which 
it is attached to and operated upon by the extrusor mechanism de- 
scribed above; dorsally it is convex to a varying degree and often 
depressed, the sides are more or less inflexed beneath for the attach- 
ment of the ventral membrane—strongly so in Embaphion and 
Eleodes planipennis; the apex is more or less emarginate and adapted 
to the base of the apicale to which it is attached by articulating mem- 
brane. (See Plate 7, fig. 2; Plate 2, figs. 1 and 8; Plate 9, figs. 3 to 9.) 

The apicale is always triangular, ravely subpyriform, more or less 
depressed and usually subacute at apex, sometimes obtuse; the base 
is more or less lobed and received into the emargination at the apex 
of the basale; the dorsal surface is more or less convex, frequently 
membranous at middle, but in the higher forms entirely chitinous; 
ventral surface is more or less membranous, but chiefly occupied by 
the edeagus and its protecting clave, the inflexed and chitinous sides 
constitute the ale which occupy the greater part of the surface; in 
one genus the ale support subale. (leodimorpha, Plate 7, fig. 8.) 

The alw are inflexed to a varying degree, strongly so when the 
ventral surface of the apicale is concave, as in Lleodes cordata 


REVISION OF ELEODITNI—BLAISDELL. 21 


(Plate 7, fig. 1) and /. dentipes (Plate 9, fig. 6) ; convex ventrally in 
Embaphion (Plate 7, fig. Ty) and Eleodes planipennis (Plate 9, 
fig. 7); in Lleodes scabricula the inferior surface of the apicale is 
concave apicad to the ale, which are suddenly prominent. In the 
different species the ale are continued basad to a varying degree, be- 
coming more and more membranous. (See figs. 6, 7, and 9, Plate 9.) 

The Hdeagus (Plate 9, fig. 11a) is usually flattened, sublingui- 
form in shape and subacute at apex; the ventral surface is narrowly 
prominent in the median line and more or less concave laterally where 
it is in contact with the clave in repose; the median ridge, divides 
toward apex to become bilabiate, between which is probably the ori- 
fice of the vas deferens. At base the structure is more chitinous and 
there is a small median process for articulating with the clave. 
The dorsal surface is feebly convex and more or less membranous 
and marked by a median groove: opposite the attachment of the 
clave ventrally the surface membrane is reflected upon the internal 
wall of the apicale, forming the fornix edcagi. In fig. 12 of Plate 
9 the orifice of the detached vas deferens can be seen entering the 
edeagus from the basale just beneath the fornix. 

At the base the sides of the edeagus are strongly chitinized and 
produced into two long processes (Plate 9, figs. 11a@ and 12) which 
extend cephalad within the interior of the basale, giving attachment 
to muscles. 

The clave are usually quite spiculiform (Plate 2, figs. 1 and 8), 
sometimes sublaminately spiculiform (see figs. 6, 11d and 12, Plate 9) 
and in Embaphion truncate at tip and concave beneath ; in 7eodimor- 
pha they are rather liguliform (see figs. 7 and 8, Plate 7). 

In repose when the edeagus is retracted they he parallel to the 
median line and more or less contiguous; with the edeagus pro- 
truded they are at right angles to the axis of the edeagophore. 

Each clava may be said to have a base and an apex, a dorsal and a 
ventral surface, and internal or posterior surface and an external or 
anterior margin. 

The base is obliquely truncate at the posterior or internal surface 
and articulated to the ventral surface of the edeagus at base; the 
internal or posterior surface or edge is much thicker than the anterior 
or external, and triangular in basal half, and thence gradually be- 
coming more ventral as it attains the apex, where it is feebly con- 
vex, the basal portion is quite plane; apex subacute to rounded or 
rather obliquely truncate; the external or anterior edge is usually 
thin, feebly arcuate and slightly notched near the apex; the dorsal 
surface more or less convex and the ventral rather concave. (/. 
dentipes, Plate 9, figs. 11d and 12. 

In repose the clave may conceal the edeagus, except at apex, 
where they slightly diverge, when they are ventral as in //eodes,; or 


22 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


the median ventral surface of the edeagus, may be visible between 
them when they are lateral as in Embaphion, ang ventrad but not 
contiguous as in Lleodimorpha. 

The female genital segment (Plate 7, figs. 3 and 4).—Viewed from 
above it has an outline varying somewhat according to the subgenus: 
triangular in Blapylis, quadrate in Melaneleodes, trapezoidal in 
Eleodes and subparabolic in Steneleodes. 

The dorsal surface varies but is usually more or less plane to sub- 
ogival; ventrally convex—strongly so at base and gradually less so 
apically. 

The segment consists of two chitinous valves concealing and pro- 
tecting the pudenda between them. The valves are movable on a lat- 
eral plane, in abduction exposing the genital orifice and its sur- 
rounding pudendal membrane. 

{ach valve (valvula) consists of a body (corpus valvule) and an 
apex (apex valvule). 

The body is always chitinous; the apex usually membranous and 
distinctly define from the body, sometimes completely chitinous and 
not differentiated. 

The superior wall of each valve is formed by the dorsal plate (Jamina 
dorsalis) which is variable in form; its surface usually more or less 
plane, but may be quite strongly concave. Each plate presents an 
external, internal, and apical border; at the junction of the external 
and apical borders an angle is formed, which may be distinct or more ~ 
or less obsolete. 

The internal borders of the dorsal plates are not usually contiguous 
but distant at base, converging posteriorly, and between which is the 
more or less exposed superior pudendal membrane. The apical mar- 
gin is generally defined from the surface of the apex; in some species 
it is not. The external border is generally more or less explanate, 
and beneath which is situated the submarginal groove (sulcus sub- 
marginalis ). 

The lateral and ventral walls of a valve are formed by a continuous 
plate, the lamina ventro-lateralis, which is connate above with the 
dorsal plate. It presents for examination a surface and an internal 
border. The surface is strongly convex at base, frequently prominent 
laterally, forming the basal prominences, much less convex apically 
and generally more or less impressed before the apex, with which it 
is directly continuous. 

The internal margins of the ventro-lateral plates of the two valves 
are not usually contiguous throughout their entire length in the mid- 
ventral line, but separated by the more or less fusiform genital 
fissure (fissura genitalis). When this fissure is basad to the middle 
it is spoken of as being subbasal; when at the middle it is median, and 


REVISION OF ELEODITNI—BLAISDELL. 23 


when apicad to the middle it is subapical. The fissure when present is 
closed by the inferior pudendal membrane. 

The basal margins of the ventro-lateral plates are strongly chiti- 
nized and together form a loose articulation at their basal angles, 
basad to the genital fissure. 

The basal margin of each ventro-lateral plate articulates about its 
middle with the process of the paranal plate, and is connected by 
articulating membrane with the posterior margin of the same plate 
(Plate 3, fig. 4a); dorsally the dorsal plates do not articulate with 
each other, but on the contrary are widely separated and connected 
only by membrane. 

The apex of each valve is separated by a fossa into two lobes, an 
external and an internal. The external l6be is only developed in the 
subgenus JZ/elaneleodes; in all others of our fauna it is obsolete. The 
internal lobe is always developed and only atrophic in J/elaneleodes. 

The fossa gives attachment and lodgment to the always short 
cercopod, 

Each valve is more or less finely pubescent about the apex, the 
dorsal plate is most noticeably pubescent in the subgenus Promus. 

The cereopoda are variable in form and size, usually mammilliform, 
frequently compressed, punctiform in Déscogenia and obsolete in 
Heteropromus. Usually penicillate. 

The pubescence on the apex is usually longer than elsewhere, and 
the internal lobe is often somewhat penicillate. 

The valvular body is never very strongly punctate, most noticeably 
so on the external moiety of the dorsal plate in Promus. The punc- 
tures are rather strong in some species of Stene/eodes, and the superior 
pudendal membrane is broadly exposed. 

From the sides of the vulva the lateral pudendal membrane is 
reflected upon the internal surface of the valve and there becomes the 
valvular membrane (Plate 7, figs. 3v and 4”), apically, directly 
continuous with that of the internal surface of the apex. 

The angle of reflection between the lateral pudendal membrane and 
valvular membrane is the fornix valvule. In some species the lateral 
pudendal membrane passes backward directly upon the internal sur- 
face of the valve without forming an angle of reflection. 

Function—The edeagophore is the agent by which the repro- 
ductive organs are brought into apposition. The apicale by its sub- 
acute apex is capable of more or less forcibly separating the tergal 
and sternal sclerites of the sixth and seventh somites and protects 
the essential organ from injury. 

With the protrusion of the edeagus and its approximation to the 
genital fissure, the simultaneous abduction of the clave forcibly ab- 
ducts the valvulee and blocks their closure. 


24 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Homology.—The examination of the male reproductive organs of 
Hydrophilus triangularis (Plate 9, figs. 1 and 2) suggests the homol- 
ogy between the reproductive organs in the Hleodiini, and also be- 
tween the former and those of the latter. 

In Hydrophilus the edeagophore consists of a basale (Plate 9, fig. 
1d) and two apical valves. The basale is impressed along the me- 
dian line dorsally and is more or less membranous at that point, and 
it therefore appears to have been developed by the partial coalescence 
of two lateral sclerites, termed by Packard the outer lobes; the two 
apical lobes he terms the inner lobes (Plate 9, fig. 1a), they are valvu- 
lar in character and capable of free abduction and adduction; be- 
tween them is the edeagus. 

I consider the outer lobes homologous with the basale, and the 
inner lobes with the apicale in the /7eodiini. 

If fig. 1 of Plate 9 be compared with fig. 12, Plate 2, a similar 
structural condition of the valves in the two sexes and in very widely 
separated series will be observed. It is evident that in Hydrophilus 
there has been less sexual differentiation than in the Aleoddini, and 
also if the male characters of the latter be considered. 

The female genital segment of Zydrophilus (Plate 9, fig. 10) pre- 
sents a complicated arrangement of sclerites which are noticeably 
divergent from anything observed in the Veodiini, and must be 
considered as a more primitive condition, in this agreeing with the 
male characters. 

It has been made known by Nussbaum that unpaired parts lke 
the edeagus, etc., have been developed from paired ectodermal out- 
growths which have fused together, and in all probability there is 
analogy between these outgrowths in the Insecta and those observed 
in the Vertebrate Series; if this be so, then there must at some period 
have been an indifferent stage in the development of these organs, 
and a time of beginning sexual differentiation. 

It appears to me that the apical lobes of Zydrophilus and the api- 
cale of the Hleodiini are homologous with the genital segment of the 
female. In Hydrophilus there are the two valves capable of free 
»bduction and adduction with the edeagus between and protected by 
them; in the female of the /Jeodiini the two valves are as capable of 
free movement as in the above and enclose and protect the sexual 
orifice. 

In the female the valves articulate with large paranal plates; in 
the male the valves or apicale articulate with the outer lobes or basale, 
and the paranal plates are small and articulate with a Y-shaped 
chitinous structure that is internal, the basal process of the basale 
articulating with the arms of the Y by ligaments. 


REVISION OF ELEODITNI—BLAISDELL., 25 


It appears to me that the outer lobes (Plate 9, fig. 1d) which unite 
to form the basale are the modified greater part, at least, of the 
paranal plates. 

There must be a definite law governing the deposition of chitin; it 
ean not be diffusely deposited—at random, as it were—but it must be 
first laid down at certain points, called the centers of chitinization, 
and from these centers spread, or have inherent limitations; different 
centers coalescing to form larger sclerites or one continuous whole, as 
in the prothorax. A simple sclerite would then be one formed by the 
secretion of chitin about a single center; a compound sclerite from 
two or more centers. The small but symmetrically arranged sclerites 
observed in the cuticle of larvee may be cited as simple sclerites. 

It might be assumed that the paranal plate in the female is com- 
pound; in the male these centers have become shifted, to form the 
small paranal plate, and the other to become the lateral sclerite of the 
basale. 

The origin of the several sclerites in the female genital segment of 
Hydrophilus may be accounted for in this way—they have not coa- 
lesced and have been shifted according to the laws governing adapta- 
tion to function ; during divergence and differentiation under changed 
conditions of environment these several sclerites fuse—of course it 
is possible for some to be suppressed, others hypertrophied or 
even revived—and in this manner produce the apparently simple 
chitinous valves of the Eleodiini. These suggestions are to indicate 
a line of investigation in the coleopteral series. 

An examination of the edeagophore of Blaps (Plate 9, figs. 3 and 
4) will be instructive. It will be seen that the apicale is divided 
longitudinally by a well-marked membranous impressed groove and 
that the apex is bifid; the basale is also marked by a similar medium 
groove. These characters indicate a structural condition somewhat 
intermediate between the /7eodiini and a more primitive state as 
suggested by Hydrophilus. 

If it be imagined that the apicale of Blaps was completely divided 
along the groove, and the ale not strongly inflexed, the edeagus and 
its pallium would bear the same relation to the valves as does the 
edeagus to the valves in /ydrophilus. There is greater specializa- 
tion in the “Veodiini than in Hydrophilus or Blaps. 

In £leodes and other members of the //eodiini the apicale is fre- 
quently marked dorsally by a membranous area or groove, and less 
frequently the basale has an impressed median line. Many species 
do not normally have this groove and such I consider to be the more 
highly differentiated, although occasionally it occurs on the apicale 
even in them, and is then to be considered as an instance of ataval 
reversion. 


26 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


It is to be noted that Escherich, in his paper on the internal male 
organs of beetles, notes that those of the Carahjde illustrate the 
simplest, most primitive condition. Packard figures Blaps as inter- 
mediate between the Carabide and Hydrophilus.¢ 

Sculpturing—From simple punctures the sculpturing passes 
through the different degrees of modification to distinct murication 
or tuberculation, or as in a few species to distinct hairiness with com- 
paratively simple punctures. 

From a careful study of the sculpturing under high power in the 
species of the Hleodiini, I have observed the following: 

A simple puncture is a more or less rounded depression, variable in 
size and depth, and bounded by a more or less distinct, although 
fine wall or margin; from the floor of this depression arises a minute 
hair or seta, which scarcely passes beyond the mouth of the pune- 
ture. The surface with such a punctuation is smooth. 

Any modification will affect the puncture in one of two ways: 
Either by a hypertrophy of its margin or by an increased growth or 
hypertrophy of the hair-like process. In the first instance the hair 
may remain small or atrophic; in the second the hair-like process 
is alone affected. Some punctures may become completely aborted. 

The hair-like process arising from the hypodermis may develop 
into a soft, flexible hair, or by a slightly increased chitinization of its 
cuticle into a bristle or seta, and by still greater increased chitiniza- 
tion, with or without hypertrophy, into a spinule or spine. 

In the muricate type of sculpturing the anterior lip of a puncture 
is thickened and more or less produced into a blunt or acute point; 
from the sides of this prominence the margins gradually diminish 
so that the posterior boundary of the puncture will be but slightly 
modified; the thickened edge is directed upward and backward, 
chiefly the latter; the hair or seta may be aborted, remain small, or 
become more chitinous to become a seta, and be carried upward by 
the hypertrophied margin to project from the posterior surface of 
the more prominent anterior margin of the puncture. 

When the margin of the puncture is equal and much thickened a 
tubercle is formed, which varies in size from a granule to a well 
marked and rounded eminence; the puncture may be entirely closed 
or obliterated, or may still be evident as a minute pore-like depression 
with a correspondingly small hair or seta projecting from it, and 
from some point, usually the posterior wall of the tubercle. There 
are all the intermediate degrees of variation to be observed. 

The punctures may remain simple and the hair become long and 
flexible. 

If the anterior lip of the puncture becomes minutely muricate the 
sculpturing is more or less asperate. 


@'Text-Book Ent., p. 494, fig. 465, A, B, and C. 


REVISION OF ELEODITNI—BLAISDELL. 27 


On the femora the proximal margin of a puncture is transversely 
raised so that the seta arising from it is more or less appressed to the 
surface and directed toward the apex of the femur. 

On the tibix the murications are more erect, the walls of the punc- 
tures being more equally raised. The hypodermal process is usually 
much thickened to become a spinule or spine—the latter may be very 
short and stout, and with the produced margins of the puncture from 
which it arises, very tightly fitted to it, so that both appear as a single 
element. 

The rugulose condition or reticulations frequently observed on the 
elytral surface is most usually due to the rapid evaporation of the 
body fluids during and immediately after the casting of the pupal 
skin, while the integuments are undergoing chitinization. It is the 
result of shrinking of the cuticular wall. 

Often depressions arise in the exoskeletal wall from muscular 
exertion on the part of the insect while the body surface is yet soft. 
The contracting muscle dimples in the integument at the point of its 
attachment; the movements being kept up while the body walls are 
hardening, the depressions finally become permanent. Such uni- 
lateral or bilateral impressions are frequently observed on the pronota 
of the Eleodiini. Eleodes impressicollis of Bohemann, a synonym of 
E.. clavicornis is an instance of this kind—merely a specimen with 
- two thoracic fover. 


TABULATED MODIFICATIONS OF A PUNCTURE, 


With hypertrophy of the an- 
terior margin into a more or 
Hypertrophy of} less acute point. 
the margin. Margins greatly thickened and 
rounded, puncture obliterated, }Tuberculation. 
hair or seta atrophied. 


Muricate punc- 
tuation. 


Simple punc- 
ture, hair Hairs flexible, not chitinous -- -. 
minute. 


Hairy or pu- 
bescent. 
the hair-like) chitinized. ct so aan eae 
paces of the 
ypodermis. |Still greater degree of chitiniza- 
tion. 


Spinules or 
spinulose; 
spines or 


Modification of|Hair-like processes in a 
| spinose. 


CLASSIFICATION AND SYNOPSIS. 
Family TENEBRIONID.®. 


Tribe HLEODIINI. 


This tribe contains insects of greater size than any other of the 
heteromerous subfamily Tenebrionine. 

The body is oblong to ovate, rarely oval, apterous; head prominent, 
slightly narrowed behind the eyes; epistoma covering the base of the 
mandibles at the sides; labrum prominent; mentuni small, inserted 


28 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 2 


upon a gular peduncle; maxilla exposed; ligula partly concealed ; 
maxillary palpi with the last joint securiform, met very large; eyes 
transverse, reniform, tolerably finely granulated; antennze 11-jointed ; 
elytra embracing widely the planks of the abdomen, epipleure narrow 
to moderately wide, middle coxew with large trochantine, side pieces 
attaining the coxal cavities; metasternum very short, episterna nar- 
row, epimera quite distinct; hind cox widely separated; intercoxal 
process of abdomen rectangular; third and fourth ventral segments 
not prolonged behind at the margin. 

Legs long; anterior femora frequently toothed; tibial spurs dis- 
tinct; tarsi channelled beneath; first, second, and third joints of the 
anterior tarsi are at times thickened beneath, more or less obliterating 
the groove, setose beneath. 

All are black or dark brownish, rarely reddish along the suture; 
variously though never very roughly sculptured; a few species are 
pubescent. 

The third and fourth ventral abdominal segments are more or less 
broadly sinuate at apex, the angles appearing somewhat prominent, 
but not noticeably prolonged. 

The tribal comparative genital characters are reserved for a future 
and special study. 

Habits —Our species are terrestrial and cursorial, apparently 
strictly nocturnal, or are about in the early morning or late in the 
day. On warm cloudy days I have seen Hleodes grandicollis and 
FE. acuticauda walking about. : 

The larger species when disturbed place their bodies nearly vertical, 
the head near the ground, tail erect, and when touched or irritated 
will emit a pungent dark oleaginous offensive secretion, which coming 
in contact with ‘the skin of the hands will stain brownish and 
remain for a long time, and apparently fixed by an alkaline sub- 
stance (Horn). 

The species of the present tribe rarely ascend plants; the smaller 
species (Blapylis) may be found on the stems or under the bark of 
shrubs. 

When alarmed they run off with their bodies elevated. Probably 
it is this habit which has suggested the name “ circus bugs,” often 
applied to them in the West (Wickham). In California they are 
often spoken of as “ stink bugs,” “ beetle bugs,” and “ tumble bugs.” 
The latter term is applied no doubt on account of their frequently 
tumbling over when excited; also, pinacate bugs (V. Kellogg). They 
are a characteristic feature of the arid regions west of the Mississippi 
River. 

Longevity.—Some of the larger and more resisting species evidently 
live to be several years of age; especially in the warmer regions of 


REVISION OF ELEODIINI—BLAISDELL. 29 


California (Horn). I have kept several Hleodes dentipes in cap- 
tivity for over four years. Doctor Horn states that specimens have 
been kept pinned two months without food or water. //eodes clavi- 
cormis dies quickly in confinement and apparently lives but one 
season. 

Food.—All are vegetable feeders, apparently preferring dried vege- 
tation and fungi, although eating green plants. 

Fnemies.—Quadrupeds like skunks will feed upon them; chickens 
devour them readily, as well as ground owls; butcher birds impale 
them on thorns. 

Economics.—As far as I have been able to determine they are 
neither injurious nor beneficial, unless the larve are in some way 
troublesome, but as they appear to prefer decaying vegetation to the 
living I doubt that they can be considered objectionable. 

Origin and diffusion —lt is impossible for me at this time to ex- 
_ press any opinion upon the origin and diffusion of the species making 
up the tribe E/eodiini; but I believe the species to a great extent had 
their origin in Mexico, or in adjoining desert regions, and that diffu- 
sion has been chiefly northward and westward. 

I know of no fossil remains of species referable to the tribe, and a 
few that were supposedly so were from a very recent deposit and 
identical with species now existing. (See Appendix.) 

It remains for the study of the Mexican fauna to aid us. The 
South American Vycterinus will be referred to elsewhere. 

Distribution —Abundant both in species and individuals through- 
out the region west of the Mississippi River, from the northern 
boundary line of the United States, southward into Lower Califor- 
nia and Mexico to South America, where they are replaced by 
Nycterinus. They also inhabit the islands adjacent to the western 
coast of the territory above named. 


ANALYTICAL KEY TO GENERA OF ELEODIINI, 


Our genera are five in number, one having been introduced from Europe. 


Outer joints of antenne broader (Hleodiini)_..._.___-__________________ fet | 
Outer joints of antenne not broader; 8-10 moniliform, suddenly shorter than 
PmmoCenine- WiLL (Biaptwe ) =. Dre Tide _Blaps. 

1. Sides of the epistoma not dilated, margin straight or sinuate, converging 
NE Slee See re ge Se oe eS aa 
Sides moderately dilated, margin Breathe ts i hore Ales 2 1 Trogloderus. 
2 Epipleure attaining the humeral angles___........._____________________ 3 
Epipleurze very narrow, not attaining the humeral angles______ BEmbaphion. 


3. Epipleurze broader at base, more or less gradually narrowing to apex. 
Epipleure occupying only a part of the inflexed portion of the elytra; 
buccal processes of the gene not produced Bleodes. 
Epipleure occupying the whole of the inflexed portion of the elytra; 
buccal processes of the genz acutely produced________ Eleodimorpha, 


30 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


The genera can also be defined by the male genital characters as 
follows: ~ 
Sternite (fig. 6, Plate 6) with lobes fully and strongly connate; clave dorsal 

to the edeagus, connate forming a pallium (similis, mucronata) (Blaptini) 

Blaps. 
Sternite distinctly bilobed, lobes connected by a membrane that at most is but 
feebly chitinized, lobes capable of abduction (leodiini). 
Clave ventrad to the edeagus. 

Clavie spiculiform, notched at side of the tips; alze more or less strongly 
inflexed, not prominent ventrally, plane or somewhat concave, never 
strongly convex (Plate °7, fig. 1)------ eee Bleodes. 

Clavie flattened (liguliform), of equal width throughout, tip obtuse 
and rounded; alee short, not strongly inflexed, giving attachment to 
subale (Pinte Tt. 1s Sie: 2 Se See Eleodimorpha. 

Clay laterad to the edeagus, truncate at tip, more or less concave beneath ; 
alee moderately and evenly inflexed, rather prominent ventrally (Plate 
qo fies, Q)igs ete a et ee oe A a) ee ee Embaphion. 

Trogloderus has the lobes of the sternite united by membrane and 
‘rapable of abduction. Unfortunately the clave could not be studied 
without destruction of the fragile material at hand. 

Genealogy —The antecedents of the present tribe are not suggested 
to my mind by the data at hand. 

The male sexual characters are the most distinctly differentiated 
for the generic subdivision of the tribe. The above synoptic table 
indicates these relationships just as forcibly as a diagram could do 
where there are so few elements involved. 

The genera are related, first, by the bilobed sternite, the lobes 
being capable of abduction; second, by the relation of the clave to the 
edeagus; third, by divergence having taken place along the line of 
degree of development in the ale. ) 

In Eleodimorpha the subale have been differentiated as distinet 
sclerites from the ale. 

In Eleodes and Embaphion the extrusion of the edeagus with 
simultaneous abduction of the clave, is brought about by muscular 
action—certain extrusor muscles; the return of these structures is 
accomplished by the inherent elasticity of the ale, the latter being 
the continuously produced and inflexed sides of the apicale. 

It is logical to suppose that accompanying the differentiation of 
the subale as distinct exoskeletal elements that there has been a 
corresponding muscular differentiation; for when the continuity of 
the ale underwent solution, they in all probability were no longer 
capable to functionate by their own inherent resiliency—of course, 
it is possible for the continuity of the elements to be restored by 
becoming connate at a sutural line; but this would evidently still be 
a weak point and correspondingly diminish the functional activity 
of the part. 


REVISION OF ELEODIINI—BLAISDELL. 31 


In all probability there has been a compensatory development of 
retractor muscles to overcome this loss of power. 


Genus ELEODES Eschscholtz. 


Mentum variable in form, more or less trilobed, the middle lobe 
larger and more convex; lateral lobes frequently inflexed and more 
or less concealed. Last joint of labial palpi triangular, narrower 
than the maxillary, which are broadly triangular or securiform. 

Head not deeply inserted, suture between epistoma and front fre- 
quently distinct; epistoma truncate or feebly emarginate, sides 
straight or sinuate and converging anteriorly. Eyes narrow, subreni- 
form. Antenne eleven jointed, the last three or four usually notably 
compressed, last joint frequently though not always smaller than the 
preceding. 

Prothorax variable in form, usually applied against the bases of 
the elytra. Scutellum always present, distinct and equilateral. 

Elytra of variable form and sculpture, in many species prolonged 
into a cauda behind; epipleure always distinct, broader at base and 
always visible at the humeral angles, gradually narrowing to apex. 

Legs moderately long, but somewhat variable in this respect; 
femora never strongly clavate, profemora most so, the mesofemora 
slightly, the profemora frequently armed in one or both sexes with a 
tooth of variable form, rarely all the femora armed; tarsi usually 
channelled and setose beneath, first and second joints of the protarsi 
frequently slightly thickened beneath and often the first two or three 
joints are clothed with a tuft of silken hairs, or dilated and densely 
covered with a spongy pubescence beneath. Spurs of the meso- and 
metatibie well developed, moderately slender, narrowing from base 
to apex, acute, the internal more or less longer than the external. 
Varying more or less in the different species. 

The above characters are those applicable to the whole genus; 
characters peculiar to certain groups will be given as of subgeneric 
value. 

The genus //eodes is one of the largest of the Tenebrionide and 
contains many more species than all of the other genera of the present 
tribe put together (United States). It is also the most polymor- 
phous and has been the most difficult to subdivide of any in our fauna. 
The study of the genitalia has greatly. reduced this difficulty, but 
extreme variations in many individuals of each species will yet be a 
source of great perplexity. 

The most salient extreme characters observed in the genus may now 
be reviewed : 

Color—Black, sometimes reddish along the elytral suture as in 
acuta, obsoleta, and hispilabris; rarely fusco-piceous as in longicollis 


32 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


and insularis, probably from immaturity, or from aging in the 
cabinet ; in one instance the elytra are describedgas castaneous (sub- 
cylindrica Casey) ; this unusual coloration is undoubtedly due to im- 
maturity. Dead specimens from long exposure to climatic inclemency 
become fuscous and even castaneous. In a limited number of species 
the legs are dark brown throughout, as in brunnipes Casey. 

Form.—Notably large and elongate as in the males of gigantea, 
oblong as in tricostata, elongately oblong as in acuta and texana, oval 
in veterator, inflated as in ventricosa and depressed in tibialis. 

Size —Largest in acuta, texana, and grandicollis; smallest in clavi- 
cornis. 

Surface—Glabrous and shining, as in dentipes, gigantea, and 
omissa,; opaque as in tricostata and opaca. 

Sculpturing—While usually simple it becomes tuberculate in gra- 
nosa and granulata; rugoso-muricate in cordata and muricate to a 
varying degree in many. 

Pubescence.—Body clothed with long flying hairs as in longipilosa, 
pilosa, and hirsuta, or confined to the elytra as in letcheri; hairs 
short, recumbent and abundant in veterator and opaca. 

Characters that vary within specific limits——These are the factors 
which make the limitations of species extremely difficult. Each 
species may have its extreme large (gigantism) and small (nanism) 
forms; its smooth and rough forms, elongate and robust forms, while 
the sculpturing varies from comparatively smooth to rough inde- 
pendently of size or form. This can be accounted for by environment 
to a great extent and to certain inherent factors. 

With robustness there is a relative shortening of the appendages 
and also of their segments, as for instance the joints of the antenney 
fromthe fourth to the seventh or eighth tend toward equality of 
length, while in the elongate forms there is a more or less inequality 
of length evident, the joints decreasing in length more or less from 
the fourth to the eighth, these joints being comparatively and rela- 
tively longer as a part of the general habitus. 

These remarks apply most emphatically to the tarsi and their 
joints as well; the antenne and legs as a whole exhibit this variation 
of length; the head varies in size and relative length and breadth 
even in any particular form. 

So it can be seen why in this particular genus (and this fact is 
worthy of broader application) that the relative length of parts of 
any appendage can not always be given as evidence of specific dis- 
tinctness. These conclusions have been arrived at after a long and 
tedious recourse to micrometrical treatment of hundreds of specimens. 

In many specimens, especially in the subgenus //eodes, there is to 
be observed a more or less evident smooth median pronotal line 


REVISION OF ELEODIINI—BLAISDELL. 33 


that may extend from the apical to the basal margins; this line is 
most pronounced as a rule in /. acuticauda var. laticollis, where it 
often becomes a distinctly impressed line; usually it is only observable 
as a line in surface luster. The significance of this character is 


obscure. 


Another very interesting character is that observed in the hind 
tibiae of the males of certain species of the subgenus /Jeodes. The 
metatibiz are slightly arcuate and narrow in basal half, rather ab- 
ruptly dilated and straight in apical half. In the literature I find 
this character mentioned by Doctor Horn as one of the male char- 
acters of £. wickhami. This is not a staple nor a specific character, 
but is observed as peculiar to the male of certain species, but not in 
all forms of those species; it is seen in the caudate forms of F. esch- 
scholtzi and luce, in armata and in the elongate form of dentipes; 
other males of the same species have the metatibie perfectly straight, 
some have them arcuate in basal half but not abruptly dilated 
apically. 

Individuals of many species exhibit what must be considered as a 
normal tibial arcuation, which must be distinguished from a common 
fortuitous arcuateness; the latter usually appears exaggerated and 
is generally a part of a general abnormal habitus, and it frequently 
accompanies unusual shortness of the legs. 


ANALYTICAL KEY TO SUBGENERA OF ELEODES, 


The genus Eleodes may be subdivided by clearly defined female genital char- 
acters into the following subgenera: 

Internal lobe of the valvular apex well developed, membranous, although fre- 

quently chitinous; external lobe obsolete; appendage directed backward 


NTE ey ae 2 i a Re Re nae oe ee eee ee eS tS 1 
External apical lobe well developed, always membranous; internal lobe feebly 
developed; appendage directed backward and inward________/ Velaneleodes. 
1. Superior pudendal membrane not broadly exposed_____________-_-_-_-_-__ 2 
Superior pudendal membrane broadly or fully exposed; body and apical 
lobe of valve fully chitinized; appendage minute ____________ Steneleodes. 
2. Appendages present, apical lobe defined from body of the valve____________3 


Appendages obsolete; apical lobe not defined from body of the valve. 
Heteropromus. 


3. Appendages moderate in size, subconical to semiellipsoidal and flattened: 
inferior margins of the valves not contiguous throughout their entire 


length but separated by a fusiform genital fissure___...__-__-_____-_ 4 
Appendages minute and punctiform; inferior margins of the valves con- 
tiguous throughout; genital fissure closed___________________- Discogenia. 

4. Apical lobe subacute, triangular, membranous or more or less semichit- 
ae See Rt eS Aa ae Rates ee 5 


597SO—Bull. 683—09——3. 


34 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Apical lobe rounded at apex, depressed, well chitinized, more or less concave 
beneath, convex above: 2 3 Jett oS ee eee Metablapylis. 
Dorsal plate variable in form, not broadened externall® finely, very sparsely 
punctate, more or less feebly and very sparsely pubescent___~--------__ 6 
Dorsal plate broadened externally, margin strongly arcuate, rather coarsely 
punctate and sparsely clothed with rather long flying hairs on outer 
moiety 2.2 a2 5 ee ee re Promus. 
6. Submarginal groove not visible from above; surface of dorsal plate at most 
but feebly concave, generally plane___-----_+.__--) >= > eee 7 
Submarginal groove dorsal or visible at some point from aboye; surface of 
dorsal plate more or less deeply concave_______-—_____-J5=_ == Bleodes. 
Superior outline of genital segment quadrate, triangulo-parabolic to para- 
bolie.. = ee ee ee ee 8 
Superior outline more or less triangular. 

Outline triangular; dorsal plate oblong-triangular, external border 
at most but slightly arcuate, internal border not strongly arcuate 
at. base. 228k 2 ean ee eee Blapylis. 

Outline subequilaterally triangular; external border of dorsal plate 
arcuate, angle obsolete; superior membrane reaching to middle 
of dorsal. plates. 222-53.) SS eee Tricheleodes. 

Superior outline of genital segment quadrate to triangulo-parabolic; superior 
membrane variable in length; dorsal plate oval-oblong, internal border re- 
flexed and impunctate; external border more or less arcuate, angle evident; 
apex small and membranous; appendage large, more or less semielliptical 
to subconical: (depressed). ===. 2. ee Eee Litheleodes. 

Superior outline parabolic, short; dorsal plate suboyate, extending upon the 
apical lobe beyond the appendage, sides not reflexed; appendage short, pro- 
jecting from beneath the outer border of the apical margin of dorsal plate; 
fenital fissure’ subapicales_- 325 24S Ses Se ee Pseudeleodes. 


on 


=i 


9 2) 


The student must bear in mind in resorting to the use of the above 
table that care must be taken to familiarize himself with the general 
structure of the genital segment. 


Besides the distinct generic differences there are in each subgenus 
specific characters which can only be worked out positively with large 
series, and, while of scientific interest, would not be of practical im- 
portance in the separation of species. 

The male genital characters may be summarized as follows: 


Apicale of the edeagophore broadly triangular, depressed on dor- 
sum and more or less broadly membranous centrally; apex 
more ‘or ‘less wounded! Shee oe eee ee ee eee 

Apicale triangular and slightly elongate, convex dorsally; apex more or less 
produced and subacute. 


Steneleodes. 
Metablapylis. 


Melaneleodes. 
Dorsum with an evident median, more or less membranous | Hleodes, 
groove. 6.062 eee ea ne Litheleodes. 
Promus. 


Dorsum without median groove, except in tibialis____-__------~~- Blapylis. 


: REVISION OF ELEODIINI—BLAISDELL. 35 


SYNOPSIS OF GENERAL SUBGENERIC CHARACTERS, 
Satine 
Anterior femora at least, armed in both sexes (except in caudifera and longi- “~~ . 
pilosa where the teeth are abortive) ----------____. Fis LES. : Bleodes. - 
Anterior femora armed only in the male or mutic. ° 
Anterior tibial spurs dissimilar in the sexes; femora mutic__J/elaneleodes.. 5.4% 
Anterior tibial spurs similar in the sexes. 7.40 oh 
Tarsi similar in the sexes, or nearly so. 
Middle lobe of the mentum large, lateral lobes rudimentary, i- 
visible without dissection; anterior tarsi with first joint more or _ 
less slightly thickened at tip beneath; anterior femora armed *" 
Wa, SPRL TN ROR EOS Felt fey 32 Ta P ais ey Fh ee ek Be tk Discogeniar * 
Middle lobe of the mentum small; anterior tarsi comparatively 
simple beneath, groove entire. ; 
Lateral lobes of the mentum fully exposed; sculpturing com-§ 77 


} aber 


paratively simple; femora mutic__ 2.27 __Metablapylis.( -..... 


‘ 


Lateral lobes moderately exposed; species opaque to shining; 
elytra tuberculate; anterior femora not sinuate, mutic. 


A. J¥6 Pseudeleodes. Frt*e<- 


7 


Anterior tarsi dissimilar in the sexes. 
Species pubescent throughout. 
Hairs long and flying; femora mutic; anterior tarsi with 
first joint scarcely produced ventro-apically, ventro-apical 
spinules noticeably produced in the female, not so in the ©. ,.. 


male; plantar grooves distinet______ ps 13% Tricheleodes. bev 


Hairs short, yellowish and recumbent; form oval, opaque, 


a 


ots 


Am 


ovis 


Po) 


hme 


5/ a 
RHO RT CRNO == 5 Ju_t79____Heteropromus, ¥etorace 


Species not usually pubescent, rarely so. 
Form elongate, usually large; first joint of the anterior tarsi 
slightly thickened at tip beneath, bearing a small trans- 
verse tuft of yellowish or brownish modified spinules which ra 


“4 
interrupt the groove in the male; simple in the female. an alae 


A. ¥09 Steneleodes.| srms 
Form ovate or fusiform; the first one or two joints of an- 


terior tarsi in the male more or less thickened and some- 


times flattened beneath, densely clothed with fine pubescence ; 2°" 


tarsi simple and unmodified in the female; anterior femora _4e7y* 


4 


ake 


. 2 a Fr. Le é 
SEPANCH (Gl TOMING (oe <8 eS ee 2 eS. fe Promus. } 4, sie 
: . ; ’ 
Form short ovate, moderate in size to small, robust (elon-) ¢{a °c. 


gate and depressed in tibialis); anterior tarsi of the male 
with first two or three joints feebly thickened at tip be- 


rer 
neath and clothed with dense silken or brownish pubescent | “°°. 


See 
tufts, obliterating the groove; joints simple with grooves «4 
entire in female; femora mutic________— 3 ~Aee___Blapylis.; ‘x 


Form ovate, moderate in size, less robust : first joint of the 
anterior tarsi more or less thickened and slightly more 
prominent ventrally than the others, pubescent tuft variable, 
most evident in ertricata; in the male first joint with a 


minute tuft of silken pubescence at tip beneath__Litheleodes. po Me be 


Az le, 


; ra 


36 BULLETIN 63, UNITED STATES NATIONAL MUSEUM, 
Subgenus MELANELEODES, new. 


Anterior spurs of the anterior tibiz dissimiliar in the sexes, and 
more or less curved. Longer and slightly thicker than the posterior 
in the male; notably longer, much broader and stouter and most 
strongly curved in the female. 

In the males of Aumeralis and rileyi the anterior spurs are much 
more strongly developed than in the same sex of the other species of 
the subgenus. 

The anterior femora are mutic and the first joint of the anterior 
tarsi is more or less thickened and feebly produced at tip beneath in 
both sexes. The male generally has a minute pencil of yellowish or 
brownish modified spinules on the produced tip; this is surrounded 
by a few closely placed, ordinary piceous spinules; in the female the 
produced tip bears only a few ordinary spinules. 

Subgeneric genital characters—Male.—Apicale of the edeagophore 
triangular, distinctly longer than wide, more or less convex above, 
with a more or less well marked semimembranous median groove 
apex more or less acute and rather attenuated ; base more or less lobed 
at middle. 

The membrane closing the interlobar sinus of the sternite is always 
more or less sparsely setose across the bottom of the sinus. This 
character is rarely observed in any other subgenus. 

Female.—Genital segment quadrate to trapezoidal. Each valve 
has the dorsal plate oblong and more or less distinctly explanate ex- 
ternally. The external lobe of apex is strongly developed, the inter- 
nal small and rudimentary. The appendage is mammilliform, rarely 
visible from above, and directed backward and inwards. 

Superior pudendal membrane is longitudinally rugulose, reaching 
apically to the base of the internal apical lobe. 

The greater development of the external lobe of the apex is the 
most distinctive subgeneric character and is not observed in any other 
division of the genus FJeodes. 

I can not at present devise a scheme by which the genital charac- 
ters can be used in the diagnosis of species; they apparently vary 
sufficiently to obscure the few constant specific characters. 

Variations—The individuals constituting the specific aggregations 
of the present subgenus are particularly prone to pronotal variations. 
This apparent instability is due to the degree to which the pronotal 
sides are arcuately deflexed, and a certain amount to the degree to 
which the extreme margin is carried downward and more or less 
inflexed against the propleure as it were. The actual marginal curve 
is quite constant in each species making up Group I. 


REVISION OF ELEODIINI—BLAISDELL. 37 


Morphologically the pronotum is widest at the middle in both 
sexes; but in the female there is a stronger lateral declivity, with a 
varying amount of lateral compression behind the middle to make 
the sides appear more or less straight from middle to base; con- 
versely, there is less deflexion anterior to the middle so that indi- 
viduals of the same species and sex may be more or less strongly 
arcuate before the middle. In Awmeralis there is less deflexion than 
in any other species of J/elaneleodes, the marginal bead being nearly 
always visible in the typical form when the pronotum is viewed 
vertically from above. 

The degree to which the angles are deflexed also gives a varying 
aspect to the pronotum. By a careful study of the different species 
in this respect the student will observe that really the pronotum is 
broadly, evenly, and rather strongly arcuate in the anterior two- 
thirds and thence to base more or less feebly sinuate. 

The above remarks apply to what I call a normal subgeneric varia- 
bility. 

Certain fortuitous deviations do occur which are not always easy 
to explain. The unique specimen described by me a number of years 
ago as interrupta is a case in point where environment has given evi- 
dence of lusus nature. 

The legs vary considerably in regard to length and stoutness of the 
different segments; sometimes these variations depend upon certain 
inherent traits of ancestral or developmental origin, constituting an 
abnormal condition in the relative proportions of the appendages 
and body. These evasive factors may affect any appendage or 
somitic part, and a student must bear these data in mind when deal- 
ing with the diagnosis of species or referring specimens to their place 
in the schema tawonomicum. 

On account of this variability within specific limits, I have given 
attention to the general relative proportions of parts under each 
species. I have been induced to do this from the extraordinary large 
series of specimens at hand. 

Distribution—The species of Melaneleodes inhabit the follow- 
ing States and countries: Texas, New Mexico, Arizona, Colorado, 
Nebraska, Kansas, Oklahoma, Montana; South Dakota, California, 
Lower California, Washington, Oregon, Idaho, Nevada, Utah, Indian 
Territory, British America (Medicine Hat, Riley) (?), Mexico (see 
humeralis and quadricollis) , lowa, Wyoming. 

Distributed through 19 States out of the 23 west of the Mississippi 
River. I have no record of specimens from Missouri, Arkansas, 
Minnesota, and Louisiana.- I expect to hear of their occurrence in 
one or more of those regions. 


38 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Diagram showing distribution. 


sees Orn te es ~ 

DEE: ee | é 

Name. Hiieie| |lalslo lela bela : 

e/=leiSl2\2/3/218(2/2/813|/slelalalale 

RIEFIS|S/SISZ(EISlElS(SlelS|Piglaiaieie 

a) o be = ° feed o Es ° a o i! & S = - 

RIZ/S/S ICIP IAlO|AIHIAIE/E/SO|ElAlAlalo 

| | 

CCDLIS! sctaisle\sie'n = ='=1e'r 4 lied Omen) Reno! eens eeies een) ero ree Mean seers eet apa SS) eg bee 
carbonaria.....---- x | X | X |----] x jovals catlepee|zcnlest elycel See Ac Ul er Pea bes | 
Mate CONOR aa kia Pawiv.<= x “|: ees ees ees fees ey sry eed rey Pa 
RUIN fawn oe Be lie. 9e0 (ee. eee Ree Ged eee Gene pee pees bere peerineee pee se ek. 
VAP IOOLOBUs ss os ss -> Be Ren Bie el een eee been es lenis bred bed breed heed ery lees eer) rey ee) bo) es 

OUSOLELG) =<. sues ane »| KM je sae|a cael cen| HC eciee|ecefyemelcoeel: Ka) Hy | WOK tS oS 
var. porcata......- meer ee KX |ncenleccalesee[sec-[one-[ooe]--n=]e<cc]-an-}ene «lane -laeee eee 
ITLOUILS TO erecta ste = Joe se] OX Jewnalpecafencs|ewccfemec|sces|stocs[ocetiecnc|ecas|s cea] Goce) eee) aaa ae 
ONUISSE) Rete oe me ser Bare) ed Pore bh eal ae | bes | Te be > ee (ey Pee Pe ae ere me) Ee) ke 
var. pygmxa.....-- bal etal ile al a meet ete a ee ee ee oe ee Se 
var. peninsularis...).... Ser, SR ee ere te eee eo be Ge rer) pes ae ees [sak on ee 
quadricollisa....... 2 Herlesec |e SseRect bee (B85) See, HES |S ee) SEA See eeh e Ree ee S eee eee Moe een ae 
IV ELasCUIR UU CLCUIROL wie sia) orm, wn! ota shel |G Mere eel oreo meester Bree een pend peed Ged boas eco eer cee 
var. lustrans ....-.. (aes PPO a oR ec 4 (D ~tas|ssaa|eece|ececlasc|ocecles 02] eee eaten 
cuneaticollis.......- Ee eee BASS Rae see Oa Roce IX Vicecloncc|sce-|eeec|tccs|eene}iaes a= 
humeralisb.......-. AEE) BS 5 Sale | Kb KX fesselocesfese.|oee fx | 1 ele ee 
TULCYURSD tok Gon Sener Ba Sel ai Dare is eerl Me 5h eR! Meee) CEM Be sceefae fe] 2 154 ce] a oirel eeteel ena et eee 
tricostata.....-.-..- Jet a Seletel Rh se cals a eee eee XK | K [soe] K fan-cflacoel KR | X | KM feee- 
pedinoides .......-. £7 SE) Led ii FR! ars sete A MIE PS Ws He TD le 9 ae) eee ES) Fees =) Se 
neomexicand ...---- ee es settee cet ees ee sey | eee [ee sefoese| apes bess le 

| 


“ Quadricollis—Sonora, Mexico. In the Biologia Cent. America (Vol. IV, Pt. 1, p. 
80) the author writes that he had seen specimens from Mexico. They probably belonged 
to a race or a closely related species. I have not seen an authentic example from Lower 
California. A specimen so given by Dr. George Horn—and examined by me—is probably 
a closely related species (Proc. Cal. Acad. Sciences, 2d ser., IV, Pt. 1, p. 350). The 
specimen was anatomically defective and has since been destroyed. 

» Humeralis.—Mexico ? In the Biologia (reference as above) the author writes: 
“There is an example of this species in Mr. F. Bates’s collection labeled as from Mexico; — 
possibly some mistake about the locality.” I agree with this. 

Dr. George Horn, in the Proce. Cal. Acad. Sciences, 2d ser., IV, Pt. 1, p. 350, gives this 
species as occurring at Sierra Laguna, San F'rancisquito, La Chuparosa, and San José del 
Cabo,.Lower California. I have studied these specimens in the collection of the academy 
and find them not humeralis at all. (See tibialis.) 


Genealogy.—In the accompanying diagram I have presented my 
present views regarding the descent and relationships of the species 
and races of the subgenus J/elaneleodes. 

From the subgeneric trunk divergence occurred along the lines of 
Groups I and II. 

In Group I the principal divergence is manifest in the calearine 
development as set forth for the Carbonaria and Quadricollis sec- 
tions. 

In the Quadricollis section the calearine modification is most 
strongly perpetuated, the specific differentiation resulting in the mod- 
ification of surface sculpturing, cuneaticollis being intermediate be- 
tween guadricollis and humeralis, but retaining greater affinity with 


REVISION OF ELEODIINI—BLAISDELL. 39 


the former; antennal modification resulted in racial divergence in 
quadricollis, while greater calcarine : Saar characterized the 
humeralis ramus. 

In the Carbonaria section crural development seems to have taken 
place, not affecting the main carbonaria stem where surface sculptur- 
ing has undergone modification along two lines, the smooth and the 
asperate as indicated in the synopsis of the species. 

Group I appears to be very plastic and reactive to modifying 
forces; Group ITI less so. In Group I there is evidence of greater 
specific and racial differentiations. 


Porcata 
Obsoleta 
matt Knausii Onrissa. 
Devils ou ra 
Jape 
Cuneaticollis 
Weak Humeralis 
Pedinoides 
aera Neomexicana 


Subgeneric Trunk 


Fie. 1.—GENEALOGICAL DIAGRAM OF THE SUBGENUS MELANELODES. 


The species of the present subgenus may be divided into two groups 
as follows: 


Elytra elongate oval, convex, comparatively narrow, feebly emarginate at base, 
humeral angles not rectangular; thorax not conspicuously broadened at base, 
angles obtuse, not overlapping humeral angles of elytra, more or less arcuately 
declivous at the sides, marginal bead rarely entirely visible when viewed 
Reptenl oa FONT DOVE ay. eke eb a eke Group lI. 

Elytra broad, flattened, truncate at base, humeral angles rectangular; thorax 
conspicuously broadened at base, angles rectangular, overlapping the humeral 
angles of the elytra; pronotum but slightly arcuately declivous at the sides, 
marginal bead distinctly visible from above_.__------------------ Group II. 


Group I may again be conveniently divided into two sections: 


Anterior spur of anterior tibix more or less gradually narrowed from base to 
RenpreIrT ED DOL ITT SOS OG! 65k ee Se ot eee Carbonaria Section. 
Anterior spur of anterior tibis more or less decidedly thickened in both sexes; 
gradually narrowing from base to apex in the male; much thickened and sub- 
parallel, narrowed more or less gradually beyond the middle third or basal 
half, subacute or obtusely rounded when worn__ ~~~ ~~~ Quadricollis Section, 


40 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


GROUP I.—CARBONARIA SECTION. 


Legs moderate in length'.2W2_ 2.) ee Eee 
Legs long; elytra rather more pointed behind 
Surface smooth, shining, punctures simple. _°___-__ 2 2_ = = eee 
Surface more strongly sculptured, punctures more or less muricate 
Smaller “species {= 2.324 te ae ee eee 
Larger -species — 2c ek ee ee eS ee 4 
Elytra with distinct and rather distant series of moderately sized punctures, 
with a single interstitial series of irregularly spaced, fine punctures (form 
Sthorlar “to \er(riea1a)_ 2 ee eee debilis, 
Elytra with distinct series of rather coarse puncturés, punctuation never 
diffuse. : 
Pronotum usually widest just before the middle (females at least). East 
of the ‘Colorado River=.-= 285 eet Eee carbonaria,. 
Pronotum quite quadrate, sides evenly rounded, equally narrowed before 
and behind (form somewhat similar to a smooth, finely punctate 
goryt}: TURRBR. 2.55 eee ee ane var. soror. 
Elytra usually more or less diffusely punctate, series frequently more or less 
evident, punctures generally subequal in size. West of the Colorado River. 
Larger size: 1 
Apical angles of thorax obtuse, apex feebly emarginate or truncate; 


won & e 


elytra. widest: at Iniddle::_._. .. 1-2» +. 2 ee eee omissa, 
Apical angles subacute, apex moderately emarginate; elytra usually 
widest behind the middle; surface polished______ var. peninsularis. 


Smaller size. Thorax more or less quadrate; males slender__var. pygmed. 
Elytra more or less subasperately sculptured; surface moderately shining, 
punctures irregularly, rather densely placed and submuricate, rarely sub- 
SETTLE soe Be Bo a ee Lg I ee knausii. 
Elytra muricately punctate; surface dull. 
Rather, robust. Elytra striato-punctate; sometimes estriate and dif- 


fusely punctate; intervals more or less moderately convex____obsoleta, 

More elongate. Elytra sulcate, intervals strongly Convex___var. porcatd, 
Surface more or less opaque througheut; pronotum broader _________~_ ampla, 
Surface more or less shining, pronotum quadrate and polished____var, dolosa, 


GROUP I.—QUADRICOLLIS SECTION, 


Anterior spur of the anterior tibiz, moderately produced in both sexes___-1 
Anterior spur of the anterior tibize, strongly produced in both sexes (less in 
the male) W..2e ese L eee eeeeeee 2 


. Elytra striato-punctate to diffusely punctate. 


Antenne moderate in length; surface more or less shining; epistoma 


coarsely and confluently punctate___._-.~._-__----=--==— quadricollis. 
Antennie long. Epistoma more finely and not confluently punctate. 

Larger species: Ssurtace @ulla) 2") === eee var. anthracina. 

Smaller species; surface polished and shining __----_~ var. lustrans. 


Elytra with punctures irregularly distributed, without serial arrangement, 
but producing a strongly rugulose appearance by mutual semicoales- 


PENCE Gio 2 ee ee See cuneaticollis, 
Elongate species___---~- =a ee eee bet ee 5 3 


Shorter, more robust species... =. i. ___ i ee eee 


REVISION OF ELEODIINI—BLAISDELL. 41 


3. Elytra opaque, densely muricately punctate (Washington); frequently 
granulato-muricately (Nevada and California) or tuberculo-muricately 


GAG Py Mc ano a Se ee ee humeralis. 
4, Elytra shining, subasparately punctate, punctures rather sparse, often with 
a feeble Hineal arrangement____.__--------_-_----_=--_~- ste ley. 


SEROUS NO at Sea See epics So ped to ee a 1 
RPO iN) Lie WeHRE *COMLTCC Ln 82 ot ete ee a ee 2 
1. Elytra with alternate interspaces more elevated, at times all quite evenly 
PMG INOS: DOUG: 60-8 te acres 2 Ce nb ere aces eee tricostata. 

2. Elytra glabrous and shining, with regular strie of moderately large 
punctures, interstices equally and very feebly convex ; pronotum with basal 

Re INEIERTLIIN rae ness acre Se SAP es a pedinoides. 
Elytra opaque, diffusely punctate; basal depressions of pronotum obso- 
(CDS I SRS a Se vip SERA ia ad eee eh eS eee SS var. neomericana, 


GROUP I.—CARBONARIA SECTION. 


ELEODES DEBILIS LeConte. 


Eleodes debilis—LErContTr, Proc. Nat. Sci. Acad. Phila., 1858, p. 185.— 
Horn, Trans. Amer. Philos. Soc., XIV, 1870, p. 808.—Casry, Annals 
N, Y. Acad. Sci., V, Nov., 1890, p. 396.- 

Ovate, moderately robust, glabrous and shining. 

Head twice as wide as long, feebly convex, rather coarsely punctate, 
punctures very sparse on the vertex, denser on the epistoma; surface 
more or less impressed laterally and along the frontal suture. Avn- 
tenne moderate, reaching to the base of the prothorax, subequal in 
the sexes, outer four joints very feebly compressed, scarcely dilated, 
third joint about equal to the next two taken together, fourth hardly 
longer than the fifth, the latter, sixth and seventh subequal, eighth 
feebly triangular, ninth and tenth suborbicular, scarcely transverse, 
eleventh ovate. 

Pronotum subquadrate, widest at the middle, about one-fifth to 
one-third wider than long; disc rather strongly and evenly convex 
from side to side, feebly so antero-posteriorly and more or less de- 
clivous at the apical angles, finely and sparsely punctulate, rather 
more strongly so laterally; apea feebly and evenly emarginate, usu- 
ally obsoletely margined; sides converging a little anteriorly, rather 
evenly and moderately arcuate in apical two-thirds, thence to base 
obliquely straight or feebly sinuate, finely margined; base rather 
broadly rounded and finely margined, about a third wider than the 
apex; apical angles subacute; basal angles obtuse. 

Propleurx smooth, sparsely and more or less obsoletely punctulate, 
slightly rugulose. 

Elytra oval, widest at the middle, about a third longer than wide; 
base broadly and very feebly emarginate, scarcely wider than the 


42 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


contiguous base of the prothorax; Auwmeri more or less acute: sides 
evenly arcuate, apex obtusely and narrowly roumded; disc evenly 
and moderately convex, laterally more strongly rounded, evenly and 
quite strongly arcuately declivous posteriorly, punctate, punctures 
slightly separated, moderate sized and arranged in rather distant 
series; intervals flat with a single series of irregularly spaced and 
usually fine punctures, both series becoming more or less confused 
at the sides and apex. 

Epipleure moderate in width, superior margin feebly sinuate be- 
neath the humeri, very gradually narrowing to apex; surface gla- 
brous and more or less finely and sparsely punctulate. 

Sterna more or less glabrous, irregularly and more or less densely 
punctate, rugulose. 

Parapleure rather coarsely and somewhat densely punctate. 

Abdomen shining and glabrous, sparsely, and finely punctulate, 
more or less rugulose. 

Legs moderate. Anterior femora mutic in the sexes; anterior tibial 
spurs slightly dissimilar; first joint of the anterior tarsi produced at 
tip beneath and feebly dissimilar in the sexes. 

Male——Slightly robust. Thorax quite quadrate; abdomen  sub- 
horizontal, first two segments more or less broadly impressed at 
middle, intercoxal process slightly concave; anterior tibial spurs 
comparatively small and slender, the anterior one scarcely widened. 
First joint of the anterior tarsi with a minute pencil of brownish 
modified spinules surrounded by dark spinules on produced tip be- 
neath. 

Female.—Moderately robust. Thorax slightly transverse. Abdo- 
men horizontal and rather strongly convex. Anterior tibial spurs a 
little larger and longer, the anterior spur slightly widened but grad- 
ually narrowed from base to apex. First joint of the anterior tarsi 
with dark spinules on the produced tip beneath. 

Measurements—M ales: Length, 12-13 mm.; width, 5.5 mm. Fe- 
males: Length, 12.8-14.8 mm.: width, 6-6.5 mm. 

Genital characters, male—Edeagophore elongate oval, acutely 
pointed at the apical extremity and slightly arched. 

Basale oblong, evenly convex, sides slightly arcuate. 

Apicale triangular, rather evenly convex above, with a narrow 
median membranous groove at the middle third; sides feebly arcuate 
at base, thence broadly and more or less strongly sinuate to apex, the 
latter appearing attenuated; base strongly and moderately narrowly 
arcuate at middle, laterally broadly sinuate. 

Sternite rather short and transversely parabolic. Each lobe with 
the external border evenly arcuate to apex, the latter narrowly 
rounded; internal border oblique; surface feebly convex, shining, 


REVISION OF ELEODIINI—BLAISDELL. 43 


very sparsely punctate and setose apically, setae moderately short. 
Sinus triangular, membrane slightly setose at bottom. 

Female.—Genital segment quadrate, valves reflexed at the sides, 
setose, glabrous and shining. 

Valvula (Plate 4, fig. 26).—Dorsal plate oblong, explanate exter- 
nally and slightly deflexed, internally reflexed and impunctate in 
apical half; surface convex externally, very sparsely punctate, punc- 
tures denser on convexity, each with a long seta; external margin 
straight; angle rectangular and prominent, sometimes feebly rounded, 
with a few moderately long set; apical margin squarely truncate to 
moderately and arcuately produced in outer half, more or less sin- 
uate over the appendage; internal margin more or less arcuate. Apex 
short, triangular and acute, set with a few moderately long sete. 

Appendage (cercopod) short and moderate in size, conical or feebly 
depressed, bearing moderately long sete. 

Basal prominences not evident. 

Superior pudendal membrane feebly and distinctly longitudinally 
rugulose, reaching to about the middle of the dorsal plate. 

Ventrolateral surface moderately convex, scarcely concave before 
the apex; submarginal groove broad and shallow beneath the ex- 
planate external border of dorsal plate; surface finely punctate and 
setose, sete minute. The internal borders of the valvule are con- 
tiguous in basal sixth; genital fissure fusiform and moderately wide, 
with the inferior pudendal membrane visible in basal half. 

Habitat—New Mexico (Santa Fé, C. V. Riley) ; Arizona (Sulphur 
Spring Valley, Hubbard and Schwarz); Texas (Alpine, elevation 
4,400-6,000 feet, H. F. Wickham). 

Number of specimens studied, 3 males, 6 females. 

Type in the LeConte collection. 

Type-locality.— New Mexico; collected by Mr. Fendler. 

Salient type-characters—Thorax subquadrate, slightly narrowed 
anteriorly; sides broadly rounded; apical angles acute, the basal ob- 
tuse; base broadly rounded. Elytra oval and convex, strongly de- 
clivous posteriorly, not acuminate at apex; base broadly emarginate ; 
humeri acute; dise quite strongly striato-punctate, intervals with a 
single series of fine punctules (LeConte). 

Diagnostic characters—In size and shape debilis resembles a 
smooth extricata, from which it differs in having the anterior femora 
unarmed in both sexes, and by the distinct and rather distant series 
of moderate punctures, with a single interstitial series of irregularly 
spaced fine punctures. 

The punctuation in debilis somewhat resembles that observed in 
carbonaria, from which it can be separated by its smaller size and 
female genital characters; the latter also separates it from all other 
members of the subgenus. 


44 BULLETIN 68, UNITED STATES NATIONAL MUSEUM. 


A specimen before me has been compared with the LeConte type 
and pronounced typical by Mr. F. Blanchard. ° 

This species has been considered as identical with carbonaria, and I 
agree with Col. Thomas Casey “ in considering it a distinct species. 

The genital characters are somewhat aberrant to the present sub- 
genus, but on account of the greater development of the external lobe 
of the valvular apex, I have placed it here; the apical margin of the 
dorsal plate is quite squarely truncate and the appendage more 
strongly exposed. In all specimens examined the female genital 
characters were constant. 

On account of the homogeneous structure exhibited by the small 
series before me, I am unable to indicate with any feeling of certainty, 
its relationships and descent. A large series illustrating the lines of 
variation is necessary for this purpose. I might tentatively suggest 
that debilis may tend toward the subgenus Promus. I would about 
as soon accept this postulate as to consider it identical with car- 
bonaria. 

The mentum” is moderate in size, and more or less triangulo- 
trapezoidal in outline; surface rather coarsely and somewhat densely 
punctate, laterally with rather broad, shallow, and subfoveate im- 
pressions, with a median longitudinal subeariniform convexity ; each 
puncture bears a short inconspicuous seta. 

The prosternum between the coxe is rather prominent ventrally. 
convex to nearly horizontal and mucronate behind. The mucro is 
variable in size, sometimes pointed and at others broad and obtusely 
rounded. 

The mesosternum is more or less arcuately oblique and concave. 

The intercoxal process of the first © abdominal segment is slightly 
transverse and equal in length to the post-coxal portion. The meta- 
sternal salient is about two-sevenths narrower than the abdominal 
process. 

In the male the second abdominal segment is equal in length to the 
post-coxal part of the first and about a third of its length longer 
than the third segment, the latter being about a fourth of its own 
length longer than the fourth segment. 

In the female the second segment is noticeably longer, being about 
a fourth of its own length longer than the post-coxal part of the first. 
The fourth is about half as long as the second, while the third is 
about a fourth of its own length longer than the fourth. 

The profemora are somewhat tumid externally, the tibial grooves 
are well developed, concave, and glabrous, margins subecariniform, 
converging and meeting at basal third. 

@VII. Coleopterological Notices, II, Oct. 6, 1890, p. 396. 
>The mentum refers to the middle lobe unless otherwise stated. 
© Tn all descriptive remarks only the visible segments are considered. 


REVISION OF ELEODIINI-——-BLAISDELL. 45 


The meso- and metafemora are each subequal in width, the former 
is scarcely compressed, the latter distinctly so. The tibial grooves 
have their margins feebly subcariniform and gradually converging 
to become contiguous and evanescent at basal thirds, 

The protibiz are feebly compressed ; all the tibize are without tarsal 
grooves, and muricately sculptured. 

The tarsi are moderate in length and comparatively slender. The 
protarsi are about a fourth of their length shorter than a mesotarsus; 
first four joints subequal in width; the second, third, and fourth are 
about as long as wide, the fourth slightly smaller than the third; the 
fifth is about as long as the preceding three taken together; the first 
is longer than wide, and rapidly narrowing to the base. 

The mesotarsi are about a fifth of their length shorter than a 
metatarsus. First four joints are subequal in width, gradually and 
slightly diminishing in length from the second to the fourth. Joints 
one and five are each separately subequal to the combined lengths of 
the second and third. 

The metatarsi are about two-fifths of their length shorter than their 
metatibize. The fourth joint is about equal to the combined lengths 
of the second and third; the first is a little longer than the same, and 
the second is just a little longer than the third. 

The comparative tarsal measurements were made on the female. 


ELEODES CARBONARIA (Say). 


Blaps carbonaria Say, Journ. Acad. Nat. Sci. Phila., IIT, 1823, p. 260. 

Eleodes carbonaria LeContr, Complete Writings of Thomas Say, II, 1859, 
p. 125; Proc. Acad. Nat. Sci. Phila., 1851, p. 181.—Horn, Trans. Amer. 
Phil. Soc., XIV, 1870, p. 308.—CHamprion, Biol. Centr.-Amer., IV, 
Pt. 1, 1884, p. 808.— Casey, Annals N. Y. Acad. Sci.. V, Nov., 1890, p. 395. 

Bleodes soror LEContr, Proc. Acad. Nat. Sci. Phila., 1858, p. 185. 

Eleodes immunis LeContr, Proc. Acad. Nat. Sci. Phila., 1858, p. 186.— 
Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 308. 

Oblong-ovate, moderately elongate, smooth, more or less shining, 
estriate, elytral punctures arranged in series. 

Head twice as wide as long, more or less convex, rather densely and 
irregularly punctate, punctures denser anteriorly, sparser about 
vertex, surface more or less impressed along the frontal suture and 
laterally. Antenne moderately long, reaching to or very slightly 
beyond the base of the prothorax, subequal in the sexes, outer four 
joints feebly compressed, just perceptibly dilated, third joint about 
equal to the next two taken together, fourth just. noticeably longer 
than the fifth, the latter, sixth and seventh subequal, eighth triangular 
and as wide as long; ninth and tenth more or less feebly transversely 
oval; eleventh ovate. . 

Pronotum subquadrate, widest at or a little in advance of the 
middle, and about one-fifth to one-third wider than long; disc evenly 


46 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


and moderately convex, more or less arcuately declivous laterally and 
at the apical angles, finely and sparsely punctulate,ppunctures slightly 
denser and coarser along the margins laterally; apex slightly emargi- 
nate or truncate, finely or more or less obsoletely margined; sides 
evenly and more or less moderately arcuate from base to apex, or 
more strongly arcuate in the anterior two-thirds, and thence to base 
oblique and convergent, or very feebly arcuate, sometimes obsoletely 
sinuate, finely beaded; base feebly arcuate, finely to more or less 
obsoletely margined, and one-fifth to one-fourth wider than the apex; 
apical angles distinct, scarcely at all rounded; basal angles obtuse. 

Propleurew finely and very sparsely submuricately punctate and 
more or less distinctly rugulose. 

Elytra oval, widest at the middle, less than twice as long as wide; 
base slightly emarginate, more or less feebly sinuate laterally, a little 
wider than or equal to the contiguous base of the prothorax: humerét 
subacute or feebly obtuse; sides evenly arcuate, apex obtuse; disc 
moderately convex on the dorsum, at times slightly depressed, later- 
ally quite strongly and evenly rounded, evenly and arcuately decli- 
vous posteriorly, serially punctate, strial punctures usually rather 
large and somewhat deeply impressed, at times rather small and not 
impressed, the individual punctures are usually more distant from 
each other than their own diameters; intervals flat, frequently feebly 
convex, with a single series of very small, distantly and irregularly 
spaced punctures; laterally the interstitial punctures become larger, 
and both series more or less irregular and somewhat confused. 

Epipleure gradually narrowing from base to apex, superior margin 
feebly sinuate beneath the humeri; surface finely to obsoletely or dis- 
tinctly and sparsely punctate. 

Sterna shining to subopaque, finely and rather densely punctate, 
at times rugulose. 

Parapleure rather coarsely and quite densely punctate. 

Abdomen more or less glabrous, finely and sparsely punctulate, 
first segment more strongly sculptured; surface obsoletely to dis- 
tinctly rugulose. 

Legs moderate. Anterior femora mutic in the sexes; anterior 
tibial spurs and tarsi slightly dissimilar in the sexes. 

Male——Body comparatively slender. Elytra rather gradually 
narrowed and evenly arcuato-obliquely declivous posteriorly. Ab- 
domen slightly oblique, very moderately convex, first two segments 
more or less impressed at the middle. Anterior spurs of the anterior 
tibiz slightly the stouter, almost twice as long as the posterior and 
gradually narrowing from base to apex, both comparatively slender 
and acute. Tarsi distinctly longer than in the female, the anterior 
with first joint bearing a minute pencil of brownish modified spinules, 
surrounded by ordinary piceous spinules on the thickened tip beneath. 


REVISION OF ELEODIINI—BLAISDELL. 47 


Female.—Body quite robust. Elytra rather broadly oval, rounded 
and more or less arcuately and vertically declivous posteriorly. Ab- 
domen horizontal, evenly and rather strongly convex. Anterior 
_spurs of the anterior tibie distinctly the stouter, more gradually 
narrowing from base to apex and about a third longer than the 
posterior. First joint of the anterior tarsi transversely clothed with 
ordinary spinules on thickened tip beneath. 

The material before me and referred to the present species is quite 
variable and presents a number of incipient races, of which three 
may be recognized to aid in the placing of specimens: 

Forma glabra.—Smooth and shining. Elytral punctures fine and 
not impressed, with a tendency to some irregularity on the disc, in- 
tervals flat. 

Forma typica.—Elytra with rows of rather large, impressed and 
distinetly defined punctures, intervals flat. 

Forma interstitialis——Elytra with very strongly impressed punc- 
tures, intervals feebly convex. 

The above forms or incipient races are connected by an abundance 
of mesotypes and when properly arranged make an_ instructive 
morphological series. 

Measurements —Males: Length, 15.8-18 mm.; width, 5.5-6.5 mm. 
Females: Length, 17-20.5 mm.; width, 7.8-9 mm. 

Genital characters, male-—Edeagophore rather elongate, fusiform, 
somewhat strongly arched and well chitinized. 

Basale oblong, rather strongly convex and more or less gibbous 
basally; sides evenly and rather moderately arcuate. 

Apicale triangular and feebly elongate; surface evenly convex, 
with a median longitudinal groove extending from near the apex to 
near the base, membranous in apical half, linear towards base: sides 
broadly and very feebly sinuate at middle third, apex subacute: base 
broadly and arcuately lobed at middle, very feebly sinuate laterally. 

Sternite parabolic in outline and slightly transverse. Each lobe 
with the external border more or less evenly arcuate, apex more or 
less evenly rounded, angle feebly evident at times; internal border 
short; surface rather evenly convex and glabrous, quite coarsely and 
densely punctate apically, more sparsely so towards base, setose, sete 
moderate in length, longer about apex, and not very dense. 

Sinus short, with the membrane very sparsely setose at bottom. 

Female.—Genital segment quadrato-trapezoidal, somewhat de- 
pressed and setose. 

Valvula (Plate 3, fig. 16).—Dorsal plate oblong, sometimes slightly 
widened at base, frequently deflexed apically, gradually explanate 
externally from basal third to apical margin with which it is directly 
continuous; surface plane, finely to rather coarsely and sparsely 
punctate, setose, setee rather short; external border feebly sinuous to 


48 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


scarcely arcuate; apical margin not defined from the external apical 
lobe. ~ 

Apex with the outer lobe well developed, the latter three-fourths 
as wide as the dorsal plate, rather broadly rounded and slightly semi- 
membranous at tip: internal lobe small and membranous; both lobes 
set with long flying set. 

Appendage short and more or less conical, sometimes slightly 
visible from above, directed backwards and slightly inwards. Fossa 
moderate and fringed with moderately long hairs. 

Basal prominences moderate. 

Superior pudendal membrane longitudinally rugulose and reach- 
ing to the side of the internal lobe of the apex. 

Ventrolateral surface rather short, not strongly convex, but more or 
less gradually sloping to the apex, not usually transversely impressed 
but sometimes more or less so just behind the narrow transverse basal 
convexity; submarginal groove well defined beneath the explanate 
external border of the dorsal plate, not markedly bending inwards 
at apex to fossa; surfaces finely and sparsely punctate, setose, sete 
very short, at apex short and flying. Internal margins of the val- 
vule contiguous for a very short distance both at base and apex; 
genital fissure broadly fusiform, with basal half closed by the inferior 
pudendal membrane. 

Remarks on female genital characters—Forma glabra has the 
genital segment quadrate in outline and more thickly setose at apex, 
with the basal convexities more strongly developed. Forma intersti- 
tialis has the dorsal plate more strongly and coarsely punctate. 

TTabitat—Forma typica.—Texas (Alpine, June, elevation 1,400- 
6,000 feet, H. F. Wickham; Uvalde, elevation 930 feet; Granjino, 
May: Marfa, elevation 4,600-4,800 feet, July, Wickham) ; New Mex- 
ico (Albuquerque, March, Wickham; Santa Fe Canyon, August, 
elevation, 7,000 feet, F. H. Snow); Arizona (Fort Grant; Oak 
Creek Canyon, Snow) ; Colorado (La Junta, Garland, Pleasant Val- 
ley, on the Arkansas River, Snow; Wickham’s List). 

Forma glabra.—Texas; New Mexico (Highrolls, May and August; 
Clouderoft, June, Warren Knaus); Colorado (Garland, June; 
Salida); Arizona (Chiricahua Mountains, May; Sulphur Springs 
Valley, June). 

Forma interstitialis——Texas (El Paso, July, elevation 3,700-3,800 
feet, Wickham) ; New Mexico (Deming, July, Hubbard and Schwarz; 
Luna; Albuquerque, March); Arizona (San Simon, July; Sulphur 
Spring Valley, May; Tucson, April; Chiricahua Mountains, May, 
Hubbard and Schwarz). 

Number of specimens studied, 100. 

Type destroyed, 


ee 


REVISION OF ELEODIINI—BLAISDELL. 49 


Type-locality as given by Thomas Say: “ Region of the Arkansas 
River and near the Rocky Mountains.” 

Salient ty pe-characters.—Thorax subquadrate ; sides regularly arcu- 
ate; apex very slightly emarginate; base nearly rectilinear, a little 
arcuate each side near the angles, the latter obtuse. Elytra without 
impressed striz; six regular series of punctures, which are more dis- 
tant from each other than the length of their diameters; interstitial 
lines with a few remote, acute, minute punctures; sides rounded and 
irregularly punctured (Say). 

Diagnostic characters—Easily recognized from the other members 
of the subgenus by the smooth polished surface, the elytra being 
sculptured with rather distantly placed series of coarse punctures; the 
intervals are flat, with a single series of small distantly spaced 
punctures. At times the punctures of both series may be equal in 
size, and the species is then with some difficulty separated from omissu 
(see p. 72 and forma glabra) ; typically the intervals are flat, but 
often become feebly convex and approach porcata (see p. 63 and 
forma interstitialis). The extreme heterotypical specimens are apt 
to have a subasperate punctuation, 

Immunis LeConte appears to be a true synonym, probably more 
closely related to var. soror. (See p. 50.) 

The mentum is moderate in size, triangulo-trapezoidal to parabolo- 
triangular in outline; surface rather broadly foveate laterally with 
a median longitudinal ridge, sometimes feebly sculptured, more or 
less strongly punctate and not noticeably bearded. 

The prosternum is variable; frequently continuously rounded be- 
tween the cox and behind, but not with the precoxal portion, not 
mucronate ; often almost horizontally produced, at other times convex 
between the coxe and with a more or less strongly developed mucro 
behind. 

The mesosternum at times is quite vertically declivous, at others 
oblique and more or less feebly concave. 

The metasternal process is as wide as the abdominal salient is long. 

The abdominal intercoxal process (male) is slightly transverse, 
a little wider than the metasternal salient, and equal to the post- 
coxal portion; the latter is also equal in length to the second seg- 
ment; the third segment is one-third of its length longer than the 
fourth. 

In the female the abdominal process is quadrate, and about a fifth 
of its width wider than, the metasternal salient, and equal in length 
to the second segment; the latter is about twice as long as the fourth; 
the third segment is equal in length to the post-coxal part of the 
first. 


59780—Bull. 683—09——4 


50 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


The profemora (male) are clavate, gradually tumid externally 
and subeylindrical in section; less so in the fengale and frequently 
somewhat compressed. The tibial grooves are more or less well 
defined by subcariniform margins which become contiguous and 
evanescent at basal fourth; the floors of the grooves are usually con- 
cave, glabrous or sometimes minutely subasperate. 

In both sexes the mesofemora are subtumid at middle, with the 
grooves rather narrow and limited by subasperulate margins, which 
are more or less evanescent before becoming contiguous at basal 
third. 

The sexes have the metafemora usually just noticeably widening 
from base to apex; grooves as on the mesofemora. 

The protibizw are frequently subarcuate, feebly compressed. All 
the tibiwe are without tarsal grooves and more or less muricate; the 
articular cavities are closed. 

The tarsi are moderate in length and stoutness. The protarsi are 
about a half of their length shorter than a mesotarsus. Joints two, 
three, and four are subequal, and combined subequal to the fifth; the 
first is longer than wide. 

The mesotarsi are subequal to (male) or a seventh of their length 
shorter than (female) a metatarsus. Joints one and five subequal in 
length, either one is subequal to the combined lengths of the third 
and fourth; joints two to four subequal in width and length, or just 
feebly diminishing in length in the order named. 

The metatarsi are about a third of their length shorter than a meta- 
tibia. The third joint is slightly shorter than the second, both com- 
bined about equal in length to the fourth; first joint equal to the 
combined lengths of joints two, three, and four of a mesotarsus. 


ELEODES CARBONARIA var. SOROR LeConte. 


Form as in carbonaria, smooth and more or less feebly shining, 
sides of the pronotum apparently evenly arcuate; elytral punctures 
usually moderately small. 

Head twice as wide as long, more or less moderately convex, some- 
what coarsely, irregularly, and sparsely punctate, punctures slightly 
denser on the epistoma; surface usually impressed laterally. An- 
tenne moderate in length, reaching to the base of the prothorax, sub- 
equal in the sexes, outer three joints scarcely compressed or dilated, 
third joint about equal to the next two taken together, the fourth 
very slightly longer than the fifth, the latter, sixth, and seventh sub- 
equal, eighth subtriangular, and slightly shorter than the seventh, 
ninth and tenth suborbicular, eleventh subovate. 

Pronotum quadrate, widest at the middle, where it is one-eighth 
to one-sixth wider than long, slightly narrowed before and behind; 
disc moderately convex, quite strongly and arcuately declivous along 


——_ 


REVISION OF ELEODIINI—-BLAISDELL. 51 


the lateral margins, finely and sparsely punctulate; apex subtruncate 
and more or less obsoletely margined; sides evenly, broadly and 
moderately arcuate from base to apex when viewed vertically from 
above, more strongly so in the anterior three-fourths when viewed 
obliquely from the side and obsoletely sinuate before the basal angles, 
margin finely beaded; base broadly arcuate, quite finely margined, 
and about one-fourth to one-third wider than the apex; apical angles 
more or less subacute; basal angles obtuse. 

Propleure smooth, finely and sparsely punctulate, more or less 
sparsely rugulose. 

Elytra oval, widest at the middle, rather strongly emarginate at 
the base and equal to or a little wider than the contiguous base of 
the prothorax; Aumeri acute and slightly prominent anteriorly: s/des 
evenly arcuate, apex obtuse and rather narrowly rounded; désc mod- 
erately convex on the dorsum, strongly and evenly rounded laterally, 
evenly and rather strongly arcuately declivous posteriorly, punctate, 
punctures moderate in size and arranged in moderately distant, 
scarcely impressed series; intervals more or less irregularly and 
sparsely punctulate, especially along the suture, laterally the series 
usually remain distinct. 

Epipleure rather wider than in carbonara, and gradually narrow- 
ing from base to apex; superior margin feebly and broadly sinuate 
beneath the humeri; surface smooth, sparsely and irregularly punc- 
tulate. 

Sterna shining, irregularly punctate and more or less rugulose. 

Parapleure irregularly and more or less closely punctate. 

Abdomen shining, evenly and sparsely punctate, usually obsoletely 
rugulose; intercoxal process more strongly sculptured. 

Legs moderate as in carbonaria. Anterior femora mutic in the 
sexes; anterior tibial spurs comparatively small and slightly dissimi- 
lar in the sexes, the anterior spur but slightly longer than the pos- 
terior. Anterior tarsi with the first joint feebly thickened at tip 
beneath and slightly dissimilar in the sexes. 

Male.—Body moderately slender, elongate-subovate. Elytra grad- 
ually narrowing, arcuately and somewhat obliquely declivous pos- 
teriorly. Abdomen slightly oblique, moderately convex, more or less 
broadly impressed at middle of the first two segments, intercoxal 
process distinctly concave. Anterior spur of the anterior tibiwe grad- 
ually narrowing from base to apex. First joint of the anterior tarsi 
with a small pencil of modified spinules surrounded by ordinary 
spinules on the thickened tip beneath. 

Female.—Body moderately robust. Elytra rather broadly oval, 
arcuately and somewhat vertically declivous posteriorly. Abdomen 
horizontal and somewhat strongly convex. Anterior spur of the 
anterior tibiw slightly thickened, acute, and gradually narrowing 


52 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


from base to apex. First joint of the anterior tarsi set with ordinary 
spinules on tip beneath. 

Measurements.— Males: Length, 16.5 mm.; width, 6.5 mm. 
Females: Length, 18.3-19 mm.; width, 8-8.5 mm. 

Genital characters——Male characters as in carbonaria. 

Female.—As in carbonaria, except that the dersal plate of each 
valve is narrower, with the sides more parallel and less explanate 
externally; the outer lobe of the apex is less broadly rounded. 

ITabitat.—Texas (Eagle Pass and San Antonio, LeConte; Browns- 
ville, Wickham.) 

Number of specimens studied, 8. 

Type, a female in the LeConte collection. 

Ty pe-locality.—Eagle Pass, Texas. 

Salient type-characters——Thorax quadrate, slightly narrowed an- 
teriorly and posteriorly ; sides broadly rounded; anterior angles acute, 
the posterior obtuse. Elytra strongly striato-punctate, intervals 
sparsely punctate (LeConte). 

Diagnostic characters ——On account of the form of the prothorax 
it somewhat resembles a smooth goryi, and LeConte differentiated 
it from that species as follows: “ The thorax is more quadrate and 
not more narrowed towards the apex than towards the base; the 
elytra are more deeply emarginate at base, and transversely much 
less convex, and those of the female are much less dilated; the pune- 
tures are more regular and never have the appearance of fovee.” 

All the specimens that I have received have either been labeled 
goryi ov unidentified, and it is undoubtedly confused with that spe- 
cies in collections generally. In all cases of doubt the subgeneric 
and genital characters must be carefully studied. 

The scuplturing of sovor is quite like that of a typical carbonaria, 
except that the serial punctures on the dorsum of the elytra are 
smaller, although as large or slightly larger and more distinctly de- 
fined laterally, the surface is feebly duller, the pronotum more quad- 
rate on account of the strongly deflexed sides of the disc and conse- 
quently less strongly rounded when viewed from above, and usually 
not more narrowed toward the apex than the base, but this character 
may be variable. 

The actual form of the pronotal margin in both sexes is like that 
of a male carbonaria, widest at the middle and not noticeably more 
strongly arcuate in the anterior moiety, as in the female of car- 
bonaria. 

From the members of the guadricollis section of the present sub- 
genus it is to be known by the form of the anterior spurs of the 
anterior tibie. 

I am indebted to Professor Wickham for the specimens in my 
collection. 


pm . 
ae 


_ i 


Se 


REVISION OF ELEODITINI—BLAISDELL. 538 


LeConte separated the species described by him as immunis from 
soror as follows: “ Of the same form as soror, but the thorax is more 
narrowed behind, almost as in £. guadricollis; the punctures of the 
rows of the elytra are sometimes placed in distinct grooves, while in 
other specimens the surface is even.” (See carbonaria.) 

Mr. Blanchard has pronounced my specimens undoubtedly the 
same as LeConte’s type from Eagle Pass, Texas. He also writes that 
the type and a male from “ Texas” have the elytra and under side 
shining, and two other females from * Texas” are dull. 

The mental, sternal, abdominal, and crural characters are as in 
carbonaria, except that the meso- and metafemora appear to be 
slightly more slender. The protarsi appear to be subequal in length 
in the sexes, and the mesotarsi appear relatively slightly shorter. 


ELEODES AMPLA, new species. 


Subfusiform to fusiform-ovate, subopaque, and smooth. 

Head twice as wide as long, more or less moderately convex between 
the eyes, rather broadly flattened anteriorly, usually feebly impressed 
laterally, sometimes across base of the epistoma; frontal suture dis- 
tinct and frequently bisinuate; evenly and sparsely punctate, punc- 
tures coarser on the epistoma and finer on the vertex. Antenne 
reaching a little beyond the base of the prothorax, very feebly com- 
pressed in outer four joints, scarcely dilated, third joint scarcely as 
long as the next two combined; the fourth, fifth, sixth, and seventh 
subequal in length; eighth triangular and as long as wide; ninth 
suborbicular; tenth slightly transversely oval; eleventh oval to ovate, 
and generally distinctly obliquely truncate at tip. 

Pronotum widest at or a little in advance of the middle, about one- 
fourth wider than long; moderately and evenly convex from side to 
side, feebly so antero-posteriorly, more or less arcuately declivous 
laterally along the sides; finely and sparsely or almost obsoletely 
punctulate; apex slightly and evenly emarginate, more or less obso- 
letely beaded; sides almost evenly arcuate from base to apex when 
viewed vertically from above, more strongly so in the anterior three- 
fourths, and obsoletely sinuate at basal fourth when viewed obliquely 
from the side, finely beaded; base quite broadly rounded and more or 
less feebly sinuate at middle, finely beaded, and about a fourth to a 
third wider than the apex; apical angles subacute; basal angles 
obtuse. 

Propleure rather smooth, finely and sparsely punctulate, usually 
distinctly rugulose. 

Elytra oval, less than one-half to about a third longer than wide; 
base more or less emarginate, frequently sinuate laterally, scarcely to 
a little wider than the contiguous prothoracic base; hwmeri subacute, 
searcely prominent anteriorly; sides evenly arcuate, apex obtuse and 


54 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


rather narrowly rounded; disc more or less moderately convex, some- 
times slightly flattened on the dorsum, more gtrongly and rather 
evenly rounded laterally, arcuately declivous posteriorly, punctate; 
the strial punctures are rather small to moderate in size and some- 
what closely placed in moderately distant series, usually not at all, 
but sometimes moderately impressed; intervals flat, rarely slightly 
convex. and with a single series of rather widely spaced, very fine 
punctures which at times become slightly irregular, especially later- 
ally and about apex. . 

Epipleurew moderate in width, gradually narrowing from base to 
apex, superior margin very broadly and slightly sinuate beneath the 
humeri; surface smooth, obsoletely to finely and sparsely punctulate. 

Sterna more or less shining, strongly punctate, and rugulose. 

Parapleure rather densely, finely, and subscabrously punctate. 

Abdomen usually shining, sparsely and quite regularly punctulate, 
more or less rugulose; intercoxal process and fifth segment more 
strongly sculptured. 

Legs long, comparatively moderately stout. Anterior femora mutic 
and the anterior tibial spurs moderately long, slightly dissimilar in 
the sexes. Anterior tarsi with the first joint feebly thickened at tip 
beneath. 

Male.—Body moderately slender and more or less fusiform, an- 
tenne reaching a short distance beyond the base of the prothorax. 
Elytra quite gradually narrowed in posterior fourth, arcuately and 
obliquely declivous. Abdomen distinctly oblique, moderately convex, 
first two segments more or less impressed at middle, process concave. 
Anterior spur of anterior tibiz about a third longer and a little 
stouter than the posterior, and narrowing moderately from base to 
apex. First joint of the anterior tarsi with the minute pencil of 
modified spinules on tip beneath scarcely evident and surrounded by 
the ordinary spinules. 

Female.—Robust. Antenne reaching just beyond the base of the 
prothorax. Elytra broadly oval, somewhat gradually narrowed and 
arcuately declivous posteriorly. Abdomen horizontal and rather 
strongly convex. Anterior spurs of the anterior tibize about a half 
longer than the posterior and feebly broadened, with sides rather 
slowly but evenly narrowing to apex, the latter acute. Anterior tarsi 
with ordinary spinules on the thickened tip beneath, the latter trans- 
verse and narrow. 

Measurements.—M ales: Length, 19-23 mm.; width, 6.5-8.5 mm. 
Females: Length, 23-25 mm.; width, 6.2-9.5 mm, 

Genital characters, male-——Edeagophore elongate, flaxseed-shaped 
(flattened, oblong-ovate) and more or less arched. 

Basale oblong; surface strongly convex towards base; sides feebly 
arcuate. 


= 


REVISION OF ELEODITNI—BLAISDELL. 55 


Apicale triangular and slightly longer than wide; surface evenly 
convex, with a narrow median groove in apical half; sides broadly 
sinuate at the middle, apex gradually narrowed and subacute; base 
broadly lobed and feebly sinuate laterally. 

Sternite transversely parabolic in outline. Each lobe subtriangular 
with the external border evenly and broadly arcuate, apex rounded 
but not narrowly; internal border rather short; surface evenly con- 
vex, glabrous at basal third, sparsely punctate and setose apically, 
setee moderate but longer at apical border. Membrane sparsely setose 
across the bottom of the sinus. 

Female.—Genital segment subquadrate, sometimes slightly deflexed 
at apex, setose. 

Valvula (Plate 4, fig. 27).—Dorsal plate oblong, moderately ex- 
planate externally; sides subparallel, and slightly sinous; surface 
nearly plane, slightly longitudinally concave, glabrous and im- 
punctate in basal third, elsewhere coarsely, sparsely, and irregularly 
punctate, setose, sete rather long and reclining; apical margin on 


external lobe strongly arcuate in outer three-fourths, internally 


sinuate. Both apical lobes set with long hairs; external lobe large, 
the internal very small. 

Appendage submammilliform and scarcely projecting beyond the 
fossa, hardly visible from above. Fossa rather large. 

Basal prominences scarcely evident. 

Superior pudendal membrane \ongitudinally rugulose and reach- 
ing about to the base of the internal lobe. 

Ventrolateral surface rather strongly convex at base, scarcely to 
feebly transversely concave before the apex; submarginal groove 
broad extending beneath the apex to the margin of the fossa. In- 
ternal margins of the valves contiguous at basal eighth; fissure 
broadly fusiform and closed in basal half by the inferior pudendal 
membrane. 

Habitat—Arizona (Oracle, July 15, Hubbard and Schwarz; 
Pinal Mountains, collection, University of Nebraska; Santa Rita 
Mountains, elevation 5,000-8,000 feet, July, F. H. Snow); New 
Mexico (Santa Fé Canyon, August, elevation 7,000 feet, I, H. Snow) ; 
Texas (Brownsville, June, Ff. H. Snow). 

Number of specimens studied, 13. 

Type in my own collection. 

Type-locality—Pinal Mountains, Arizona. 

Salient type-characters—Subopaque. Prothorax widest at (male) 
or in advance (female) of the middle. Elytra noticeably narrowing 
at apical fourth in both sexes; striato-punctate, stria not impressed, 
strial punctures rather small, the series rather distant, interstitial 
punctures very fine. Legs long. 


56 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Diagnostic characters.—In collections the females of the present 
species are usually associated with subnitens, while the males are 
placed with guadricollis or carbonaria. 

On account of its large size and dull integuments ampla has to be 
carefully differentiated from suvbnitens, from which it differs in the 
sides of the pronotum, being more strongly rounded and widest at or 
in advance of the middle, by the less prominent apical angles, and the 
subgeneric characters. 

Ampla bears no resemblance to a typical carbonaria; the dull luster, 
fine and unimpressed elytral punctuation readily separates it from 
the latter species, and besides the males are quite fusiform in outline 
and the legs are distinctly longer, characters never observed in car- 
bonaria. A male from Santa Fé canyon, New Mexico, has the strial 
punctures of the elytra moderately large and slightly impressed, but 
the form is elongate and fusiform. 

From guadricollis it differs by the anterior spurs of the anterior 
tibie of the females, being much narrower and evidently narrowed 
from base to apex, and by the long legs. 

Specimens from about Brownsville, Texas, are more shining than 
those observed from elsewhere. 

For characters separating it from its race dolosa, see below. 

In fact, ampla is the largest species in the present section of the 
subgenus. 

The mentum is very moderate for so large a species and varies quite 
a little in form—triangulo-trapezoidal to trapezoido-parabolic; the 
surface is moderately strongly punctate, and the setw are very small, 
laterally with shallow foveate impressions, rather broadly and longi- 
tudinally convex at the middle. 

The prosternum is variable, usually feebly convex between the coxe, 
rounded behind and very feebly mucronate at middle, or with a mod- 
erate mucro; sometimes horizontally produced, compressed, and ob- 
liquely truncate. 

Mesosternum more or less declivous and feebly concave. 

The metasternal process is about as wide as the abdominal salient 
is long. ‘ 

The abdominal intercoxal process is slightly transverse (male) or 
subquadrate (female), and about a fifth of its width wider than the 
metasternal process, and subequal in length to that of the post-coxal 
part of the same segment, the latter being quite equal to the second 
(male), the second segment a little longer (female) than the former. 
The third segment is about a third longer than the fourth. 

The profemora are feebly tumid to rather strongly so (male), less 
strongly and as variable in the female, in both sexes usually more or 
less slightly compressed, or subcylindrical, in transverse section in 
certain specimens (males) ; tibial grooves not strongly limited by the 


tt 


REVISION OF ELEODITNI—BLAISDELL. 57 


moderate subcariniform margins that converge, become contiguous, 
and then evanescent at basal fourth. The grooves are not notably 
concave. 

The mesofemora are moderately compressed, subfusiformly and 
not strongly tumid (male), or gradually wider to near apex in outer 
half (female) ; grooves not usually well defined, margins rather feeble. 

The metafemora are feebly widened outwardly with sides sub- 
parallel; grooves less defined, margins more or less asperulate and 
evanescent near the middle. 

The protibiw are scarcely compressed; all the tibize are without 
tarsal grooves. 

The tarsi are moderately long and comparatively stout. 

- The protarsi in the male are about a third longer than in the 
female. 

In the male the protarsi are about a fifth of their length shorter 
than a mesotarsus. Joints subequal in width, two, three, and four 
subequal in length and width, the fifth scarcely as long as the three 
preceding taken together; the first not as long as the two following 
combined. In the female the joints are relatively and proportionally 
the same. 

The mesotarsi (male) are a little shorter than a metatarsus. Joints 
one and five subequal in length; two and three are subequal in length, 
the fourth a little shorter. 

A metatarsus (male) is about a fourth of its length shorter than 
its metatibia. Joint one, three times as long as the third, the fourth 
quite equal to the combined lengths of the second and third, the latter 
two are subequal in length. In the female the joints are proportion- 
ally the same. 


ELEODES AMPLA var. DOLOSA, new. 


Elongate-ovate, more or less shining, pronotum polished. 

Head with the frontal suture obsolete or more or less well marked. 

Pronotum quadrate, widest at the middle in both sexes, and a little 
wider (5) than long; disc polished and shining, very finely and 
sparsely punctulate, punctures a little denser laterally; sides rather 
evenly and not strongly arcuate from apex to base, obsoletely sinuate 
at the basal fourth; base more or less finely and obsoletely beaded 
and about a fourth wider than the apex. 

Elytra with the dise striato-punctate, strial punctures feebly im- 
pressed, each puncture moderate in size and separated by about their 
own diameters; interstitial punctures small, distantly spaced, both 
series more or less irregular at the periphery. 

Legs long. Anterior tibial spurs moderately short. Otherwise as 
in ampla, 


58 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 
Male—Body moderately narrow, elongate-ovate. Elytra moder- 


ately narrowed in posterior fourth. Otherwise as in ampla. 

Female.—Moderately robust. Elytra less gradually narrowed in 
the posterior fourth. Otherwise as in ampla. . 

Measurements.—Males: Length, 18-21 mm.; width, 7-7.5 mm. 
Females: Length, 21 mm.; width, 9 mm. 

Genital characters, male-—Edeagophore scarcely at all arched. 

Basale elongate and suboval. 

Apicale elongately triangular ; surface extremely sparcely and finely 
punctulate. 

Sternite somewhat transverse. Each lobe triangulo-quadrate, with 
the external border broadly and feebly sinuate at basal three-fourths, 
thence arcuate to apex, the latter more or less rounded; internal 
border short and oblique. Otherwise as in ampla. 

Female.—Genital segment quadrate, 

Valvula.—Dorsal plate scarcely concave. 

Appendage very small. 

Ventrolateral surface quite broadly concave before the apex. 
Otherwise as in ampla. 

Habitat—Arizona (along the Colorado River, Beverly Letcher). 

Number of specimens studied, 7. 

Sexitypes in. my own collection. 

Type-locality—Western Arizona. 

Salient type-characters—More or less shining, elongate-ovate. 
Pronotum quadrate, polished, widest at the middle. Elytra striato- 
punctate, strial punctures not impressed (male) or feebly impressed 
and coarser (female). Legs long. 

Diagnostic characters.—In surface lustre and sculpturing resembles 
carbonaria; by its larger and more elongate form and long legs it is 
most closely related to ampla. 

Mr. Blanchard writes me that it is not to be referred to any species 
in the LeConte collection. It has heretofore been referred to car- 
bonaria, but the long legs will quickly separate it from that species. 

A specimen in Mr. Blanchard’s collection has the sides of the pro- 
notum more strongly arcuate. 

The mental, sternal, abdominal, and crural characters are practi- 
cally the same as in ampla. The anterior tarsi are apparently less 
elongate in the male, and in both sexes for that matter: in the female 
joints two, three, and four are comparatively a little smaller. 


ELEODES OBSOLETA (Say). 


Blaps obsoleta Say. Journ. Acad. Nat. Sci. Phila., III, 1823, p. 261.— 
LeContr, Complete Writings of Thomas Say, II, 1859, p. 155. 
Eleodes obsoleta Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 308. 
Eleodes obsoleta var. porcata Casey, Ann. New York Acad. Sci., V, Noy., 
1890, p, 396, 


EE — 


REVISION OF ELEODIINI—BLAISDELL. 59 


Oblong-ovate, slightly elongate, black, feebly shining to subopaque ; 
elytra striate and slightly scabrous, frequently reddish along the 
suture. 

Head twice as wide as long, moderately convex, frontal suture 
usually distinct, more or less feebly impressed laterally, rather finely 
and not very densely punctate, punctures slightly sparser on the ver- 
tex. Antenne moderate and subequal in the sexes, about reaching to 
the base of the prothorax, outer four joints feebly compressed, 
slightly and gradually dilated, third joint equal in length to the next 
two taken together, fourth just perceptibly longer than the fifth, 
the latter, sixth and seventh subequal, eighth slightly triangular, 
ninth and tenth irregularly orbicular in outline, eleventh oval and 
very slightly longer than wide. 

Pronotum widest at or slightly in advance of the middle, sub- 
quadrate one-fourth to one-third wider than long; disc moderately 
and quite evenly convex, more or less declivous at the sides, finely, 
usually irregularly but quite evenly punctate; apex feebly and evenly 
emarginate to subtruncate, finely to obsoletely beaded; sides nearly 
evenly arcuate from base to apex when viewed vertically from above, 
or more strongly so in the anterior three-fourths, thence oblique and 
more or less feebly sinuate to base, finely beaded; base more or less 
evenly and slightly rounded, finely margined and about a third wider 
than the apex; apical angles distinct and subacute; basal angles 
usually obtuse. 

Propleure finely, more or less irregularly and submuricately punc- 
tate, rugulose. 

Elytra oval, less than twice as long as wide, usually widest at about 
the middle; base feebly emarginate, scarcely to slightly wider than 
the contiguous prothoracic base; Awmeri subobtuse, scarcely at all 
prominent anteriorly; sides evenly arcuate, apex obtusely rounded; 
dise rather evenly convex, slightly flattened on the dorsum, laterally 
more strongly and rather evenly rounded, arcuately declivous pos- 
teriorly ; striate, rarely estriate or smooth, strize impressed and rather 
coarsely, uniseriately punctate near the suture to more irregularly so 
externally; intervals flat to moderately convex, sparsely and irregu- 
larly punctate, punctures more or less feebly scabrous; striz less de- 
fined and punctures more irregular and denser laterally. 

E'pipleure moderate in width, gradually narrowed from base to 
apex, superior margin beneath the humeri very feebly and broadly 
sinuate; surface finely and muricately punctate. 

Sterna finely to obsoletely punctate. 

Parapleure rather densely and finely punctate. 

Abdomen more or less polished, finely, sparsely punctulate, and 
more or less rugulose; intercoxal process more strongly sculptured. 


60 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Legs rather slender and moderate in length; anterior femora mutic 
in the sexes; anterior tibial spurs dissimilar; apterior tarsi feebly 
dissimilar in the sexes, first joint slightly thickened at tip beneath. 

Male.—Elongate, elytra rather gradually narrowed posteriorly, 
quite evenly and arcuately declivous behind. Abdomen moderately 
oblique, not strongly convex, more or less feebly impressed at middle 
on first two segments; intercoxal process somewhat concave. Anterior 
spurs of the anterior tibize about twice as long as the posterior, 
slightly curved, just feebly widened and gradually narrowed from 
base to apex, the latter acute. First joint of the anterior tarsi with 
the minute tuft of modified spinules scarcely evident, ordinary 
spinules present on the thickened tip beneath. 

Female-—Robust. Elytra somewhat broadly oval, and slightly 
narrowed posteriorly, usually arcuately and rather vertically decli- 
yous behind; abdomen horizontal and strongly convex. Anterior 
spurs of the anterior tibize usually about a third to a half longer than 
the posterior, moderately curved and rather gradually narrowed from 
base to apex, noticeably widened. First joint of the anterior tarsi 
with ordinary spinules on the thickened tip beneath. 

Four forms may be recognized : 

Forma glabra.—Elytra estriate and more or less smoothly seulp- 
tured. 

Forma typica. 
subasperate. 

Forma annectans.—Elytra rather strongly striato-punctate, in.er- 
vals strongly convex; sculpturing subasperate. 

Forma punctata.—Elytra estriate, irregularly and more or less 
muricately punctate. 

Measurements —Male: Length, 12.5-16 mm., width, 5-7 mm. Fe- 
male: Length, 14.5-18 mm.; width, 6.5-7.5 mm. 

Genital characters, male—Edeagophore (Plate 3, fig. 1) flax-seed- 
shaped (flattened oblong-ovate), scarcely arched. 

Basale oblong-oval, glabrous, moderately convex; sides more or 
less feebly arcuate. 

Apicale slightly elongate and triangular; surface in basal moiety 
evenly convex, in apical half a median longitudinal membranous 
groove; sides moderately arcuate in basal half, sinuate at middle, 
hence feebly or scarcely arcuate to apex, the latter acute but not 
acuminate; base broadly lobed at middle two-fourths, broadly and 
feebly sinuate laterally. 

Sternite (Plate 3, fig. 2) parabolic in outline. Each lobe with the 
external border more or less evenly arcuate from base to apex, fre- 
quently slightly sinuate in basal half; apex evenly rounded, angle 
usually not evident; internal border more or less oblique and feebly 
sinuous; surface rather sparsely punctate in apical moiety, setose, 


Elytra striato-punctate, intervals slightly convex, 


REVISION OF ELEODIINI—BLAISDELL. 61 


sete not dense and moderate in length, longer on apical border. 
Membrane very sparsely setose across the bottom of the sinus. 

Female.—Genital segment (Plate 3, fig. 3) quadrate, somewhat de- 
pressed and setose. 

Valvula—Dorsal plate oblong, explanate externally from base to 
apex, glabrous; surface more or less plane, never excavated, some- 
times with the apical portion slightly deflexed, sparsely punctate, 
sete moderate in length, longer apically; external margin feebly 
sinuate towards base, feebly arcuate towards apex, the latter with 
the external lobe strongly developed, three-fourths as wide as the 
dorsal plate and broadly rounded; internal border straight to feebly 
sinuous, the inner lobe of apex minute and separated from the ex- 
ternal by a small sinuation; apex set with long flying sete. 

Appendage minutely mammilliform, usually invisible from above ; 
fossa fringed with fine and rather long hairs. 

Basal prominences not conspicuous. 

Superior pudendal membrane longitudinally rugulose, reaching to 
the base of the internal apical lobe. 

Ventrolateral surface (Plate 3, fig. 4) with the basal swollen por- 
tion quite short, more or less broadly and transversely concave before 
the apex; surface glabrous, sparsely punctate and minutely setose; 
submarginal groove broad and shallow beneath the expanded sides, 
curving inward beneath the external lobe to the base of the internal 
lobe. Internal margins of the valves not contiguous at apex: genital 
fissure broadly fusiform, and nearly closed by the longitudinally 
rugulose inferior pudendal membrane. 

Habitat—Forma_ glabra.—Arizona (Williams, Barber, and 
Schwarz); Colorado (Arboles, C. F. Baker). 

Forma typica—Texas (Mobeetee, H. S. Barber): Nebraska 
(Grant, July, C. V. Riley; Pine Ridge, Sand Hills, Sioux City, col- 
lection University of Nebraska); Kansas (F. H. Snow): Colorado 
(Golden, E. J. Oslar; Colorado Springs, June, elevation 6,000-7,000 
feet, H. F. Wickham; Sterling, collection University of Nebraska; 
Platte Canyon, October, Greeley, H. Soltau; La Junta, Durango, 
South Park, San Luis Valley, Pueblo, West Cliff, Canyon City, Den- 
ver; Wickham’s List) ; Wyoming (Cheyenne, C. V. Riley and H. F. 
Wickham): Oklahoma (Fort Supply, October, H. S. Barber) ; Mon- 
tana (Assiniboine, Hubbard and Schwarz; Moose Jaw, August, 
A. N. Caudell; Helena); South Dakota (Badlands, Pine Ridge, 
September, L. Stejneger; Hot Springs, collection University of 
Nebraska). 

Forma annectans.—Colorado (Gulnare, Las Animas County, July 
8). 

Forma punctata.—New Mexico (Coolidge, Wickham); Colorado 
(Edith; Denver, collection of E. C. Van Dyke). 


62 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Number of specimens studied, 200. 

Type destroyed. “ 

Type-locality.—Say received his specimens from the “Arid plains 
of Arkansas and Missouri, in the vicinity of the Rocky Mts.” 

Salient type-characters—Body oblong-subovate, elytra with im- 
pressed striw, which are slightly scabrous with minute elevated points 
and impressed punctures; interstitial lines also punctured; sutural 
margin obsoletely reddish brown (Say). 

Diagnostic characters—This common species is to be recognized 
from the other members of the group by its duller integuments and 
striato-punctate elytra. In the typical form the elytra are feebly 
scabrous at the central part of the disc and more strongly so at the 
periphery; the strize are very obvious; in some specimens the striz 
are obsolete or feebly evident and the sculpturing slightly more 
scabrous (forma punctata), while in some there is a more or less dis- 
appearance of the asperate punctuation so that they are quite smooth 
and feebly shining and the striz are feebly marked (forma glabra) ; 
this form appears to be rare, as I have only seen four samples, and 
has no doubt heretofore been referred to quadricollis. 

The small and slightly more robust form, reddish along the elytral 
suture, is to be considered the typical form. Say’s measurements are 
one-half to three-fifths of an inch in length. This form in Colorado 
and elsewhere has very convex elytral intervals and is here spoken 
of as forma annectans, and is the homologue of porcata. 

The var. porcata is a larger and much more elongate form with 
strongly convex elytral intervals. (See p. 63.) 

The mentum is moderate in size and varies in form from triangulo- 
trepezoidal to trapezoido-parabolic; the surface is moderately pune- 
tate, setee minute, convex at middle, and more or less foveate laterally. 

The prosternum is rather short before the coxee and protuberant 
ventrally with the coxe, usually rounded antero-posteriorly between 
the same; in the small forms not mucronate, but in a few specimens 
there was a very slight mucro. In the larger individuals and in the 
punctate form the mucro becomes evident, but I have never seen it 
well developed. 

The mesosternum is at times feebly convex, never strongly concave, 
more or less oblique, but at times it is rather vertically declivous. 

The metasternal process is about as wide as the abdominal salient 
is long (male) or slightly narrower (female). 

The abdominal process is quadrate (male) or slightly transverse 
(female) and about a fourth of its width wider than the metasternal 
salient, equal in length to the post-coxal part of the first segment, the 
latter being equal in length to the second (male); the second is a 
little longer in the female, where it is a half longer than the third; 


i 


REVISION OF ELEODIINI-——BLAISDELL. 63 


in the male the third is a little longer than the fourth, and in the 
female a little less than twice as long as the fourth. 

The metasternum laterally between the coxw is as long as the width 
of a mesotibia at apical third. 

The femoral and tibial characters are practically as in ampla, ex- 
cept that the legs are much shorter. 

The tarsi are moderate in length and rather slender. 

The protarsi in the male are a little stouter than in the female; the 
difference in length is less marked, those of the male being just a 
little longer. 

The protarsi are about a third (male) or a little less (female) 
of their length shorter than a mesotarsus. Joints two, three, and four 
are subequal in size, each about as wide as long and together scarcely 
as long as the fifth; the first is a little longer than wide. 

The mesotarsi are about a third (male) or a sixth (female) shorter 
than a metatarsus. Joints two to four diminish just a little in length 
in the order named; the fifth is about equal to the combined length 
of the second and third, and subequal to the first. 

The metatarsi are about a fourth of their length shorter than a 
metatibia. Joint three is just noticeably a little shorter than the 
second, and the two together are subequal in length to the fourth, 
the latter subequal with the first. 


ELEODES OBSOLETA var. PORCATA Casey. 


Oblong-ovate, moderately elongate, black, elytra sulcate. 

Head twice as wide as long, frons slightly flattened to feebly con- 
vex, faintly impressed laterally, frontal suture usually finely im- 
pressed ; surface somewhat densely punctate, more coarsely so on the 
epistoma and finely and sparsely on the vertex; antenne moderate 
in length and stoutness, outer four joints very slightly compressed, 
more or less feebly dilated, third joint about equal to the next two 
taken together, the fourth slightly longer than the fifth, the latter, 
sixth, and seventh subequal, eighth feebly triangular, ninth and tenth 
circular in outline, eleventh oval to ovate and very slightly longer 
than wide. 

Pronotum subquadrate, one-fourth to one-third wider than long, 
widest in advance of the middle as viewed vertically from above, at 
the middle when viewed obliquely from the side; dise gabrous and 
shining or feebly alutaceous, moderately and evenly convex, more or 
less arcuately declivous laterally, finely and rather sparsely punctate, 
the punctures becoming a little coarser, but not very dense laterally: 
apex slightly emarginate in circular are, finely or obsoletely mar- 
gined; sides quite evenly and moderately arcuate in anterior two- 
thirds, thence less arcuate and oblique to base as viewed vertically 


64 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


from above, or more strongly arcuate at the middle, thence oblique 
and moderately convergent, feebly sinuate to basg as viewed obliquely 
from the side, finely beaded; base quite evenly and feebly rounded, 
finely to somewhat obsoletely margined and about one-fifth wider 
than the apex; apical angles distinct, not acute, frequently narrowly 
rounded; basal angles obtuse, sometimes more or less rounded. 

Propleurw finely, sparsely to rather densely submuricately punctate 
and more or less rugulose. 

Elytra oval, usually widest at about the middle, less than twice as 
long as wide; base broadly and feebly emarginate, as wide as or 
slightly wider than the contiguous prothoracic base; huwmeri more or 
less feebly prominent; s¢des evenly arcuate, apex not very narrowly 
rounded; disc quite evenly convex, occasionally slightly flattened on 
the dorsum, laterally rather evenly and strongly rounded, arcuately 
declivous posteriorly, deeply sulcate, the sulci finely, uniseriately to 
irregularly and muricately punctate; intervals about equal in width 
to the sulci, very convex, usually strongly defined, finely, sparsely, 
and irregularly punctate; inflexed sides feebly sulcate, intervals 
feebly convex, irregularly and rather more densely punctate. 

Epipleure moderate in width and gradually narrowing from base 
to apex, superior margin feebly and broadly sinuate beneath the 
humeri; surface finely, more or less sparsely, submuricately punctate. 

Sterna more or less finely, densely, submuricately punctate and 
rugulose. 

Parapleure finely and rather densely punctate. 

Abdomen somewhat shining, sparsely to more or less densely punc- 
tate, especially on the first and last segments, more or less rugulose. 

Legs moderate, comparatively rather longer than in *obsoleta. 
Anterior femora mutic in the sexes. Anterior tibial spurs dissimilar. 
Anterior tarsi dissimilar in the sexes, first joint slightly thickened at 
tip beneath, with a small tuft of spinules. 

Male.—Body somewhat slender. Elytra rather gradually nar- 
rowed posteriorly, evenly arcuately and somewhat obliquely declivous 
at apex. Abdomen slightly oblique, moderately convex and more or 
less broadly flattened at middle on first two segments, intercoxal 
process more or less concave. Anterior spurs of the anterior tibie 
about a half longer than the posterior, distinctly wider and gradually 
narrowing from base to apex. Anterior tarsi longer than in the 
female, first joint with a minute and inconspicuous tuft of modified 
spinules, surrounded by ordinary spinules on the thickened tip 
beneath. 

Female.—Body robust. Elytra rather broadly oval, rather rapidly 
narrowed, arcuately and vertically declivous posteriorly. Abdomen 
horizontal and strongly convex. ‘Anterior spurs of the anterior 
tibie about a third longer than the posterior, distinctly broadened, 


REVISION OF ELEODIINI—BLAISDELL. 65 


gradually but less rapidly narrowing from base to apex. First joint 
of the anterior tarsi with tuft of ordinary spinules at tip beneath. 

Measurements—Males: Length, 12-18 mm.; width, 6-7 mm. Fe- 
males: Length, 18.5-21 mm.; width, 7.5-9.5 mm. 

Genital characters, male——Edeagophore flaxseed-shaped (flattened 
eblong-ovate), slightly elongate and not usually arched. 

Basale oblong, glabrous, evenly convex; sides more or less parallel. 

Apicale triangular, slightly elongate, apical region more or less 
deflexed ; surface glabrous, rather strongly convex at middle, with a 
long median membranous groove reaching nearly to the base, groove 
linear in basal third; s?des broadly sinuate at middle third, slightly 
arcuate toward base; apex acute and rather gradually narrowed; 
base very broadly lobed at middle and very feebly sinuate laterally. 

Sternite slightly transverse. Each lobe with the external border 
feebly arcuate and oblique, becoming arcuate at apex, the latter 
broadly rounded or feebly subtruncate, angle scarcely evident; inter- 
nal border nearly straight; surface rather sparsely punctate in apical 
two-thirds, setose, setee moderately long on apical border, shorter 
toward base. Membrane sparsely setose across the bottom of the sinus, 
which is narrow and triangular. 

Female—Genital segment quadrate and setose. 

Valvula (Plate 3, fig. 14).—Dorsal plate somewhat broadly oblong, 
moderately explanate externally and at apex, frequently with a 
marked antero-posterior convexity; surface plane, sparsely punctate 
along the internal moiety and at apex, sete not long; external and 
internal borders more or less straight and quite parallel; apex with 
the external lobe broadly rounded and apparently semi-chitinous, the 
internal lobe small and membranous as usual and separated from the 
external by a small sinuation at junction of the first and second inner 
fourths of the apical margin of the dorsal plate. External lobe set 
with long flying sete, a few on the internal lobe. 

Appendage small, inferior, and more or less compresso-mammilli- 
form. Fossa rather large and fringed with sete. 

Basal prominences not conspicuous. 

Superior pudendal membrane as in obsoleta. 

Ventrolateral surface feebly convex toward base, broadly and shal- 
lowly concave before the apex, glabrous, finely and very sparsely 
punctate, sete minute, denser at apex: submarginal groove shallow 
and not well defined at the apex, not reaching the internal lobe. 
Internal margins of the valves contiguous at basal sixth. Genital 
fissure quite broadly fusiform. Inferior pudendal membrane regulose 
and largely visible. , 

It is to be noted that the sides of the dorsal plates are not as 
strongly explanate as in obsoleta. . 

59780—Bull. 683—09——5 


66 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Habitat—Arizona (Fort Apache, Casey; Galiuro Mountains, May; 
Fort Grant, July; Chiricahua Mountains, May, Hubbard and 
Schwarz; Williams, July; Ash Fork, June; Flagstaff, July, Barber 
and Schwarz: Kearn’s District, Navajo Indian Reservation, April 
A. W. Barber; Peach Springs, Walnut, Winslow, July; Seligman, 
July, H. F. Wickham; Prescott, H. C. Fall) ; New Mexico (Pecos, 
July). 

Number of specimens studied, 85. 

Type is in Col. Thomas Casey’s collection. 

Type-locality—Fort Apache, Ariz. 

Salient type-characters.—Prothorax with the dise evenly convex, 
finely, sparsely punctate, the punctures becoming rather coarse later- 
ally but not very dense; apex feebly emarginate; sides more strongly 
arcuate before the middle, thence moderately convergent and gradu- 
ally feebly sinuate to the basal angles, which are very obtuse but not 
distinctly rounded; base feebly and evenly arcuate. Elytra with 
the dise very deeply sulcate, the sulci finely, rather sparsely and 
muricately punctate, the intervals equal in width to the sulci, very 
convex, finely and sparsely punctate (Casey). 

Diagnostic characters——The strongly sulcate elytra separates this 
race from all others of the group. The individuals are unusually 
larger and more elongate than odsoleta. 

Well developed specimens appear quite distinct from obsoleta, but 
they mark a heterotypical variation which is connected to the typical 
form of obsoleta by an abundance of mesotypes of gll sizes and 
sculpturing, so that it can only be considered as a good variety. 
Some individuals are nearly as large as ampla, notably a series from 
Fort Grant, Arizona, collected by Hubbard and Schwarz. I have 
authentic examples before me kindly contributed to Mr. Charles 
Fuchs and myself by Colonel Casey. 

The mentum is moderate and varies in form from trapezoidal to 
trapezoido-parabolic; surface is rather strongly punctate, sete small, 
moderately convex and more or less strongly foveate laterally and 
narrowly impressed along the apical margin, the latter being fre- 
quently deflexed. 

In Casey’s types the prosternal process is prolonged and prominent. 
In the series before me it is very variable—from a small mucro to 
well developed, or compressed and subtruncate behind. 

The mesosternum varies relatively with the foregoing, concave to 
convex, slightly oblique to vertically declivous. 

The intercoxal process of the first abdominal segment is subquad- 
rate (male) or slightly transverse (female), and about a third of its 
width wider than the metasternal salient. 

The metasternum laterally between the coxe is as long as the width 
of a mesotibia at apex. 


— 


ES Ss shU et S”S— 


REVISION OF ELEODIINI—BLAISDELL. 67 


The post-coxal part of the first segment is equal in length to the 
process, equal in length to the second segment in the male, and the 
third segment in the female. ; 

In the male the third segment is a third of its length longer than 
the fourth; in the female the second is twice as long as the fourth. 

The legs are usually quite strongly sculptured. The profemora 
and the metafemora are as in carbonaria. The mesofemora are usu- 
ally, gradually and feebly widened from base to apex. 

The protibizw are subcylindrical in transverse section. All the 
tibia are without tarsal grooves and the articular cavities are closed. 

The tarsi are moderate in length, comparatively a little stouter 
than in obsoleta. 

A protarsus in the male is about a sixth of its length longer than in 
the female, and subequal in stoutness in the two sexes. 

The relative proportions of the tarsi to each other, and the con- 
stituent joints of each one to each other is practically the same as in 
obsoleta. 

ELEODES KNAUSII, new species. 


Oblong-ovate to ovate, more or less shining, estriate and moder- 
ately convex. 

Head twice as wide as long, feebly convex, frontal suture feebly 
marked, frons very slightly impressed laterally, somewhat coarsely, 
irregularly and more or less densely punctate, especially on the epis- 
toma. Anteyne moderate in length, outer four joints feebly com- 
pressed, distal three slightly dilated, third about as long as the next 
two taken together, fourth scarcely longer than the fifth, the latter, 
sixth and seventh subequal in size and about as wide as long, eighth 
shorter and apparently a little wider than long, ninth subtriangular 
and slightly transverse, tenth transversely oval, eleventh ovate. 

Pronotum subquadrate, widest just in advance of the middle when 
viewed vertically from above, at the middle when viewed obliquely 
from the side, and about a third wider than long; disc evenly and 
moderately convex, rather arcuately declivous at the sides and apical 
angles, quite finely, irregularly and somewhat sparsely punctate. 
punctures slightly denser at the sides; apea more or less feebly 
emarginate in circular arc, finely and more or less obsoletely beaded ; 
sides moderately arcuate in anterior three-fourths, thence obliquely 
convergent to base as viewed vertically from above, or more strongly 
arcuate anteriorly, thence convergent and more or less sinuate to 
base when viewed obliquely from the side, finely beaded; base feebly 
arenate, sometimes feebly sinuate at the middle, finely and more or 
less obsoletely beaded, One-fifth to one-third wider than the apex: 
apical angles distinct, not at all prominent anteriorly, sometimes very 
feebly and narrowly rounded; basal angles obtuse. 


68 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Propleurw obsoletely to finely, sparsely submuricately punctulate 
and rugulose. : 

Elytra oval, distinctly less than twice as long as wide, widest at the 
middle; base more or less emarginate, usually feebly sinuate each side 
of the middle, and generally slightly wider than the contiguous 
prothoracic base; Awmeri more or less exposed and obtuse; sides 
evenly arcuate, apex obtuse and rather narrowly rounded; disc mod- 
erately convex on the dorsum, quite evenly and strongly rounded at 
the sides, arcuately declivous posteriorly; almost coarsely, irregu- 
larly and rather densely punctate, the punctures are subequal in size 
and obsoletely muricate, frequenty there is a tendency to an arrange- 
ment into rather distant rows on the dorsum; laterally there is a 
shght tendency to rugulosity. 

Epipleure moderate in width and gradually narrowing from base 
to apex; superior margin feebly and broadly sinuate beneath the 
humeri; surface dull, finely and irregularly punctate. 

Sterna obsoletely to rather densely and finely punctate, more or less 
rugose. 

Parapleure finely and quite densely punctate. 

Abdomen finely, sparsely, and irregularly punctate, more or less 
rugulose, more coarsely sculptured on the first segment, the fifth 
coarsely punctate. 

Legs rather short and somewhat slender. more or less strongly 
sculptured. Anterior femora mutic; anterior tibial spurs compara- 
tively small and feebly dissimilar in the sexes. Anterior tarsi with 
the first joint slightly thickened at tip beneath and a little dissimilar 
in the sexes. 

Male—Oblong-ovate, moderately slender. Antenne reaching to 
the base of the prothorax. Elytra moderately narrowed in apical 
fourth, arcuately and scarcely obliquely declivous posteriorly. Ab- 
domen moderately oblique and convex, more or less broadly im- 
pressed at middle of the first two segments. Anterior spur of 
the anterior tibie about a half longer than the posterior, rather 
slender and gradually narrowing from base to apex, acute and 
feebly curved. First joint of the anterior tarsi with a small in- 
conspicuous pencil of modified spinules, surrounded by ordinary spin- 
ules on the thickened tip beneath. 

Female.—Ovate and robust. Antenne not quite reaching to the 
prothoracie base. Elytra feebly narrowed and arcuately, vertically 
declivous in apical fourth. Abdomen horizontal, strongly convex. 
Anterior spurs of the anterior tibie about a half longer than the 
posterior, slightly broadened, gradually narrowed from base to apex: 
both spurs rather strongly curved. First joint of the anterior tarsi 
with ordinary spinules on the thickened tip beneath. 


CO 


—_ 


REVISION OF ELEODIINI-——-BLAISDELL. 69 


Measurements.—Males: Length, 14.5 mm.; width,6 mm. Females: 
Length, 14-16 mm.; width, 7-8 mm. | 

Genital characters, male—Edeagophore subfusiform and rather 
elongate, not arched, 

Basale oval-oblong, evenly convex, with sides feebly arcuate. 

Apicale triangular, slightly elongate; surface evenly convex with a 
median membranous groove extending from near the tip almost to 
the base, where it becomes slightly less membranous; sides arcuate 
in basal half, thence more or less sinuate to apex, the latter very 
gradually and slightly decurved and very narrowly rounded. 

Sternite quite parabolic in outline. Each lobe with the external 
border quite evenly arcuate; apex rather narrowly rounded, angle 
not evident; internal border short, straight and oblique; surface 
slightly convex, glabrous, rather coarsely punctate in apical half,. 
setose, setee moderate in length; punctures denser and sete longer at 
apex. Membrane slightly setose at bottom of the sinus. 

Female—Genital segment quadrate, somewhat depressed, setose 
and not strongly chitinized. 

Valvula (Plate 4, fig. 24.)—Dorsal plate oblong and glabrous; 
surface plane, finely and sparsely punctate, with rather short reclin- 
ing sete arising from the same; external border straight or feebly 
sinuous, gradually explanate from the basal third to the apical 
margin, the latter rounded and explanately produced upon the ex- 
ternal lobe in outer three-fourths, directly and arcuately continuous 
with the external border; external apical lobe rather feebly mem- 
branous beneath and concealed from view from above; internal lobe 
moderate and separated from the external by a small sinuation in the 
apical margin; internal border straight or sinuous; apical region 
set with a few long, flying, soft sete. 

Appendage slightly visible from above, directed backward and 
feebly inward, conico-mammilliform. Fossa inferior, rather large, 
and fringed with long, flying sete. 

Superior pudendal membrane longitudinally rugulose and reaching 
to the base of the internal apical lobe. 

Basal prominences not noticeable. 

Ventrolateral surface not strongly convex in basal moiety, more or 
less concave laterally before the apex, but not transversely so; sub- 
marginal groove broad and shallow beneath the explanate external 
border of the dorsal plate and apical lobe, meeting the fossa at its 
external edge; external apical lobe not strongly developed beneath ; 
surface finely and irregularly punctulate and very finely setose. Inter- 
nal margins of the valves contiguous for a short distance at base and 
apex, between which the fissure is rather broadly fusiform, and closed 
in basal two-thirds by the inferior pudendal membrane, the latter 
feebly rugulose. 


70 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Habitat—New Mexico (Cloudcroft, James Canyon, June, Warren 
Knaus). 

Number of specimens studied, 8. 

Sexitypes in my own collection. 

Ty pe-locality—Cloudcroft, New Mexico. 

Collector, Warren Knaus. 

Salient ty pe-characters.—More or less shining, estriate and moder- 
ately convex. Pronotum subquadrate; disc rather finely, irregularly 
and sparsely punctate, punctures denser at the sides; apex very feebly 
emarginate; apical angles distinct and not at all prominent ante- 
riorly ; basal angles obtuse. 

Elytra with the humeri more or less exposed; disc almost coarsely, 
irregularly, and rather densely punctate, punctures subequal in size 
and obsoletely submuricate; laterally denser, with a slight tendency 
to rugulosity. Legs rather short and somewhat slender. 

Diagnostic characters.—Resembles the robust form of extricata or 
vileyé nv general outline. If it inhabited the region where parvicollis 
is found it might be mistaken for that species; some forms of leconte? 
resemble it. First of all, the genital characters distinguish it from 
all the above and associates it with obsoleta which it resembles, but 
less strongly in general habitus. It is usually confounded with 
extricata, from which it differs in the mutic anterior femora of the 
male, the more strongly arcuate sides of the pronotum at anterior 
two-fourths, and besides the punctuation is quite different. 

In vileyi the female has the anterior spurs of the anterior tibie 
strongly developed, while in knausii they are but feebly broadened. 
The spurs have greater development in the male of rileyi also than 
in the male of Anausii. 

In parvicollis and lecontei the lateral margins of the pronotum is 
distinctly visible from above, while in navsii the pronotal sides are 
arcuately declivous. The genital and tarsal characters are also dis- 
tinctive. 

Knausii differs from typical obsoleta in its smooth and shining sur- 
face and estriate elytra. It is also more robustly ovate than in the 
smooth form of obsoleta. 

I am indebted to Mr. Warren Knaus for this interesting species, 
and I take pleasure in dedicating it to him. 

Anausii appears to bear the same relation to the Carbonaria section 
that ri/ey? bears to the Quadricollis section. In this respect the two 
species are analogous. ; 

There is a specimen before me from Cananea, District of Arizpe, 
State of Sonora, Mexico, that resembles /:nausci in form, but it is more 
alutaceous and smoother, the striz of punctures are more evident and 
the interstitial punctures smaller and less conspicuous. The anterior 
tibial spurs are also feebly developed, and while the study of the 


REVISION OF ELEODITNI—BLAISDELL. 71 


United States specimens leaves the relationships in doubt, a know]- 
edge of the Mexican contingent would’ undoubtedly be more elucida- 
tive. At present /nausii appears as a derivative of obsoleta. 

The mentum is more or less trapezoido-parabolic, rather densely 
punctate and subfoveate laterally, somewhat convex at the middle; 
the sete are scarcely evident. 

The prosternum is moderately produced behind, feebly convex or 
horizontal between the cox and triangularly dilated behind the 
median transverse axis of the acetabula; the mucro is small, at times 
the process is compressed and more or less vertically truncate behind. 

The mesosternum is quite vertically declivous and more or less 
feebly concave. 

The intercoxal process of the abdomen is subquadrate (male) or 
slightly transverse (female), and about a fourth (male) or a third 
(female) of its width wider than the metasternal salient. 

The metasternum laterally between the coxe is as long as the width 
of a mesotibia at apical third. 

The post-coxal part of the first abdominal segment is subequal 
(male) or equal (female) in length to that of the process. 

In the male the second segment is equal to the length of the process; 
the third is about equal to the post-coxal part of the first; the fourth 
is about two-thirds as long as the second. 

In the female the second segment is about twice as long as the 
fourth, the third is equal to the post-coxal portion of the first. 

The legs are noticeably short. 

The profemora of the male are moderately clavate and feebly com- 
pressed; in the female feebly tumid and moderately compressed; the 
tibial grooves are limited by feeble but distinct subcariniform mar- 
gins which meet and become evanescent at the basal fourth. 

In both sexes the meso and metafemora are quite similar. The 
mesofemora have the surface lines feebly but distinctly arcuate, and 
the femora in outline may be said to be subfusiform. The meta- 
femora have the superior and inferior surface lines quite parallel. 
The grooves are limited by asperulate margins which converge at 
basal third on the mesofemora and become evanescent, while on the 
metafemora they become evanescent about the middle without be- 
coming contiguous. 

The tibizw are all more or less feebly arcuate, and rather more 
strongly widened apically than is usually observed; the tarsal grooves 
are absent and the articular cavities are closed. The protibiz are 
observed to be feebly compressed or subcylindrical. 

The tarsi are moderately long and rather stout when considered in 
a comparative sense. The protarsi of the male are about a fourth 
longer than those of the female. 


7] 


12 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


The protarsi are about a fourth (male) or two-thirds (female) of 
their length shorter than a mesotarsus. 

The mesotarsi are about a ninth—less in the female—of their 
length shorter than a metatarsus. 

The metatarsi are about a seventh (male) to two-fifths (female) 
of their own length shorter than their respective metatibia. 

The relative proportions of the joints of each tarsus are practically 
the same as in obsoleta. 


ELEODES OMISSA LeConte. 


Eleodes omissa LEContTrE, Proc. Acad. Nat. Sci. Phila., 1858, p. 186.—Horn, 
Trans. Amer. Philos. Soc., XIV, 1870, p. 308. 
Eleodes interrupta BLAISDELL, Ent. News, III, Dec. 1902, p. 241. 


Oblong-ovate to ovate, elongate, and more or less shining. 

‘Head twice as wide as long, more or less convex, feebly impressed 
laterally, frontal suture fine and usually evident, rather finely and 
not densely punctate, punctures sparser on vertex. Antenne 
moderate, outer four joints very feebly compressed, scarcely dilated, 
third joint equal in length to the next two taken together, fourth 
slightly longer than the fifth, the latter, sixth, and seventh subequal, 
eighth just the least shorter, subtriangular, and longer than wide, 
ninth and tenth somewhat circular in outline, eleventh ovate. 

Pronotum subquadrate, widest at or in front of the middle as 
viewed vertically from above, one-fourth to one-third wider than 
long; disc evenly and moderately convex, more or less arcuately 
declivous at the sides, finely, sparsely, and more or less irregularly 
punctulate, punctures just a little coarser and denser laterally; apex 
feebly emarginate or subtruncate in circular are, finely or obsoletely 
beaded; sides evenly and moderately arcuate in the anterior two- 
thirds, thence to the base more or less straight and converging 
(viewed vertically from above), or evenly and rather broadly arcuate, 
converging and more or less feebly sinuate (viewed obliquely from 
the side), finely beaded; dase slightly rounded to subtruncate, finely 
margined, one-ninth to a fourth wider than the apex; apical angles 
subacute to obtuse, rarely feebly prominent anteriorly; basal angles 
obtuse, frequently apparently rounded when viewed vertically from 
above. 

Propleure finely and very sparsely punctate, more or less rugulose, 
rarely rugose. 

Elytra oval, widest at the middle, less than twice as long as wide; 
base feebly emarginate to truncate, usually scarcely wider than the 
contiguous prothoracic base; hwmeri obtuse and not prominent; sides 
evenly arcuate, apex obtusely rounded; disc moderately convex on 
the dorsum, rather strongly and evenly rounded laterally, arcuately 
declivous posteriorly, punctate, punctures frequently of the same 


REVISION OF ELEODIINI—BLAISDELL. 73 


size, fine, diffusely arranged, not dense, generally some evidence of 
a serial order; the strial punctures are frequently slightly the larger, 
the series are moderately distant and rarely impressed. 

Epipleuvre moderate in width, gradually narrowing from base to 
apex, very feebly sinuate along the superior margin beneath the 
humeral region; surface finely and sparsely punctulate. 

Sterna more or less shining, rather densely punctate and more or 
less rugose. 

Parapleure quite densely punctate. 

Abdomen glabrous, finely and very sparsely punctulate, usually 
more or less distinctly rugulose—almost rugose at times. 

Legs moderate in length and stoutness, sometimes moderately 
thickened ; anterior femora mutic and the anterior tibial spurs slightly 
dissimilar in the sexes; anterior tarsi with the first joint slightly 
thickened at tip beneath and slightly dissimilar in the sexes. 

Male—Oblong-ovate, rather slender and elongate. Elytra grad- 
ually narrowing in the posterior third, arcuately and somewhat 
obliquely declivous posteriorly. Abdomen oblique, moderately con- 
vex, more or less flattened at middle of the first two segments. 
Anterior spurs of the anterior tibix slightly thickened, nearly twice 
as long as the posterior, both comparatively short. First joint of the 
anterior tarsi-set with a small tuft of spinules, the modified spinules 


scarcely evident. 
Female.—Ovate and robust. Elytra broadly oval, and somewhat 


gradually narrowing in the posterior fourth, arcuately and vertically 
declivous behind. Abdomen horizontal and strongly convex. An- 
terior tibial spurs rather variable in length, the anterior distinctly 
broadened and gradually narrowing from base to apex, sometimes 
with sides slightly arcuate in basal half, and about a third longer 
than the posterior, both acute, tapering, and stouter than in the male. 
First joint of the anterior tarsi set with a small transverse tuft of 
ordinary spinules on the thickened tip beneath. 

I have observed only four forms worthy of distinction in the hun- 
dreds of specimens before me of this variable species. 


Forma typica.—Thorax somewhat transverse, apex feebly emar- 
ginate or truncate, angles somewhat obtuse and not at all prominent 
anteriorly. Femora distinctly thickened. 

Male.—¥lytra rather more elongate than usual. 

Female.—Elytra with the sides more strongly arcuate anteriorly 
than usual. 

Forma cataline.—Pronotum more strongly punctate, especially 
laterally. Femora more or less thickened. 

Forma communis.—Thorax as in the typical form. Elytra with 
the sides less strongly and suddenly arcuate anteriorly. Femora not 
noticeably thickened. 


74 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Forma emarginata.—Thorax quadrate, apex distinctly emarginate, 
angles subacute and somewhat prominent anteriogly. 

Measurements.—M ales: Length, 16-19 mm.; width, 6-7 mm. Fe- 
males: Length, 18-23: mm.; width, 8-10 mm. 

Anomaly.—The specimen described by me* as interrupta belongs 
here. The prothorax has the side margin at middle rather abruptly 
interrupted. The elytra has the dise suleate posteriorly. Length, 
16.6 mm.; width, 7.1 mm. 

Genital characters, male-—Edeagophore (Plate 2, fig. 1) elongate- 
ovate to fusiform, and more or less arched. 

Basale feebly oblong-oval to oblong; surface evenly convex; sides 
feebly arcuate, sometimes converging slightly toward the apex. 

Apicale triangular, slightly elongate; surface moderately and 
evenly convex, with a median longitudinal grove in apical half, 
sometimes extending from apex to base, slightly dilated and mem- 
branous apically, gradually narrowing to become linear basally; 
sides moderately and evenly arcuate in basal half, thence rather 
feebly sinuate to apex, the latter subacute; base rather broadly lobed 
at middle and feebly sinuate laterally. 

Sternite (Plate 2, fig. 2) transversely parabolic. Each lobe more or 
less subparabolic in outline; external border straight to evenly 
arcuate, the apex being continuously but more narrowly rounded, 
angle sometimes evident; internal border quite short and feebly 
arcuate; surface scarcely convex, sparsely punctate and setose in 
apical three-fourths, sete rather long on the apical border. Mem- 
brane distinctly and sparsely setose at the bottom of the sinus, the 
latter short. 

Female.—Genital segment (Plate 2, fig. 3) quadrate, moderately 
explanate laterally and setose. 

Valvula—bDorsal plate oblong, moderately narrow, sides subpar- 
allel; surface plane, glabrous, sparsely punctate, impunctate at basal 
third, setose, setee moderate and reclining; external border straight to 
feebly arcuate, continuously so with the more strongly and evenly 
rounded apical border, which is more or less inwardly oblique; in- 
ternal lobe short and separated from the external by a small sinua- 
tion; external lobe set with rather long sete, the internal with a 
few rather short ones; internal border slightly sinuous. 

Appendage just visible from above, short mammilliform, with a 
pencil of sete at tip. Fossa moderate and in the internal wall of 
the external apical lobe. 

Basal prominences scarcely evident. 

Superior pudendal membrane longitudinally rugulose, reaching 
about to the internal lobe of the apex. 


a4—Ent. News for Dec., 1902. 


—- ache —_ 


REVISION OF ELEODIINI—BLAISDELL. 75 


Ventrolateral surface (Plate 2, fig. 4) moderately convex, slightly 
concave laterally before the apex; surface sparsely punctate and finely 
setose; submarginal groove well developed beneath the explanate ex- 
ternal border of the dorsal plate, passing obliquely inward beneath 
apex to the inner angle of the fossa, ending at the internal apical 
lobe. Internal margins of the valves contiguous at basal sixth; geni- 
tal fissure quite broadly fusiform, and closed at basal half by the 
inferior pudendal membrane. 

Habitat—Southern California; Santa Catalina Island. 

Number of specimens studied, 500. 

Type is in the LeConte collection. 

Type-locality.—San Diego, California. 

Salient type-characters.—Longer than usual. Thorax slightly con- 
vex, sides broadly rounded, somewhat narrowed posteriorly, finely 
and sparsely punctulate. Elytra declivous and obliquely narrowed 
behind, sparsely, finely, and very irregularly punctate. Femora un- 
usually thick (LeConte). 

Diagnostic characters.—The most common species in Southern Cali- 
fornia and always heretofore referred to qguadricollis, from which it 
differs in the smaller and more slender anterior spurs of the anterior 
tibie (females) ; from carbonaria, to which it is most closely related, 
by the punctures of the elytral disc having a less distinct serial ar- 
rangement, and by the serial punctures being scarcely larger than 
those of the intervals, the tendency being toward diffuse pune- 
tuation. 

Omissa is with greater difficulty separated from the smooth form 
of carbonaria, where the elytra punctures are small and of the same 
size; here locality must be relied upon for their separation. All 
those specimens from the eastern side of the Colorado River are 
forms of carbonaria, while all those from the western side are to be 
referred to the present species. 

I do not know of a single specimen taken west of the Colorado 
River ever having been identified as an authentic carbonaria. In 
fact I consider omissa as a western modification of carbonaria. 

Omissa is a variable species in regard to the form of the pronotum, 
but in the several hundreds which I have studied I found an abun- 
dance of all necessary mesotypes, both as regards the form of the 
pronotum and elytral punctuation. 

In a large percentage of those individuals that have the sides of 
the pronotum evenly arcuate from apex to base, it will be found 
that they really present a more normal form of the side margin than 
do those that are widest in front of the middle; for here it will be 
seen that the sides of the pronotal dise are most strongly declivous 
behind the middle and that the true margin is displaced downward 
and not visible when the pronotum is viewed vertically from above ; 


76 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


when viewed obliquely from the side the normal marginal curve will 
be seen and that in reality the pronotum is widest at the middle. 

It must also be borne in mind that the pronotal dise is normally 
moderately declivous laterally in the present subgenus. 

In many specimens (mostly male, but some females as well) the 
pronotal sides are less strongly deflexed and appear quite evenly 
rounded from apex to base, and as a result the pronotum is broader 
as compared to those with strongly deflexed sides where the thorax 
is less rounded and more narrowed to the base. ; 

Although the punctuation is variable, usually there is always some 
evidence of a serial arrangement; rarely, the strial punctures are 
rather large and even moderately impressed; these specimens ap- 
proach carbonaria in this respect, but oméssa is as a rule more elongate 
in the male and more broadly ovate in the female. 

The strongly punctate individuals are not. necessarily confined to 
the region bounding the Colorado River on the west, but are to be 
taken at any part of the distributional area. 

The larger Southern California specimens are to be referred to 
omissa, as the characters exhibited by the anterior tibial spurs are 
those characteristic of the present section of the subgenus; with these 
conceptions of oméssa and the elimination of quadricollis everything 
becomes clear. 

Ampla is a larger, more elongate species with elongate legs. 

For the differential characters of the varieties pygme@a and penin- 
sularis see below. 

I am indebted to Prof. H. C. Fall for notes on comparisons of 
specimens with the LeConte type. 

The mentum is variable, rather small to moderate in size, “na tri- 
angulo-trapezoidal to trapezoido-parabolic; surface rather strongly 
punctate, more or less foveate laterally and convex at middle. 

The prosternum is convex or horizontal between the coxe and 
mucronate behind; the mucro may be small and subacute; larger and 
conical in contour, or compressed and obliquely to nearly vertically 
truncate behind. 

The mesosternum is more or less concave, and varies in the degree 
of its obliquity. 

The intercoxal process of the abdomen is quadrate (male) or a 
little transverse (female), and about a fifth (male) or a sixth (fe- 
male) of its width wider than the metasternal salient. 

The metasternum laterally between the coxe is as long as the width 
of a mesotibia at the middle. 

The post-coxal part of the first abdominal segment is equal (male) 
or subequal (female) in length to that of the process. 

In the male the second segment is about a half longer than the 
third and twice as long as the fourth, and equal to the length of the 
process. 


REVISION OF ELEODITINI—BLAISDELL. 77 


In the female the second segment is about a third of its length 
longer than the third and twice the length of the fourth. 

The profemora in the typical form are distinctly tumid and clavate, 
and in the males of the other forms and all females more or less mod- 
erately so. The grooves are rather broad and the margins are feebly 
subcariniform, converging quite close to the base. 

The mesofemora are gradually and feebly thickened externally ; 
the metafemora have their superior and inferior borders subparallel. 
The grooves are dull and feebly subasperate, the margins finely 
asperulate and more or less evanescent before becoming contiguous. 

The protibiw are more or less feebly arcurate and slightly com- 
pressed. All the tibiz are usually without tarsal grooves, although 
an occasional specimen exhibits rudimentary grooves on the protibiz 
with the articular cavities slightly open, otherwise the cavities are 
closed. 

The tarsi are moderate both as to length and stoutness. 

The protarsi in the male are just noticeably longer than in the 
female, and specimens of corresponding size must be selected, as a 
large female compared with a small male would not show the true 
difference. 

The protarsi are about two-fifths (male) or three-sevenths (female) 
of their length shorter than a mesotarsus. Joints two, three, and four 
are subequal, just noticeably smaller in the female, and together about 
equal to the fifth in length; the first is about equal to the combined 
lengths of joints three and four. 

The mesotarsi are about a seventh (male) or an eighth (female) 
of their length shorter than a metatarsus. Joints one and five are 
subequal in length, and the combined lengths of two, three, and four 
just a little longer than either. 

The metatarsi are about three-fourths (male) or a half (female) 
of their length shorter than a metatibia. Joint two is a little longer 
than three; the first is as long as the combined length of third and 
fourth. 


ELEODES OMISSA var. PYGM/A, new. 


Oblong-ovate to ovate, more or less shining and smooth. 

Head finely punctate, punctures denser on the epistoma, frons 
feebly convex to flat. Antenne moderate, outer four joints scarcely 
dilated, fourth, fifth, sixth, seventh, and eighth subequal in length, 
ninth and tenth suborbicular and slightly transverse. 

Pronotum quite quadrate, widest at or just in front of the middle, 
one-fifth to one-fourth wider than long; disc evenly and moderately 
convex from side to side, finely and sparsely punctate; apex truncate 
in circular arc; sides evenly and not strongly arcuate in anterior two- 
thirds, thence more or less straight and converging to base, or some- 


78 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


times apparently quite evenly arcuate from apex to base, the latter 
quite truncate, equal to the length, a seventh toga fourth wider than 
the apex; apical angles obtuse, not in the least prominent anteriorly, 
sometimes feebly rounded ; basal angles obtuse. 

Elytra oval, quite evenly and more or less moderately convex on the 
dorsum, more strongly rounded laterally; ase scarcely emarginate, 
equal to or shghtly wider than the base of the prothorax; humeré 
obtuse, scarcely prominent or rounded; sides evenly arcuate, apex 
obtusely rounded ; disc finely and diffusely punctate, a serial arrange- 
ment usually evident, punctures generally of the same size, serial 
punctures sometimes slightly larger. Otherwise as in oméssa. 

Male—wUsually quite slender and oblong-oval, frequently more 
ovate, about three times longer than wide. Elytra scarcely wider 
than the prothorax in the typical form, at other times wider. Both 
sexes otherwise as in omissa. . 

Measurements—Males: Length, 13-15 mm.; width, 4.5-5.2 mm. 
Females: Length, 13-16.5 mm.; width, 6-7 mm. 

Genital characters as in omissa (Plate 2, figs. 1, 9, and 10.) 

Habitat—Southern California (San Diego to Kaweah, Tulare 
County). 

Number of specimens studied, 1,000. 

Sexitypes in my own collection. 

Type-locality—San Diego, Cal. 

Salient type-characters—Much smaller than omissa. Thorax quite 
quadrate. Elytra at the widest point scarcely wider than the pro- 
thorax in the male. 

Diagnostic characters —Typical specimens are much smaller than 
omissa, the larger individuals becoming indistinguishable from it. 

Pygmea is of especial interest from the fact that it has been 
generally distributed as gentilis. 

The female type before me was identified as gentilis by Colonel 
Casey, and is the species referred to by him in his Coleopterological 
Notices, VII,* where he writes “that a very large series of gentilis 
which I took at San Diego shows quite clearly that this species should 
be associated with guadricollis and vicinus, and is out of place in the 
present arrangement.” 

Gentilis is a different species altogether and is a race of gigantea. 

Pygmea is the most abundant species about San Diego, where I 
have seen it so abundant beneath boards, tin cans, sacks, and cow- 
chips, that they actually carried these things about over the ground. 
These remarks apply to the early “ seventies,” when San Diego con- 
sisted of scattered houses and the saline flats, covered by ice plants 
(Mesembryanthemum crystallinum) formed the greater part of the 


“Annals N. ¥. Acad, Sci., V, Nov., 1890, p. 395. 


ie sent igen 


REVISION OF ELEODIINI-—BLAISDELL. 79 


maritime landscape, then these insects were gregarious in untold 
hundreds. I have never found them so plentiful in recent years. 

Specimens from Tulare and Kern counties represent a slightly 
different incipient race, which may be characterized as follows: 

Forma borealis.—Males less oblong and a little more ovate. 

T am indebted to Mr. Ralph Hopping for the privilege of studying 
a large series. : 

In Professor Fall’s collection there is a female that is more opaque 
than usual, but otherwise it does not differ to any extent; it was col- 
lected in the San Bernardino Mountains, California. 

Var. peninsularis is larger, glabrous, and shining, found farther 
south upon the peninsula of Lower California (See below). 

I can not determine that Pygmea has any other characters worthy 
of mention as distinguishing it from om7ssa, with which it agrees in 
mental, sternal, abdominal, and crural characters. 

It might be mentioned that the prosternum is more or less convex 
between the coxe and mucronate behind; the mucro is small. The 
mesosternum is feebly oblique and slightly concave. 


ELEODES OMISSA var. PENINSULARIS, new. 


Suboblong-ovate to ovate, glabrous and shining, elytra obsoletely 
subsuleate. 

Head quite finely punctate; antennw somewhat slender, eighth joint 
searcely shorter than the seventh. 

Pronotum subquadrate, widest at the middle, a fifth to a fourth 


‘wider than long; disc evenly and moderately convex from side to 


side, scarcely more arcuately declivous laterally, but distinctly so at 
the apical angles, somewhat obsoletely, finely and sparsely punctu- 
late; apex truncate to feebly emarginate in circular arc; sides 
moderately and rather evenly arcuate, somewhat oblique posteriorly, 
briefly and very feebly sinuate before the base, as seen obliquely from 
the side; base feebly rounded, and scarcely to one-fifth wider than 
the apex; apical angles subacute, with a tendency to become feebly 
prominent anteriorly: basal angles obtuse and not rounded. 

Propleure obsoletely punctulate and more or less rugulose. 

Elytra oval, usually widest behind the middle; disc striato-punc- 
tate, punctures generally not distinctly defined, more or less eroded, 
striae somewhat impressed, intervals apparently somewhat feebly con- 
vex; strial punctures rather moderate in size and rather closely 
placed, intervals minutely, irregularly and obsoletely punctulate, 
punctules rather more distinctly defined laterally, but not usually 
confused. . 

E pipleure obsoletely punctulate. 

Sterna obsoletely punctate and more or less rugose. 


80 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Legs rather somewhat slender. Femora moderately and rather 
sparsely punctate; anterior tibial spurs rather feebly developed. 
First joint of the anterior tarsi very feebly thickened at tip beneath. 
Otherwise as in omissa. 

Male.—More or less oblong-ovate and somewhat slender. Abdomen 
feebly oblique. First joint of the anterior tarsi with a minute and 
pointed tuft of modified spinules on tip beneath. 

Female.—Ovate, anterior spur of the anterior tibize not strongly 
differentiated and not elongate as usual. 

Measurements.—Males: Length, 15-17 mm.; width, 6-6.5 mm. 
Females: Length, 16-18 mm.; width, 7-8 mm. 

Genital characters.—M ale——Edeagophore as in omissa. 

Sternite parabolic in outline, slightly transverse. Each lobe sub- 
triangular; external border quite evenly arcuate, with apex introrsely 
placed and narrowly rounded; internal border straight; surface 
slightly convex, glabrous and not densely punctate in apical half, 
setze moderate in length and not dense. Membrane sparsely setose at 
bottom of the sinus, the latter rounded and subparabolic. 

Female.—Genital segment quadrate; dorsal surface plane and 
setose. 

Valvula.—Dorsal plate oblong, moderately wide and explanate 
apically; swrface nearly plane, very glabrous, sparsely and finely 
punctate, setee moderate, longer towards apex; borders subparallel, 
the external sinuous at basal third, thence feebly arcuate to and con- 
tinuously so with the broadly and strongly rounded apical margin, 
which is minutely sinuate at internal fourth; internal margin sinuous. 
Apical external lobe inferior, the internal small, both with a few long ~ 
flying hairs. 

Appendage small mammilliform, not visible from above, with few 
long hairs on tip. Fossa moderate and fringed with long soft sete. 

Basal prominences not visible laterally. 

Superior pudendal membrane longitudinally rugulose and reaching 
to the base of the internal apical lobe. 

Ventrolateral surface strongly convex at the basal third, thence to 
apex broadly concave; submarginal groove broad and well marked 
basally, thence passing into the general concavity; surface in apical 
two-thirds sparsely and finely punctate and setose. Internal mar- 
gins of the valves contiguous at basal sixth and subcontiguous in 
apical two-sixths; fissure broadly fusiform and closed at the basal 
half by the inferior pudendal membrane, the latter rugulose. 

Habitat—Lower California (Sierra San Lazaro). 

Number of specimens studied, 20. 

Types in the collection California Academy of Sciences; co-sexi- 
types In my own. 

Type-locality—Sierra San Lazaro, Lower California. 


REVISION OF ELEODIINI—BLAISDELL. 81 


Salient type-characters.—Glabrous and shining; elytra obsoletely 
subsuleate. Pronotum widest at the middle; dise scarcely more arcu- 
ately declivous laterally than on the dorsum, distinctly declivous at 
the apical angles, obsoletely and sparsely punctulate; apical angles 
subacute with a tendency to be feebly prominent anteriorly. Elytra 
usually widest behind the middle; dise striato-punctate, punctures 
more or less eroded, striae somewhat impressed, intervals apparently 
somewhat feebly convex and obsoletely punctulate. Legs somewhat 
slender. 

Diagnostic characters.—The salient type characters are sufficient 
to differentiate peninsularis from the other races of omissa. 

It is very interesting to note that it is necessary to use care in recog- 
nizing it from the peninsular form of insv/aris, which occurs in the 
same region. 

In insudaris the pronotum is more quadrate and the apical angles 
are more prominent anteriorly, and the anterior femora are feebly 
armed in both sexes. The first joint of the anterior tarsi of the male 
is clothed with a brush of golden pubescence beneath—the latter often 
dark in old specimens. The genital characters are quite different and 
subgenerically so in the two species. In the peninsular form the 
elytra are obsoletely striato-punctate as in oméssa var. peninsularis. 

Dr. George Horn, in his report of the Coleoptera of Baja Cali- 
fornia,’ referred the specimens here described, as peninsularis to gen- 
tilis, and these were confused with the peninsular form of insularis 
which were also referred to gentilis. (See Promus insularis.) 

Three specimens in the academy’s collection differed from the type 
series of peninsularis in being intensely black and highly polished. 
One female was slightly aberrant; serial arrangement of the elytral 
punctures distinct, strial punctures distinctly larger than the inter- 
stitial; the pronotum had the sides quite strongly margined, the bead 
thin and reflexed. These specimens were destroyed in the great confla- 
gration of April 18, 1906. 


GROUP I. QUADRICOLLIS SECTION. 
ELEODES QUADRICOLLIS Eschscholtz. 


Bleodes quadricollis EScHscHoutz, Zool. Atlas, III, 1833, p. 12, pl. xv, fig. 
5.—MANNERHEIM, Bull. Soc. Nat. Moscow, XVI, 1843, p. 268.—LrE 
Conte, Proc. Acad. Nat. Sci. Phila., 1858, p. 181.—Horn, Trans. 
Amer. Phil. Soc., XIV, 1870, p. 388.—Casey, Annals N. Y. Acad. Sci., 
V, Noy., 1890, p. 395.—Cuampion, Biol. Centr.-Amer., IV, Pt. 1, 1884, 
p. 80. 

Bleodes tarsalis Casey, Annals N. Y. Acad. Sci., V, Nov. 1890, p. 399. 


* Proc. Cal. Acad. Sci., IV, Pt. 1. p. 349, 
59780—Bull. 63—09--—6 


82 BULLETIN 63, UNITED STATES NATIONAL MUSEUM 


Oblong-ovate to ovate, moderately elongate, shining, sometimes 
subasperulate at the sides of the elytra. " 

Head scarcely twice as wide as long, more or less broadly and 
feebly impressed laterally, frontal suture usually evident and more 
or less bisinuate; frons broadly flattened to feebly convex, coarsely 
punctate, punctures finer and sparser on vertex, more coarsely, 
densely, and somewhat confluent on the epistoma. Antenne mod- 
erate in length and comparatively stout, outer four joints very feebly 
compressed and not dilated, third joint subequal to the combined 
lengths of the next two, the fourth just noticeably longer than the 
fifth, the latter, sixth, seventh, and eighth subequal in length, the 
eighth subtriangular, ninth and tenth suborbicular, eleventh short 
ovate. 

Pronotum more or less subquadrate, usually widest before the 
middle, occasionally at the middle, from one-sixth to one-third wider 
than long; disc evenly and moderately convex, arcuately declivous 
laterally, rather coarsely, almost evenly and rather thickly punctate, 
punctures generally denser at the sides where it is also more or less 
rugulose on the declivity; apex subtruncate to very feebly emargi- 
nate, finely and more or less obsoletely beaded; sides rather strongly 
arcuate anteriorly, thence straight and obliquely converging to the 
base as viewed vertically from above, more broadly and eyenly arcu- 
ate in anterior three-fourths, thence more or less feebly sinuate to 
base as viewed obliquely from the side, finely beaded; base subtrun- 
cate to feebly rounded, finely margined, one-fifth to one-third wider 
than the apex; apical angles obtuse and slightly rounded; basal 
angles obtuse. 

Propleure usually glabrous, sparsely punctulate, and more or less 
irregularly rugulose. 

Elytra oblong-oval to oval, frequently strongly inflated, widest at 
the middle; base truncate to feebly emarginate; humeri obtuse, at 
times slightly prominent, narrowly rounded; sdes evenly arcuate, 
apex obtusely rounded; désc moderately convex on the dorsum, 
strongly and quite evenly rounded laterally, inflexed sides broadly 
und feebly concave before the apex, dorsum arcuately declivous 
posteriorly, strongly, quite evenly and diffusely punctate to striato- 
punctate, the punctures generally simple but slightly asperate lat- 
erally and on apex. 

Epipleure moderate in width, gradually narrowing from base to 
apex, very feebly and broadly sinuate beneath the humeral region; 
surface finely and sparsely punctate. 

Sterna quite strongly and densely punctate. rugose. 

Parapleure strongly and densely punctate. 

Abdomen glabrous and shining, rather thickly punctate and 
rugulose. 


—_—_—_— 


— == 


REVISION OF ELEODIINI-——-BLAISDELL. 83 


Legs moderate, strongly punctate, anterior femora mutic; anterior 
tibial spurs dissimilar; first joint of the anterior tarsi slightly thick- 
ened at tip beneath. 

Male.—Oblong-ovate, comparatively narrow, sometimes subcylin- 
drical. Antenne reaching to the base of the prothorax.  Elytra 
rather narrowly oblong-oval, gradually narrowed in apical fourth, 
arcuately and slightly obliquely declivous behind. Abdomen dis- 
tinctly oblique, very moderately convex and broadly impressed at 
middle of the first two segments. Femora sometimes slightly thick- 
ened; anterior tibial spurs rather short, the anterior about one-half 
longer than the posterior, feebly thickened and gradually narrowed 
from base to apex, acute. Anterior tarsi with a tuft of spinules on 
the thickened tip beneath, the pencil of modified spinules scarcely 
evident. 

Female.—Body robust, ovate, antenne not reaching the base of the 
prothorax. Elytra broadly oval, frequently inflated, moderately nar- 
rowed at posterior sixth, arcuately and vertically declivous behind. 
Abdomen horizontal and strongly convex. Anterior spurs of the 
anterior tibiz robust and broad, at least twice as long as the posterior, 
sides arcuate and parallel, narrowed in about apical fourth and sub- 
acute. Anterior tarsi with a small transverse tuft of ordinary spi- 
nules on the thickened tip of the first joint beneath. 

Measurements.—Males: Length, 16.5-21 mm.; width, 6-7.5 mm. 
Females: Length, 17.5-24 mm.; width, 8-12 mm. 

Genital characters, male—Edeagophore (Plate 2, fig. 8) fusiform 
to flattened oblong-ovate, rather stout and arched. 

Basale oblong; surface rather strongly convex, sides more or less 
arcuate. 

Apicale triangular, slightly elongate; surface somewhat strongly 
and evenly convex, with a median membranous groove in apical half; 
sides feebly arcuate in basal half, thence broadly and feebly sinuate 
to apex, the latter somewhat narrow and acute; base with a small 
rounded lobe at middle and broadly sinuate laterally. 

Sternite (Plate 2, fig. 5) transverse. Each lobe slightly quadrate, 
with the external border quite straight in basal half, thence broadly 
arcuate to apex, angle narrowly rounded; internal border quite 
straight; surface moderately convex, sparsely and strongly punctate, 
setose, setae moderate, longer and denser in apical area. Membrane 
sparsely setose at bottom of the sinus, the latter broad. 

Female.—Genital segment (Plate 2, fig. 6) quadrate, not deflexed 
apically and setose, 

Valvula.—Dorsal plate oblong, slightly narrowing toward apex, 
explanate externally and at apex; surface nearly plane, impunctate 
in basal half, elsewhere sparsely punctate and setose, sete rather 
short and reclining; external border nearly straight, converging 


84 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


apically; apical border strongly arcuate on the external lobe, the 
latter occupying the entire apical margin, with aysmall sinuation sep- 
arating it from the minute internal lobe; internal border nearly 
straight, at times more or less sinuous. Apical margin clothed with 
rather long flying soft sete. 


Appendage not visible beyond the margins of the fossa, short, 


mammilliform. 

Basal prominences not evident. 

Superior pudendal membrane longitudinally rugulose, and reach- 
ing to opposite the internal apical lobe. 

Ventrolateral surface (Plate 2, fig. 7) moderately convex basally 
and feebly concave before the apex, punctuate and setose, sete rather 
short, long about the fossa; submarginal groove moderate beneath 
the somewhat explanate external border of the dorsal plate, curving 
inward beneath the external apical lobe to the fossa. Internal mar- 
gins of the valves contiguous in basal eighth; fissure rather broadly 
fusiform, closed in the basal half by the inferior pudendal membrane. 

Variations in genital characters.—In the most ventricose specimen 
that I examined the entire apical border was taken up by the exter- 
nal lobe; the internal lobe was not evident from above; the dorsal 
plate was narrower than usual. The genital segment viewed ven- 
trally showed the inferior border of the valvular membrane as form- 
ing the internal wall of the fossa, and that the internal lobe was 
therefore small and ventral. Appendage slightly compressed. 

In a specimen exhibiting marked variation, I found the sete 
shorter, and the external lobe of the apex more membranous; the 
sides of the dorsal plate were quite straight and parallel; the internal 
lobe was ventral as above, and the inferior pudendal membrane was 
longer than usual. 

Habitat—California (about the Bay of San Francisco, Esch- 
scholtz; Mount Diablo, Thomas Casey; Monte Bello, near Mountain 
View, Santa Clara County, Edward Ehrhorn; San Emigdio Canyen, 
Kern County, Fordyce Grinnell). 

Number of specimens studied, 500. 

Type, location unknown to me. 

Type-locality—Foothills near San Francisco. 

Salient type-characters.—Thorax punctulate, subquadrate, broadest 
anteriorly ; elytra punctato-striate. Body of the male subcylindrical, 
of the female obovate (Eschscholtz). 

Diagnostic characters —The strongly developed anterior spurs of 
the anterior tibize (females) characteristic of the present section, 
separate quadricollis from all the members of the Carbonaria section. 
But it will be necessary to actually scrutinize the anterior spurs of 
the very robust examples of omissa, where the anterior spurs are cer- 


_—s Se ee 


a es ax St eC CO 


REVISION OF ELEODITNI—BLAISDELL. 85 


tainly strongly developed for that section, and while it will be seen 
that the spurs are distinctly broadened, they will also be observed to 
taper more gradually from base to apex, and never so parallel nor 
so suddenly or apically narrowed as in quadricollis. Unworn spurs 
must always be taken as the criteria. 

The var. anthracina described below is duller in surface luster, less 
convex, not inflated, and in typical specimens have the elytra just 
noticeably obsoletely subsulcato-striate, characters never observed 
in the true guadricollis; the latter by its quite simple elytral punctua- 
tion is easily separated from humeralis; in cuneaticollis the elytral 
sculpturing is distinctly muricato-rugulose. i/eyi is more robustly 
ovate. - 

Quadricollis has been up to the present time confused with omissa. 
Reference to the original description of this species as given by 
Eschscholtz should quickly close all controversy. That author writes 
that the elytra are punctato-striate (elytris punctato-striatis) and 
that this species is plentiful in the foothills about San Francisco 
(“ Bei St. Franzisco auf den Anhohen hiufig”). These remarks can 
only be applied to the present species. 

What has previously been said in regard to the variation of the 
elytral punctuation in the different species must be recalled at this 
point. 

All those specimens with the elytra punctato-striate are to be re- 
garded as typical. Fully two-thirds of the large series before me 
have the “ sculpture consisting of punctures sometimes fine, at others 
rather coarse, rather densely but irregularly placed, and never muri- 
cate; never arranged in rows” (Horn). 

Doctor Horn undoubtedly included oméssa in the above definition, 
but then a large percentage of the examples of that species exhibit 
a serial arrangement of the punctures. After all, the characters 
apply to guadricollis, for the punctures are sometimes fine, but gen- 
erally rather coarse; punctato-striate specimens are not common, and 
never really muricate in the sense as it is applied to Aumeralis and 
its races. Quadricollis and omissa as well have a feeble type of sub- 
muricate punctuation at the elytral sides and apex. 

I have a male guadricollis at hand, kindly transmitted to me a 
number of years ago by Colonel Casey. It is not at all inflated, was 
taken near San Francisco, and is diffusely punctured; bears the label 
quadricollis in Casey’s handwriting. I consider it a typical specimen, 
except as regards the punctuation. 

Tarsalis is only an inflated form (forma tarsalis) of quadricollis. 
Colonel Casey himself has made this fact known.* 


4 Coleopterological Notices, V, p. 597, Annals N, Y, Acad. Sci., Dee., 1S. 


86 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


The student will at times find it very difficult to decide whether a 
given specimen (especially males) shall be referred to guadricollis or 
omissa. 

In my experience the epistoma in guadricollis is always more 
coarsely punctate than in omissa. Specific differentiation is no more 
difficult in this instance than is met with in other sections of the genus 
Eleodes. 

The extreme forms or heterotypes of carbonaria and obsoleta are 
as difficult of separation as are the above, in fact, all the constituents 
of the present section of the subgenus J/elaneleodes might be consid- 
ered as races and forms of a single species if viewed from a broad 
evolutionary standpoint. 

The mentum is usually triangulo-trapezoidal, strongly punctate 
and foveate laterally within the more or less strongly defined margins; 
the surface is longitudinally ridged at the middle. 

The prosternum is more or less strongly protuberant ventrally with 
the coxee, and more or less longitudinally convex between the same, 
usually grooved at the middle and rather distinctly margined around 
the acetabula; frequently submucronately produced behind and at 
times nearly vertically subtruncate. 

The mesosternum more or less concave and arcuately declivous. 

The intercoxal process of the abdomen is quadrate (male) to feebly 
transverse (female) and about a third of its width wider than the 
metasternal salient. 

The metasternum laterally between the coxe is as long as the width 
of a mesofemur at the base. 

The post-coxal portion of the abdomen is equal in length to that 
of the process, and also to that of the third segment. 

In the male the second segment is about equal to the width of the 
process, and the third is a half longer than the fourth. In the female 
the second is twice as long as the fourth segment. 

The profemora are usually moderately clavate in the male and less 
so in the female. The grooves are limited by rather strong margins, 
which are more or less asperulate and converge near the base. 

The meso- and metafemora have their superior and inferior sur- 
faces quite parallel. The margins of the grooves are not strongly 
marked, are finely muricate, and become evanescent before becoming 
contiguous. 

The tibie are frequently more or less arcuate and without tarsal 
grooves; the articular cavities are closed. 

The tarsi are somewhat variable as regards to stoutness, fre- 
quently rather more slender in the female. The protarsi are usually 
slightly thicker in the male and apparently subequal in the sexes. 

The protarsi are about two-fifths of their length shorter than a 
mesotarsus, Joints two, three, and four are subequal in the male; 


REVISION OF ELEODITINI—BLAISDELL. 87 


the fourth a little smaller than the preceding in the female; the fifth 
is quite equal to the combined lengths of the preceding three; the 
first about a half longer than wide. 

The mesotarsi are about two-sevenths (male) or a little less (fe- 
male) of their length shorter than a metatarsus. Joints two to four 
are just a little progressively shorter or subequal, and together a 
little longer than the fifth, which is quite equal to the first. 

A metatarsus is about five-ninths of its length shorter than a meta- 
tibia. Joint three is just noticeably shorter than the second, both 
together equal to the length of the first, the fourth a little shorter. 


ELEODES QUADRICOLLIS var. ANTHRACINA, new. 


Oblong-ovate to ovate, surface subopaque, elytra usually obsoletely 
subsulcate; antenne reaching beyond the base of the prothorax in the 
male, 

Head somewhat coarsely punctate, the coarser punctures on the 
epistoma not confluent nor crowded. Antenne rather long, slightly 
stout, feebly clavate, outer four joints slightly compressed and 
feebly dilated, third joint as long as the next two combined, fourth 
a little longer than the fifth, the latter, sixth, and seventh subequal, 
the eighth shorter and subtriangular, ninth and tenth suborbicular, 
eleventh ovate to oval. 

Pronotum subquadrate, usually widest at about the middle, near a 
sixth wider than long; disc evenly and moderately convex, arcuately 
declivous laterally, finely and rather sparsely punctulate, scarcely 
coarser or denser laterally, where it is frequently slightly rugulose ; 
apex subtruncate in circular arc, and finely or obsoletely margined ; 
sides evenly and moderately arcuate in the anterior three-fourths. 
thence converging to base, frequently rather evenly arcuate from apex 
to base (viewed vertically from above), margin finely beaded; base 
slightly rounded, often feebly sinuate at the middle, finely and more 
or less obsoletely beaded, about one-fourth wider than the apex and 
equal to the length; apical angles distinct and very narrowly rounded ; 
basal angles obtuse. 

Elytra oval, widest at the middle, less than twice as long as wide; 
base more or less emarginate, frequently slightly sinuate laterally, 
usually a little wider than the contiguous prothoracic base; humeri 
obtuse, narrowly rounded and scarcely at all prominent anteriorly; 
dise moderately convex on the dorsum, more strongly and almost 
evenly rounded laterally, arcuately declivous posteriorly, surface 
usually striato-punctate, intervals apparently very feebly convex, 
giving an obsoletely and faintly subsulcate appearance, strial punc- 
tures somewhat small, series moderately distant, rarely impressed, 
interstitial punctures more or less irregular on the dorsum, quite 
equal in size to the strial; frequently the punctures of both series are 


88 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


arranged without trace of order, nearly always so laterally and on 
the apical declivity, where they are scarcely ever subasperate. 

Otherwise as in quadricollis. 

Male.—Antenne reaching a short distance beyond the base of the 
prothorax. Pronotum usually widest at the middle. 

Female.—Antenne reaching to the base of the prothorax. Prono- 
tum not usually very noticeably widest before the middle. 

Otherwise the sexual characters are as in quadricollis. 

Measurements.—M ales: Length, 18-19.5 mm.; width, 6.5-7.5 mm. 
Females: Length, 18-22 mm.; width, 7.5-10.5 mm. 

Genital characters, male.—Edeagophore flaxseed-shaped and mod- 
erately arched. 

Basale oblong, quite evenly convex above; sides slightly arcuate. 

Apicale triangular and moderately elongate; surface moderately 
and evenly convex, with a median semimembraneous groove in apical 
half; sides nearly straight, apex subacute; base rather broadly and 
arcuately lobed at the middle, and slightly sinuate laterally. 

Sternite parabolic in outline and slightly transverse. Each lobe 
with the external border quite evenly arcuate to apex, the latter nar- 
rowly rounded; internal border short and nearly straight; surface 
feebly convex, impunctate in basal half, sparsely punctate apically, 
setose, setae moderate in length, longer about the apex, not dense. 
Membrane sparsely setose across the bottom of the sinus, the latter 
short. 

Female.—Genital segment quite quadrate, explanate externally in 
apical two-thirds. 

Valvula.—Dorsal plate oblong, moderate in width, sides subpar- 
allel; surface quite plane, glabrous, coarsely and sparsely punctate, 
each puncture with a moderately long reclining seta, impunctate at 
base; external border more or less straight to feebly arcuate to the 
apical border, which is strongly rounded on the external lobe and 
slightly sinuate just external to the minute internal lobe; external 
lobe about as wide as the dorsal plate, both lobes sparsely set with 
long sete; internal margin more or less sinuous. 

Appendage not visible from above, short mammilliform and ~ 
scarcely projecting beyond the margins of the fossa. 

Basal prominences scarcely evident. 

Superior pudendal membrane longitudinally rugulose, reaching to 
the internal lobe. 

Ventrolateral surface as in quadricollis. - 

Habitat—Arizona (Fort Grant, July 10; Hot Springs, June; 
Oracle, San Simon, Galiuro Mountains, May; Catalina Springs, 
April; Tucson, Hubbard and Schwarz; Nogales, September, Albert 
Koebele and F. W. Nunenmacher). 


Se ee 


REVISION OF ELEODITNI—BLAISDELL., 89 


Number of specimens studied, 150. 

Types (Cat. No. 12202) are in the U. S. National Museum collee- 
tion; cotypes in my own. 

Type-locality.—F ort Grant, Arizona. 

Salient type-characters—Integuments dull in luster. Epistoma 
somewhat coarsely punctate, punctures not crowded nor confluent. 
Antenne in the male reaching beyond, and in the female to the pro- 
thoracic base. Thorax in both sexes widest at about the middle and 
not more strongly arcuate before the same. Elytra striato-punctate, 
obsoletely and faintly subsulcate. 

Diagnostic characters.——Anthracina differs from quadricollis in 
its salient type-characters. For its separation from carbonaria, 
humeralis, and cuneaticollis, see differential diagnosis under qguad- 
ricollis. 

I have seen specimens of this race labeled carbonaria, and it is 
generally labeled guadricollis in collections. 

In its dull luster and long antenne it might be confused with 
ampla, but the shorter legs and large anterior spurs of the anterior 
tibiz in the female will separate it. 

Mr. Blanchard has compared it with the LeConte types, especially 
vicina, and says that it differs from all and is most closely related to 
quadricollis. 

Prosternum as in guadricollis, but less inclined to be submucronate 
and less frequently grooved at the middle between the coxe. The 
mesosternum is frequently slightly more strongly concave. 

Otherwise as in qguadricollis. 


ELEODES QUADRICOLLIS var. LUSTRANS, new. 


Oblong-ovate to ovate, moderately elongate, very glabrous and 
shining. 

Head comparatively rather small, frons moderately and quite 
evenly convex, scarcely to distinctly impressed laterally, frontal 
suture usually not evident, almost coarsely, rather thickly and evenly 
punctate, punctures usually not denser on the epistoma. Antenne 


rather long, somewhat slender, outer four joints scarcely at all com- 


pressed or dilated, third joint quite equal in length to the next two 
taken together, the fourth just visibly longer than the fifth, the latter, 
sixth and seventh subequal, eighth just the least shorter and subtri- 
angular, ninth and tenth suborbicular, eleventh short ovate or oval. 

Pronotum subquadrate, widest at or just a little in front of the 
middle, one-fourth to one-third wider than long; dise evenly and 
moderately convex, more or less arcuately declivous at the sides and 
apical angles, finely and sparsely punctate, sometimes a little more 


- 


90) BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


coarsely so laterally; apex truncate in circular arc, finely and more or 
less obsoletely beaded; sides evenly and not strongly arcuate in 
anterior three-fourths, thence convergent to the base, or more or less 
arcuate from apex to base, as viewed vertically from above, more 
broadly arcuate and. just the least sinuate before the base as viewed 
obliquely from the side, rather distinctly and finely beaded; base 
feebly rounded and more or less finely margined, a seventh to a fifth 
wider than the apex; apical angles obtuse and not in the least promi- 
nent anteriorly; basal angles obtuse. 

Elytra oval, widest at the middle, scarcely twice to one-fourth 
longer than wide; base truncate to feebly emarginate, scarcely to a 
little wider than the contiguous prothoracic base; hwmeri obtuse; 
sides evenly arcuate, apex not very narrowly rounded; dése slightly 
flattened and moderately convex on the dorsum, rather evenly and not 
very broadly rounded at the sides, arcuately declivous posteriorly, 
punctate, punctures rather fine and more or less irregular, usually 
with a serial arrangement on the dorsum, interstitial punctures more 
or less irregular, those of both series equal in size or the strial are 
slightly larger; both series always confused at the sides and on the 
apex; not at all submuricate. 

Legs rather slender. 

Otherwise as in guadricollis. 

Male somewhat depressed. 

Female.—Convex and comparatively less elongate than in an- 
thracina. 

Other secondary sexual characters as in guadricollis. 

Measurements.—M ales: Length, 16-17 mm.; width, 6-7 mm. fe- 
males: Length, 16.5-17 mm.; width, 8-8.2 mm. 

Genital characters, male—Sternite with each lobe more or less 
parabolic in outline. Otherwise as in qguadricollis. 

Female. is, except that the ventrolateral surface 
is strongly convex at basal fourth and broadly concave before the 
apex, setee rather long internally. 

Habitat—Arizona (Chiricahua Mountains, Fuster Hubbard and 
Schwarz; Catalina Springs, April; Tucson.) 

Number of specimens studied, 9. 

Types (Cat. No, 12203) in the U. 8S. National Museum collection. 

Ty pe-locality— Chiricahua saunas Arizona. 

Salient type-characters.—Glabrous and shining. Antenne reaching 
beyond (male) or to (female) the base of the prothorax. Body in 
the female less elongate than in quadricollis. Punctures on the epis- 
toma not coarser than on the frons.. Pronotum widest at the middle; 
disc finely and sparsely punctate, punctures not coarser nor denser 
laterally. Elytra with the discal punctures arranged in series on the 


REVISION OF ELEODITNI-——BLAISDELL. 9] 


dorsum, laterally confused and not submuricate; punctures of both 
series fine and equal in size. 

Diagnostic characters.—Difters from both quadricollis and its race 
anthracina by the salient type characters. In both sexes the form is 
smaller and less stout and the female is decidedly less elongate and 
more robustly ovate. The punctuation is noticeably finer in all the 
specimens that I have seen, the epistoma is more sparsely and finely 
punctate, the punctures being not at all confluent. A striking feature 
in the sculpturing is the very smooth integuments. 

The large anterior tibial spurs separate it from all the constituents 
of the Carbonaria section. 

Mr. Blanchard, after careful comparisons made with the LeConte 
material, writes: “ I do not recognize this as belonging to any of the 
named forms; it is nearest to guadricollis.” 

The prosternum is more often decidedly mucronate, although not 
strongly produced. Otherwise as in qguadricollis. 


ELEODES CUNEATICOLLIS Casey. 


Fleodes cuneaticollis Casey, Annals N. Y. Acad. Sci., V. Nov. 1890, p. 397. 


Oblong-ovate to ovate, elongate, rather shining; elytra coarsely 
rugulose. 

Head twice as wide as long, feebly convex or flattened between the 
eyes, coarsely and strongly, irregularly and more or less confluently 
punctate, punctures finer on the vertex. Antenne moderate in length 
and somewhat stout, scarcely compressed and not dilated, third joint 
fully as long as the next two combined, the fourth but slightly longer 
than the fifth, the latter, sixth, seventh, and eighth subequal, ninth 
suborbicular, tenth transverse, eleventh short ovate. 

Pronotum widest at the junction of the anterior and middle thirds, 
one-third to two-fifths wider than long; disc broadly and moderately 
convex, more or less declivous laterally and at the apical angles, rather 
sparsely, coarsely, and deeply punctate, the punctures about twice as 
large and distinct as those of humeralis, frequently more or less rugu- 
lose at the sides; apex truncate or broadly and feebly emarginate in 
circular arc, very finely and more or less obsoletely beaded; sides 
strongly arcuate anteriorly, conspicuously convergent and almost 
straight in basal half, viewed vertically from above, or more broadly 
arcuate in anterior three-fourths, thence convergent and more or less 
sinuate to the base, as viewed obliquely from the side, finely beaded ; 
base truncate to slightly rounded and finely margined, about equal to 
the length and one-tenth to one-fifth wider than the apex: apical 
angles obtuse and more or less narrowly rounded; basal angles obtuse. 


92 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Propleure sparsely punctate and more or less strongly rugulose. 

Elytra oval, widest at the middle, less than twige as long as wide; 
hase truncate to feebly emarginate, sometimes sinuate laterally, equal 
to or slightly wider than the contiguous prothoracic base; humeri 
obtuse, at times more or less rounded, again slightly prominent; sédes 
evenly arcuate, somewhat pointed at the apex, the latter not broadly 
rounded; disc more or less moderately convex on the dorsum, quite 
strongly and evenly rounded laterally, arcuately declivous posteriorly ; 
inflexed sides somewhat oblique; very coarsely, deeply, and densely 
punctate, punctures irregularly distributed without trace of a 
serial arrangement, not muricate, but producing a strongly rugu- 
lose appearance by mutual semi-coalescence. 

Epipleurw moderate in width and gradually narrowing from base 
to apex, superior margin very broadly and feebly sinuate beneath 
the humeri; surface sparsely punctate. | 

Sterna somewhat shining, densely punctate and more or less 
rugose. 

Parapleure coarsely and densely punctate. 

Abdomen glabrous, sparsely punctate, punctures coarser and denser 
on the fifth segment; surface more or less rugulose. 

Legs short and rather robust. Anterior femora mutic; anterior 
tibial spurs dissimilar in the sexes; first jomt of the anterior tarsi 
thickened at tip beneath, feebly dissimilar in the sexes. 

Male.—More or less slender. Antennz reaching to the base of the 
prothorax. Elytra arcuately and more or less obliquely declivous 
behind and gradually narrowed in the posterior fourth. Abdomen 
more or less distinctly oblique, moderately convex, broadly im- 
pressed on first segment, less so on the second. Anterior spurs of 
the anterior tibie slender, acute, and about a third longer than the 
posterior. First joint of the anterior tarsi with the tuft of spinules 
obsolete at tip beneath, groove slight and not interrupted on the 
thickened tip. 

Female.—Ovate, robust, antenne not quite reaching to the pro- 
thoracic base. Elytra broadly oval, arcuately and more or less ver- 
tically declivous and less strongly narrowed behind. Abdomen hori- 
zontal and more or less strongly convex. Anterior spurs of the 
anterior tibiz moderately broadened, sides more or less parallel, 
narrowing in apical fourth, rather acute or blunt if worn. First 
joint of the anterior tarsi distinctly thickened at tip beneath, tuft of 
spinules not evident. 

Measurements.—Males: Length, 14-17.5 mm.; width, 5.5-6.8 mm. 
Females: Length, 15-18.8 mm.; width, 7-9.2 mm. 

Edeagophore elongate and acutely 
ovate, and rather strongly arched. 


Genital characters, male. 


———=— ee eC 


REVISION OF ELEODIINI—BLAISDELL. 93 


Basale rather short, suboblong oval; sides feebly arcuate; surface 
evenly convex. 

Apicale elongately triangular and rather arched; surface evenly 
convex, with a rather narrow longitudinal groove in apical two- 
thirds; sides nearly straight, apex acute; base with a rather small 
rounded lobe at middle and feebly sinuate laterally. 

Sternite short and transversely parabolic in outline, each lobe 
somewhat transverse with the external border broadly arcuate; apex 
slightly prominent and rather narrowly rounded; internal border 
rather straight; surface feebly convex, strongly and sparsely punctate, 
sete rather long. Membrane very sparsely setose across the bottom 
of the sinus, the latter short. 

Female.—Genital segment quadrate, valves with their external 
borders feebly converging, setose. 

Valvula (Plate 3, fig. 18).—Dorsal plate oblong, feebly explanate 
externally and nearly plane; surface glabrous, finely and very 
sparsely punctate, sete short and reclining; external border more or 
less feebly arcuate; internal border straight or sinuous; apical margin 
moderately arcuate in external three-fourths on the external lobe; 
internal lobe not prominent and separated from the external by a 
feeble sinuation. 

Appendage sometimes visible from above, rather short mammilli- 
form, with a few short setw at tip. Fossa large and quite terminal, 
the inferior margin of the valvular membrane forming the internal 
margin, the latter more prominent than the internal lobe above. 

Basal prominences not in the least evident. 

Superior pudendal membrane longitudinally rugulose, long, reach- 
ing to the internal apical lobe. 

Ventrolateral surface moderately convex, not prominent at the base 
and not transversely concave before the apex; submarginal groove 
very broad and shallow, forming a lateral concavity before the apex. 

Habitat—California (San Francisco, Marin, Alameda, and San 
Mateo counties, August). Specimens have been distributed as having 
been collected in San Diego County, but all of those were undoubtedly 
collected by Mr. G. W. Dunn and sent out without true locality 
labels—at least that is the case with a series that I have had for years. 
Personally I have collected them only in the above-named localities 
and have no authentic data of their having been taken elsewhere. 
Colonel Casey’s type specimen was without an exact locality label. 

Number of specimens studied, 30. 

Type in Colonel Casey’s collection. 

Type-locality —California; probably from some place about the 
Bay of San Francisco, 


94 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Salient type-characters——Head rather large, coarsely and deeply 
punctate. Antenne short and robust. Prothorax with apex nearly 
as wide as the base; sides strongly arcuate anteriorly, conspicuously 
convergent and almost perfectly straight in basal two-thirds; disc 
widest at apical third, rather sparsely, coarsely and deeply punctate, 
punctures about twice as large and distant as those of humeralis. 

Elytra distinctly less than twice as long as the prothorax; disc 
rather depressed above, very coarsely, deeply, and densely punctate, 
the punctures irregularly arranged without trace of impressed strie, 
not muricate but producing a strongly rugulose appearance by mutual 
semi-coalescence. Legs short and robust (Casey). 

Diagnostic characters —Cuneaticollis differs from all the other 
species of the present subgenus in having the elytra strongly rugulose 
from semi-coalescence of the coarse punctures. As Casey writes, it 
belongs near humeralis, but differs in four important characters, 
namely: “ The much shorter and robust antenne and legs, the very 
much coarser and deeper elytral sculpture, and coarser, sparser pro- 
notal punctures, and finally the smaller and much less unequal spurs 
of the anterior tibiz in the male, the larger spur in humeralis being 
nearly four times as long as the smaller one.” In the largest females 
before me the humeri are exposed. 

The most noticeable disparity in the elytral characters of Casey’s 
type and the series before me, is that, while in the type the elytra 
are distinctly less than twice as long as the prothorax, in all the 
examples before me the elytra are distinctly a little more than twice 
as long as the same. My series agrees in all other particulars with 
the original description. I have not seen any other species that could 
be referred to cuneaticollis; Casey does not mention the number of 
specimens he had before him at the time of writing his description, 
but specimens do occasionally have unusually short elytra in other 
species. In some examples the elytra are vaguely substriate when 
viewed longitudinally. 

The individual specimens of cuneaticollis exhibit considerable 
variability, more so than in the other related species. The largest 
female before me has abnormally short legs, another has the sides of 
the prothorax unusually and strongly rounded in the anterior third; 
the antenne are noticeably more robust and shorter in some specimens 
than others. I believe the present species to be more closely related to 
quadricollis than to humeralis; the anterior tibial spurs suggest this 
relationship. 

The mentum is subparabolic to triangulo-parabolic, not coarsely but 
densely punctate, and not noticeably setose; surface more or less 
foveate laterally and broadly convex centrally. 

The prosternum and mesosternum are variable, as in humeralis. 


/ 


REVISION OF ELEODIINI—BLAISDELL. 95 


The intercoxal process of the abdomen is subquadrate (male) or 
slightly transverse (female) and about one-fourth of its width wider 
than the metasternal salient. 

The metasternum laterally between the coxe is as long as the width 
of a mesotibia at the extreme apex. 

The post-coxal part of the first abdominal segment is equal in 
length to the process, and the latter to that of the third segment. 

In the sexes the second segment is twice as long as the fourth. 

The profemora are more or less distinctly clavate; the grooves are 
moderately broad and the margins subcariniform, more or less asperu- 
late, becoming contiguous and evanescent at basal third. 

The mesofemora are scarcely or very feebly clavate, grooves quite 
plane, margins feeble and asperulate, evanescent at middle before 
becoming contiguous. 

The metafemora are not at all widened externally and the superior 
and inferior surface lines are quite parallel; grooves as on the meso- 
femora. 

The tibize may be more or less feebly arcuate, tarsal grooves absent, 
articular cavities closed. 

The pro- and mesotibizw are somewhat inwardly produced at apex. 
The protibie are sometimes quite distinctly carinate externally in 
basal half, noticeably so in the largest female before me. 

Tarsi variable in stoutness. The protarsi are about one-half (male) 
or one-fourth (female) of their length shorter than a mesotarsus. 
Joints two, three, and four subequal in size, and together a little 
longer than the fifth; the first slightly shorter than the third and 
fourth taken together. 

The mesotarsi are about a ninth (male) or a fifth (female) of their 
length shorter than a metatarsus. Joints two, three, and four feebly 
decreasing in length in the order named; the fifth subequal to the 
combined lengths of the second and third, the first to the third and 
fourth. 

. A metatarsus is about a half (male) or a third (female) of its 
length shorter than its metatibia. Joints two and three are subequal, 
and together about equal to the fourth, which is subequal to the first. 


ELEODES HUMERALIS LeConte. 


EBleodes humeralis LeConte, Reports of Explor. and Surveys . . . 47th 
and 49th Parallel, XII, Appendix No.1, 1857, p. 50; Proc. Acad. Nat. 
Sci. Phila., 1858, p. 182.—Horn, Trans. Amer. Phil. Soc.. XIV. 1870, 
p. 309.—CHAMPION, Biol. Centr.-Amer., IV, Pt. 1, 1884, p. 80. 
Oblong-ovate to ovate, elongate, more or less opaque: elytra 
densely, rather finely and muricately, or granulato-muricately pune- 
tate. 


96 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Head rather less than twice as wide as long, frequently feebly and 
rather broadly impressed laterally, frontal suturg obsolete or feebly 
evident and scarcely sinuate; frons feebly convex, usually more or 
less flattened, densely and finely, sometimes a little coarser and more 
or less confluently punctate, punctures fine on the vertex increasing 
in coarseness to the epistoma. Antenne long and somewhat slender, 
feebly compressed and scarcely dilated in the outer four joints, third 
joint as long as the next two taken together, fourth slightly longer 
than the fifth, the latter, sixth, and seventh subequal, eighth a little 
shorter and more or less subtriangular, ninth and tenth suborbicular, 
eleventh short ovate. 

Pronotum subquadrate, widest at or just in front of the middle, 
about one-third wider than long; disc moderately and quite evenly 
convex, feebly and rather arcuately declivous laterally, rather finely 
and densely punctate, usually more or less narrowly subgranulate 
along the sides; apex truncate or very feebly emarginate in circular 
arc, usually finely beaded; s¢des rather broadly and moderately are- 
uate in the anterior two-thirds, thence to the base convergent and 
straight or feebly sinuate, marginal bead rather thin, finely reflexed 
and visible throughout its entire length from above; base truncate or 
feebly rounded, finely margined, and about one-sixth wider than the 
apex, about equal to the length; apical angles obtuse, scarcely 
rounded; basal angles obtuse, not rounded, almost rectangular at 
times. 

Propleure quite densely punctato-rugulose. 

Elytra oval, not twice as long as wide, usually widest at the 
middle; base truncate or more or less feebly emarginate, as wide as or 
slightly wider than the contiguous prothoracic base; humeri obtuse 
or slightly prominent; sédes evenly arcuate, apex somewhat narrowly 
rounded; disc more or less slightly depressed on the dorsum, some- 
times slightly ogival, arcuately declivous posteriorly, and quite 
strongly but not broadly rounded laterally, very densely and rather 
finely muricately punctate or almost granulate, punctures more 
simple centrally along the suture, the latter frequently impressed ; 
surface at times is obsoletely striate, the inflexed sides are inwardly 
declivous and with the surface lines somewhat straight. 

Epipleure somewhat narrow, gradually narrowing from base to 
apex, the superior margin is broadly and rather feebly sinuate be- 
neath the humeri; surface densely and subasperulately punctate. 

Sterna more or less shining and very densely punctato-rugulose. 

Parapleure densely punctate. 

Abdomen more or less shining, rather densely punctato-rugulose. 
The first segment is more strongly sculptured, and the fifth quite 
densely punctate. 


EE aa ee 


REVISION OF ELEODITNI—BLAISDELL. 97 


Legs moderate in length, comparatively somewhat slender. Femora 
mutic and not thickened; anterior tibial spurs dissimilar in the sexes; 
anterior tarsi with the first joint slightly thickened at tip beneath, 
groove interrupted at the ventral apical margin, tuft of spinules 
feebly developed and slightly dissimilar in the sexes. 

Male.—Oblong-ovate to ovate, not slender. Antenne reaching a 
short distance beyond the prothoracic base. Abdomen feebly oblique, 
moderately convex, more or less impressed at middle on segments one 
and two. Anterior spurs of the anterior tibize usually about 
three times longer than the posterior, slightly broadened and slender, 
acute, gradually narrowing from base to apex. 

Female.—Ovate, robust. Antenne reaching at least to the protho- 
racic base. Elytra arcuately and vertically declivous behind. Abdo- 
men horizontal, strongly convex. Anterior spurs of the anterior 
tibizx much broadened and produced, about three times as long as the 
posterior, sides subparallel and narrowed in apical fourth, subacute, 
worn spurs are obtusely rounded at tip. | 

In the material before me three forms may be recognized: 

Forma typica.—Elytra generally flattened on the dorsum, sides of 
the dise are not broadly rounded, and the inflexed portions are ob- 
liquely and inwardly declivous. Elytral disc very densely and rather 
finely muricately punctate. 

Forma granulato-muricata.—Elytra generally moderately convex 
on the dorsum, sides more or less broadly rounded, dise moderately 
densely granulato-muricate, granules fine and shining. 

Forma tuberculo-muricata.—Pronotum more arcuately declivous 
laterally, with the margins more or less invisible from above. Elytra 
more coarsely and strongly sculptured, subtuberculately muricate, the 
punctures much sparser than in the typical form. 

Measurements.—M ales: Length, 14.2-17.5 mm.; width, 6.2-7 mm. 
Females: Length, 16-20.2 mm.; width, 7.6-8.5 mm. 

Genital characters, male.—Edeagophore elongately ovate, pointed 
and slightly arched. 

Basale oblong, evenly convex above; sides feebly arcuate. 

Apicale triangular, twice as long as wide, evenly convex above, 
with a rather wide median membranous groove in apical half; sides 
straight and convergent to the subacute tip; base with a rounded lobe 
at middle, feebly sinuate laterally. 

Sternite subparabolic, almost as wide as long. Each lobe with the 
outer border more or less straight in basal two-thirds, thence obliquely 
subtruncate to apex, tip narrowly rounded; internal margin feebly 
arcuate; surface scarcely convex, glabrous, very sparsely punctate in 
apical half and setose, set moderately long, longer on apical margin 
and rather sparse. Membrane with scattered setw across the bottom 
of the sinus, the latter quite deep. 


59T8O—Bull. 68—09-——7 


98 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Female.—Genital segment trapezoidal, valves rather distant and 
convergent apically, setose; superior pudenda, membrane rather 
widely exposed. 

Valvula (Plate 3, fig. 17).—Dorsal plate oblong, feebly explanate 
externally, moderately narrow; surface plane, finely and sparsely 
punctate, also setose in apical half, sete fine and short; borders 
parallel, straight or feebly sinuous; apical margin feebly arcuate in 
outer three-fifths on the external apical lobe, the latter slightly 
defined from the internal lobe by a slight sinuation; internal lobe 
not prominent. Apical parts set with moderately long sete. 

Appendage not visible from above, short mammilliform, not ex- 
tending beyond the margins of the fossa, the latter rather large. 

Superior pudendal membrane longitudinally rugulose and reaching 
to the internal lobe. 

Ventrolateral surface rather evenly convex, not protuberant, sur- 
face lines quite straight when viewed longitudinally, not transversely 
concave before the apex; submarginal groove broad and shallow, 
forming a lateral concavity that does not bend inward beneath the 
apex; surface finely and sparsely punctate and setose, sete very 
short. Internal margins of the valves contiguous in basal eighth; 
fissure rather broadly fusiform, open apically, closed in basal half 
by the inferior pudendal membrane. 

In the granulato-muricata form the genital segment of the female 
is more quadrate, valves quite parallel. The dorsal plate is explanate 
externally in apical two-thirds. Otherwise as in the typical form. 

Forma typica—//abitat.—Washington (Walla Walla; Yakima, C. 
V. Piper) ; Oregon (George Horn, G. W. Dunn) ; California (north- 
ern, George Horn) ; Idaho (Hubbard and Schwarz). 

Forma granulato-muricata—Nevada (Verdi, April, F. E. Blais- 
dell; Carson City, July; Reno, H. F. Wickham) ; Utah (C. V. Riley) ; 
California (Lassen County). 

Forma tuberculo-muricata—Colorado (C. V. Riley; South Park, 
elevation, 8,000-10,000 feet, Bowditch; Buena Vista, Wickham’s 
List). 

Number of specimens studied, 105. 

Type in the LeConte collection. 

Type-locality —* Wenass River to Fort Colville.” 

Salient type-characters.—Opaque, head and thorax densely pune- 
tate. Thorax subquadrate, narrowed behind, -sides rounded, base 
truncate, posterior angles obtuse. Elytra depressed on the dorsum, 
obsoletely striate, sculptured with subacute granules that are more or 
less thickly and irregularly placed, punctate on each side of the 
suture. Prosternum perpendicular behind (LeConte). 

Diagnostic characters.—Humeralis exhibits a remarkable develop- 
ment in the anterior tibial spurs. In both sexes the anterior spur is 


oo 


REVISION OF ELEODIINI——BLAISDELL. 99 


about three or four times longer than the posterior. In this it differs 
from all of the other species of the subgenus, except ri/eyi. The 
integuments are more or less opaque, and in the typical form from 
Washington the elytra are densely muricately punctate and more or 
less flattened on the dorsum. 

Specimens from Lassen County, California, have the elytra dis- 
eretely granulate; each granule under low power of magnification is 
polished and shining, under high power minutely pointed. These 
constitute the heterotypical western form; in the Nevada specimens 
the granules are smaller, less shining, but discrete. 

In the Colorado examples the granules are much larger and sparser, 
and characterizes the incipient race here designated as forma 
tuberculo-muricata. 

The Idaho specimens are feebly shining and more convex and 
resemble the species described as schwarzii. The form of the anterior 
tibial spurs will separate the two. These specimens and those from 
Colorado have the sides of the pronotum more strongly declivous, so 
that the marginal head can not be seen from above when viewed 
vertically. The relationships of these forms are really dubious, but 
the material at hand does not warrant their separation as distinct 
species. 

I have seen specimens of the granulato-muricata form labeled 
granulata; the latter species belongs to another subgenus and_ is 
different altogether. 

Rileyi is smaller, more sparsely punctate, more convex in form, and 
undoubtedly more closely related to Awmeralis than to any other 
species of the subgenus. 

The mentum is of moderate size, rather parabolic in outline in the 
typical form, and more triangulo-trapezoidal in the other forms; 
the surface is coarsely but not very densely punctate, moderately 
convex and scarcely foveate laterally; sete very short. 

The prosternum is usually quite semicircularly convex antero- 
posteriorly between the cox; sometimes feebly submucronate or 
rarely vertically subtruncate behind. 

The mesosternum is more or less feebly oblique and concave. 

The intercoxal process of the abdomen is slightly transverse in both 
sexes, and about one-third (male) or a sixth (female) of its width 
wider than the metasternal salient. 

The metasternum laterally between the cox is as long as the width 
of a metatibia at apex. 

The post-coxal portion of the first abdominal segment is equal in 
length to that of the process, and in the female to the length of the 
third segment. 


100 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


In the male the second segment is a little longer than the post-coxal 
part of the first and twice as long as the fourth.» The third is about 
one-half longer than the fourth. 

In the female the second segment is about two and one-half times 
longer than the fourth. 

The profemora are rather feebly and evenly tumid, scarcely clavate; 
the grooves have well-defined subasperulate margins that become 
contiguous at basal fourth. 

The meso- and metafemora are scarcely at all widened externally. 
‘The superior and inferior surface lines are quite parallel. The tibial 
grooves of the mesofemora are quite well defined by asperulate mar- 
gins that become evanescent before becoming contiguous. On the 
metafemora the grooves are similar, but evanescent at middle. 

In the typical form the tibie are rather slender, and stouter in the 
other forms. The tarsal grooves are obsolete and the articular cavi- 
ties closed. The tibize are usually not at all arcuate. 

The tarsi are moderate in length and stoutness. 

The protarsi in the male are apparently a little longer than in the 
female; about two-sevenths of their length shorter than a mesotarsus. 
Joints two, three, and four are subequal, and together about equal to 
the fifth; the first is scarcely as long as the second and third taken 
together. 

The mesotarsi are about a fifth of their length shorter than a meta- 
tarsus. Joints two, three, and four slightly decrease in length in the 
order named, and together a little longer than the fifth; the first is 
quite equal to the combined lengths of the second and third. 

The metatarsi are about five-eighths of their length shorter than a 
metatibia. Joints two and three are subequal, and together equal to 
the first, the fourth being slightly shorter. 


ELEODES RILEYI Casey. 
Eleodes rileyi Casrey, Annals N. Y. Acad. Sci., VI, Nov., 1891, p. 57. 


Rather robust and convex, somewhat strongly shining, intense black 
throughout. //ead scarcely twice as wide as long, feebly convex, 
rather broadly and more or less impressed laterally, frontal suture 
usually distinct, bisinuate, and more or less impressed, coarsely and 
‘ather closely punctate; punctures sparser on the vertex, rather 
coarser on the epistoma. Antennw subequal in length to the head and 
prothorax, the third joint three times as long as wide and rather 
longer than the next two combined; the outer joints but moderately 
robust. 

Pronotum from one-fourth to three-fifths wider than long, and 
widest a little before the middle; disc moderately and evenly convex, 
arcuately declivous at the sides, rather coarsely, deeply punctate, 


ee ee cer 


REVISION OF ELEODIINI—BLAISDELL. 101 


densely so toward the sides, sparsely and unevenly, with large im- 
punctate patches toward the middle; apex broadly and feebly emar- 
ginate, more or less obsoletely and finely margined; sides strongly 
arcuate at apical third, thence strongly convergent and nearly straight 
to the apex; distinctly sinuate before the base, finely and distinctly 
beaded; dase transverse, generally very broadly, feebly sinuate 
toward the middle, two-ninths to one-fourth wider than the apex, 
rather coarsely margined laterally, more or less obsoletely so*at 
middle; apical angles obtuse, but not at all rounded; basal angles 
slightly obtuse, not much rounded and rather prominent. 

Propleure sparsely, irregularly punctate and more or less rugulose. 

Elytra two and one-half times longer than the prothorax, and, in 
the middle, from one-fifth to nearly one-half wider than the latter, 
widest at the middle; dase truncate or feebly emarginate; humeri 
narrowly rounded, slightly prominent, and usually distinctly exposed ; 
sides evenly arcuate, apex narrowly rounded ; disc moderately convex 
on the dorsum, rather evenly rounded laterally, and arcuately decliv- 
ous behind, rather coarsely, feebly and subasperately punctate, some- 
times with very feebly impressed distant lines, the punctures rather 
sparse, not much denser laterally, confusedly disposed, but often hav- 
ing a feeble lineal arrangement. 

Epipleure moderate in width, gradually narrowing from base to 
apex, superior margin feebly and broadly sinuate beneath the 
humeri; surface finely, sparsely, and rather obsoletely punctate. 

Sterna densely punctate and rugulose. 

Parapleure densely punctate. 

Abdomen more or less shining, sparsely punctate and rugulose; first 
segment densely and strongly sculptured, and the fifth more coarsely 
and densely punctate. 

Legs rather short, anterior femora mutic; anterior tibial spurs 
extremely unequal, the anterior spur robust, long, subparallel, and the 
posterior spur very small and acute. Anterior tarsi with the first 
joint feebly thickened at tip beneath, with tuft of ordinary spinules 
more or less feebly developed. 

Male—Oblong-ovate, not noticeably slender. Elytra arcuately 
and somewhat obliquely declivous behind. Abdomen distinctly 
oblique and moderately convex, more or less broadly impressed on 
first two segments. Anterior spur of the anterior tibize about one- 
half as wide as that of the female, and somewhat gradually narrowed 
in apical half, and acute. 

Female.—Robust, ovate. Elytra arcuately and vertically declivous 
behind. Abdomen horizontal and rather strongly convex. Anterior 
spur of the anterior tibiz much broadened, and narrowed in apical 
fourth, and subacute. 

Measurements.—Length, 12-15 mm.; width, 5.2-7.2 mm. 


102 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Genital characters, male. 
feebly arched. ~ 

Basale oblong, about three times longer than wide; surface evenly 
and moderately convex; sides feebly arcuate. 

Apicale triangular; surface evenly and moderately convex, groove 
semi-membranous and rather broad, extending from near tip to base; 
sides arcuate in basal third, thence broadly and moderately sinuate 
to apex, the latter narrowly rounded; base subtruncate. 

Sternite subparabolic. Each lobe with the apex somewhat ex- 
trorsely terminal; external border rather straight at basal third, 
thence arcuate to apex, the latter somewhat rounded; internal border 
arcuate; surface more or less convex, punctate and setose in apical 
half, setae moderate in length and sparse. Membrane sparsely setose 
across the bottom of the sinus, the latter deep. 

Female.—Genital segment moderate in size, more or Jess trapezoi- 
dal, valves more or less reflexed externally, somewhat feebly deflexed 
at apex, setose; external border rather strongly explanate, often less 
so toward apex. 

Valvula—bDorsal plate oblong, sides slightly converging apically; 
surface rather plane, glabrous, sparsely punctate and setose, sete 
longer in apical moiety; external border usually feebly and broadly 
sinuate from the rather prominent basal angle; apical margin not 
distinct from the external lobe, the latter broadly and not very evenly 
arcuate, very feebly defined from the internal lobe by a minute sinu- 
ation; apex clothed with rather long setz; internal border more or 
less sinuous; internal lobe small. 

Appendage minute within the transverse fossa. 

Superior pudendal membrane longitudinally rugulose and not quite 
reaching the internal apical lobe. 

Ventrolateral surface moderately convex in basal two-thirds, thence 
broadly and feebly concave to apex; submarginal groove broad _ be- 
neath the explanate external border of the dorsal plate and becoming 
continuous with the general concavity; surface sparsely punctate and 
finely setose, sete longer on apex. Internal margins of the valves 
contiguous at basal fifth, subeontiguous apically; fissure rather 
broadly fusiform, and closed in basal half by the inferior pudendal 
membrane. 

Habitat—Arizona (Thomas Casey) ; Idaho (7). 

Number of specimens studied, 2 (male and female). 

Type in Colonel Casey’s collection. 

Ty pe-locality.—Arizona. 

Salient type-characters.—Rather robust. Head coarsely and rather 
closely punctate. Prothorax with apex broadly, feebly emarginate; 
base transverse, very broadly, feebly sinuate toward the middle; sides 
strongly arcuate at apical third, thence strongly convergent and 


Edeagophore elongately ovate, very 


ee 


REVISION OF ELEODIINI—BLAISDELL. 103 


nearly straight to the apex, distinctly sinuate before the base; dise 
rather coarsely, deeply punctate, densely so at the sides, with large 
impunctate patches toward the middle. Elytra with dise rather 
coarsely, feebly, and subasperately punctate, punctures rather sparse, 
not much denser laterally, confusedly disposed (Casey). 

Diagnostic characters—The structure of the anterior tibial spurs 
allies this species with Auwmeralis alone. The two specimens before 
me are smaller and shorter ovate, the punctation is sparser and much 
less asperate. The female resembles a large cordata in outline. 

I am indebted to Colonel Casey for the authentic male in my col- 
lection. 

I have before me a small series of specimens collected at Soda 
Springs, Idaho, which I refer to the present species until a larger 
series shall have been studied. One of the specimens, a male, is more 
strongly convex than the male received from Colonel Casey, while 
the females have the facies of a moderately smooth cordata, and the 
punctuation is slightly coarser, the integuments duller and slightly 
more asperate. One male from Idaho has the anterior spur of the 
anterior tibix much longer and stouter than in that sex of réleyi; in 
a single female from the same region the anterior spur is unusually 
prolonged and attenuated, being about four or five times longer than 
the posterior, which is very short and unusually slender, in the other 
females the anterior spurs are at least three times longer than the 
posterior, much broadened and obtusely rounded at tip; these are 
undoubtedly worn, and those in the female mentioned above with 
unusually prolonged spurs represent the normal condition. I believe 
that a sufficiently large series of ri/eyi would show what relation exists 
between these forms; they can be geographical races of either humer- 
alis or rileyi. The genital characters are the same as in riley. 

In the authentic male of ri/eyi the mentum is moderate and sub- 
triangular in outline, the surface not coarsely punctate, feebly convex, 
and scarcely foveate laterally. 

The prosternum is not at all produced and vertically truncate be- 
hind. The mesosternum is obliquely arcuate and very feebly and 
broadly concave. 

The intercoxal process of the abdomen is evidently slightly trans- 
verse and about a third of its width wider than the metasternal 
salient. 

The metasternum laterally between the cox is as long as the width 
of a mesotibia at basal third. 

Post-coxal part of the first abdominal segment is equal in length 
to that of the process, and also equal to the third segment in length ; 
the second is about one-half longer than the fourth. 

The profemora are clavate; the grooves have distinct margins that 
are asperulate and become contiguous at basal third. 


104 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


The mesofemora are feebly tumid, and the grooves have feebly 
formed margins that are finely muricate, becoming contiguous and 
evanescent at middle. - 

The metafemora have subparallel surface lines, and the grooves are 
as on the mesofemora. 

The tibiw are without tarsal grooves and the articular cavities are 
closed. All of the tarsi are missing on the authentic specimen under 
consideration, except a single metatarsus and that is proportioned as 
in humeralis. 


GROUP II.—TRICOSTATA SECTION. 


The species of this group differ from those of Group I in the 
broad, flattened form, with the elytra truncate at base and the 
humeral angles rectangular; the prothorax is likewise broadened at 
base, with the angles rectangular, overlapping the humeral angles 
of the elytra. The pronotal disc is much less declivous laterally and 
the marginal bead is always visible when the surface is viewed verti- 
cally from above; there is consequently scarcely any variation in the 
characteristic pronotal outline. 

The anterior spurs of the anterior tibiz are distinctly thickened and 
broadened in both sexes, less so in the male where they are always 
more so than in the same sex of the preceding group. 

The first joint of the anterior tarsi is always more. or less thickened 
at tip beneath and even slightly produced; a tuft is also evident. 

In the females the dorsal plates of the genital valves are broad and 
oblong. (See Plate 1, fig. 11, and Plate 3, fig. 20.) 

There is also a development of pubescence in one species at least. 
The hairs are setiform. 


ELEODES TRICOSTATA Say. 


Blaps tricostata Say, Journ. Acad. Nat. Sci. Phila., III, 1823, p. 262. 

Bleodes tricostata LECoNTE, Proc. Acad. Nat. Sci., Phila., 1858, p. 181.— 
Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 307. 

Bleodes planata Soxrer, Studi Ent., 1848, p. 366. 

Pimelia alternata Kirpy, Fauna Boreali-Amer., IV, 1837, p. 232. 

BRleodes robusta LeContrer, Proc, Acad. Nat. Sci. Phil., 1858, p. 183. 


Oblong to oblong-oval, black, opaque, clothed with short setiform 
hairs, each arising from a puncture. 

[Head moderate, feebly convex, rather finely, more or less densely 
and evenly punctate, punctures more or less sparse on the vertex; 
frontal suture usually defined, surface more or less broadly impressed 
across the fronto-epistomal junction. Antenne moderately short, not 
reaching to the prothoracic base, outer three joints very feebly com- 
pressed, scarcely dilated, third joint hardly as long as the next two 


REVISION OF ELEODIINI—BLAISDELL. 105 


taken together, fourth very slightly longer than the fifth, the latter 
with sixth, seventh, and eighth subequal in length, the eighth feebly 
triangular, ninth and tenth suborbicular, eleventh oval. 

Pronotum widest at the middle, a little less than one-half wider 
than long and.at least twice as wide as the head; disc moderately con- 
vex, with broad, shallow basal igepressions, finely, evenly and rather 
densely punctate, laterally feebly asperate and rugulose; apex evenly 
and somewhat deeply emarginate, marginal head rather fine; sides 
broadly, evenly, and rather strongly arcuate in the anterior four-fifths, 
thence to base oblique or more or less sinuate, marginal bead some- 
times fine, at other times rather coarse; base squarely truncate to 
feebly emarginate, rather coarsely margined and about two-thirds 
wider than the apex; apical angles distinct, not acute, sometimes 
slightly rounded; basal angles usually rectangular, rarely slightly 
rounded or feebly obtuse. 

Propleure quite evenly and rather finely muricately punctate, at 
times more or less rugulose. 

Elytra oblong-oval, usually widest at the base, sometimes at the 
middle; base truncate, slightly overlapped by and a little wider than 
the base of the prothorax; humeri quite rectangular, overlapped by 
the basal angles of the prothorax; sides subparallel to feebly arcuate, 
more or less arcuately converging in apical third, apex obtuse; dis¢ 
usually flattened, very feebly to moderately convex, laterally sub- 
acutely margined by the subhumeral costae, arcuately. declivous pos- 
teriorly, more or less costate, there being a common sutural, two discal, 
a humeral and subhumeral costa on each elytron, alternating costw 
being usually more or less rudimentary or obsolete, sulci more or less 
shallow ; surface scabrous from rather fine, densely placed muricate 
punctures, which become slightly tuberculiform on the slightly ele- 
vated intervals; inflexed sides inwardly and obliquely declivous, sur- 
face plane and muricately scabrous; pubescence dual, consisting of 
nearly erect brownish setw, and subfusiform, paler, flattened, more 
fragile and semi-reclining subsquamiform hairs. The cost are usu- 
ally more coarsely muricate. 

E’pipleure rather narrow and gradually diminishing in width from 
base to apex, the superior margin rather strongly and broadly sin- 
uate beneath the humeral region; surface evenly and muricately 
punctate. 

Sterna more or less evenly and muricately punctured. 

Parapleure rather finely and asperately punctate. 

Abdomen somewhat shining, finely, submuricately, and rather 
sparsely punctulate, sometimes rugulose. 

Legs somewhat slender. Anterior femora mutic; anterior tibial 
spurs dissimilar in the sexes, the anterior spur curved, larger, and 
longer than the posterior. Anterior tarsi slightly dissimilar in the 


106 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


sexes, first joint slightly thickened and feebly produced at tip be- 
neath with a tuft of spinules interrupting the grogve. 

Male.—Oblong-oval, somewhat elongate. Antenne reaching to the 
posterior fifth of the prothorax. Elytra usually widest at the base 
and thence arcuately narrowing to apex. Abdomen slightly oblique, 
moderately convex, distinctly impressed at base of the first segment 
and between the coxe. Anterior spur of the anterior tibia about one- 
half to one-third longer than the posterior, gradually tapering from 
base to apex, and acute. First joint of each anterior tarsus with a 
small, subacute tuft of yellowish modified pubescence on the produced 
tip beneath; second joint slightly thickened at tip beneath with a 
similar and rather inconspicuous tuft; groove interrupted. 

Female-——Oblong, robust. Antenne reaching to the posterior 
fourth of the prothorax. Elytra broadly oval and usually widest at 
the middle, scarcely narrowed but more or less arcuately rounded pos- 
teriorly. Abdomen horizontal and rather strongly convex. Anterior 
spur of the anterior tibia about twice as long as the posterior, curved, 
distinctly broadened, with sides quite parallel, somewhat narrowing 
at tip. First joint of an anterior tarsus with a tuft of ordinary pic- 
eous spinules on the thickened tip; second joint unmodified. 

The present species is quite variable, and I deem it proper to indi- 
cate four incipient races, as follows: 

Forma typica—Slightly elongate in form, pubescence usually pale 
flavate in color. Elytra distinctly flattened, normal costa well devel- 
oped, the intermediate completely obsolete or very feebly indicated, 
intervals broad and flat. 

Forma ovalis—QOval, short and robust in form. Pubescence 
usually black. Elytra rather strongly convex, normal cost well 
developed and somewhat glabrous and shining, intermediate cost 
more or less evident. 

Forma costata—General form as in typica but smaller. Elytral 
cost more or less strongly and quite equally developed. 

Forma robusta—Form broadly oblong, large. Elytra more or 
less convex, all of the coste# more or less developed, rather coarsely 
muricate, at apex becoming subspiculiferous. 

Measurements.—Males: Length, 13-22.5 mm.; width, 6.5-10.5 mm. 
Females: Length, 14-22.5 mm.; width, 7-11.5 mm. 

Genital characters, male—Edeagophore flaxseed-shaped, slightly 
elongate and not arched. 

Basale oblong, evenly convex and more or less gibbous in basal half}; 


sides evenly arcuate. 

Apicale triangular, slightly elongate, feebly arched; dorsal surface 
evenly convex, with a moderately wide median membranous groove 
in apical half; sides feebly arcuate or nearly straight; apex subacute; 
base more or less broadly arcuate, scarcely sinuate laterally. 


. 
. 


REVISION OF ELEODITNI—BLAISDELL. 107 


Sternite transversely parabolic. Each lobe with the outer border 
evenly arcuate; apex more or less rounded ; internal border more or less 
straight; surface feebly convex, shining, sparsely to rather densely 
punctate, setose, setae not dense and moderate in length. Membrane 
slightly setose across the bottom of the sinus, the latter moderate in 
size. 

Female.—Genital segment quadrate, slightly longer than wide, 
with a slight antero-posterior convexity, setose. 

Valvula (Plate 3, fig. 20).—Dorsal plate oblong, varying more or 
less in width. Surface glabrous and shining, plane to feebly concave, 
sparsely punctate in apical half and along the internal moiety to 
base, sete rather short; sides subparallel; outer lobe of apex not 
differentiated from the apical margin of the plate, evenly rounded, 
well developed, scarcely defined from the very small and hardly visible 
internal lobe. The valvular apex obliquely and inwardly truncate, 
sometimes very slightly sinuate at the inner third, apical area set 
with rather long and flying sete. 

Appendage small and conical, partly visible from above, directed 
backward and more or less inward. 

Superior pudendal membrane long, reaching to the base of the in- 
ternal lobe of the apex, and longitudinally rugulose. 

Ventrolateral surface more or less evenly and moderately convex, 
sparsely punctate and setose, sete rather short; submarginal groove 
arcuately bending inward to the fossa, well defined beneath the 
explanate external border of the dorsal plate and external apical 
lobe, the latter set with quite long flying hairs. Internal margins 
of the valves contiguous in basal fifth, genital fissure broadly fusi- 
form, margins of the valves not closely approximated at apex. In- 
ferior pudendal membrane visible in basal half of the genital fissure. 

Habitat—Texas (Clifton, Bosque County, G. W. Belfrage: San 
Diego, E. A. Schwarz; San Antonio, H. Soltau; Alice, E. A. Schwarz; 
Mobeetee, October); New Mexico (Santa Fé Canyon, 7,000 feet, 
August, F. H. Snow; Albuquerque, February, collector, H. Soltau; 
Las Vegas, August, Barber and Schwarz); Oklahoma (South Me- 
Alester, June, H. F. Wickham); Kansas (Topeka, Hubbard and 
Schwarz; Riley County, May and September, Popenoe; Finney 
County, September, H. W. Menke, collection of Ralph Hopping: 
Onaga); Colorado (Colorado Springs, June and April, H. Soltau; 
Glenwood Springs, June, Hubbard and Schwarz; Denver, H. Soltau; 
Gulnare, Las Animas County, collection of California Academy of 
Sciences; La Junta, Golden, Poudre Canyon, near Long’s Peak, Estes 
Park, Lamar, Sterling, Holly, Fort Collins, La Veta, Veta Pass, 
Wickham’s list) ; Iowa (Sioux City, April, H. Soltau) ; “ Taken at 
Ames by Professor Osborn, while Professor Shimek has taken it in 
Lyon County (June); eastern Emmet County (August); southern 


108 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Dickinson County (August) and eastern Woodbury County (Sep- 
tember) ; it extends as far east as Independence,Buchanan County, 
September, on a broad, dry sand flat, along theWapsipinicon Bot- 
tom” (H. F. Wickham, Iowa Acad. of Sci., 1899); Wyoming 
(Cheyenne, May; collectors, Hubbard and Schwarz); Montana 
(Helena, May; Assiniboine, August; collectors, Hubbard and 
Schwarz) ; Southern Dakota (Sioux Falls, May, H. Soltau) ; British 
America (Medicine Hat, September, C. V. Riley). 

Number of specimens studied, 110. 

Type destroyed. 

Type-locality given by Say, Missouri and Arkansas. 

Salient type-characters.—Body oblong-oval, with numerous minute 
prostrate hairs. Thorax transversely subquadrate, anterior edge 
concave; basal edge rectilinear, covering the base of the elytra; lateral 
edge arcuate; anterior angles rounded; posterior angles excurved. 
Elytra scabrous, with three elevated lines, a common sutural, and a 
lateral one on the edge, intermediate spaces broad, flat, not concave, 
and sometimes with the appearance of a slightly elevated longitudinal 
line alternating with the others (Say). 

Diagnostic characters.—A distinct species, dull in color, sparsely 
pubescent, and with more or less costate elytra. 

In the typical form there is a humeral, a subhumeral, sutural, and 
two discal coste, the intervening cost obsolete or very feebly indi- 
cated, with the intervening spaces flat and wide. 

In the costate form all of the costz are developed and of about 
equal prominence; the individuals are usually under the average 
size and decidedly oblong in form; males more frequently observed. 

The individuals of the oval form are more or less oval and rather 
convex, the coste are normal, strong, and frequently glabrous, the 
pubescence may be blackish or of the normal fulvous color. 

The robust form is a larger, stouter, and decidedly more coarsely 
sculptured variety. Laterally and at apex the murications are quite 
spiculiform. 

The mentum is comparatively small, triangular to trapezoido- 
triangular in general outline; surface feebly convex, more or less 
slightly impressed along the sides at times, rather finely punctate, 
each puncture with a short seta. 

Prosternum comparatively broad between the cox, usually convex 
antero-posteriorly and more or less arcuate or vertical behind, rarely 
subtruncate or submucronate. 

The mesosternum is usually arcuately vertical and more or less 
moderately concave, sometimes feebly so. 

The intercoxal process of the first abdominal segment is quadrate 
and equal in length to the post-coxal part of the same segment; it is 
also equal to the second in length; the third segment is one-half 


ee ee ee 


— =~ 


REVISION OF ELEODITNI—BLAISDELL. 109 


longer than the fourth, the latter slightly more than one-half longer 
than the second (male); in the female the second is slightly longer. 

The metasternal process is about one-fifth narrower (male) to equal 
in width (female) to the abdominal salient. 

The metasternum laterally between the cox is as long as the width 
of a mesotibia at middle. 

The profemora*are moderately robust, slightly tumid, in outline 
elongately oblong-oval, narrowing a little at base; the tibial grooves 
are well marked, concave, glabrous, and shining, margins subcarinate, 
arcuately converging to become contiguous at basal third. 

The meso- and metafemora are subparallel in their surface lines, 
respectively, scarcely at all tumid; the grooves are less strongly 
defined, the margins becoming contiguous and evanescent at near the 
middle. 

The tibiz are without tarsal grooves and muricate throughout; 
the protibial articular cavities are more or less open and slightly 
advanced upon the outer face of the tibia; the meso- and metatibial 
articular cavities are closed. # 

The tarsi are comparatively long and moderately slender. 

In the female of forma typica the protarsi are about one-half of 
their length shorter than a mesotarsus. Joints one to four gradually 
and slightly diminish in size and length; the fifth is about equal in 


length to the second, third, and fourth taken together; the first is 


about equal in length to the second and third taken together. 

The mesotarsi are one-fourth of their length shorter than a meta- 
tarsus. Joints two, three, and four gradually diminish in length 
and size; the first and fourth are subequal in length, and about equal 
to the length of the second and third taken together. Metatarsi 
about one-half of their length shorter than a metatibia. Joints one 
to three gradually diminish in width; first joint about as long as the 
second and third taken together; the fourth is distinctly longer than 
the second. 

ELEODES PEDINOIDES LeConte. 


Bleodes pedinoides LeContr, Proc. Acad. Nat. Sci. Phila., 1858, p. 183. 
Eleodes asperata LeContTeE, Proc. Acad. Nat. Sci. Phila., 1858, p. 183. 
Oblong to somewhat broadly oval, black, elytra frequently more 
or less reddened along the suture, integuments more or less shining. 
Head about twice as wide as long, more or less convex, frequently 
with small, irregular impressions, moderately coarsely but not very 
densely punctate, punctures rather evenly placed on the frons and 
epistoma, vertex with irregular impunctate areas; frontal suture 
usually distinct and more or less impressed. Anfenna@ moderate in 
length and stoutness, outer three or four joints feebly compressed, 
scarcely or very feebly dilated, third joint about equal to the fourth 


110 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


and fifth taken together, the fourth slightly longer than the fifth, 
the latter, with sixth and seventh subequal in length, eighth slightly 
shorter and feebly triangular, ninth and tenth irregularly orbicular, 
eleventh oval to subtrapezoidal and scarcely longer than wide. 

Pronotum widest at the middle and more than twice as wide as 
the head, and about one-half wider than long; disc smooth, moder- 
ately and evenly convex, frequently with small, irregularly placed 
impressions, rather finely and more or less sparsely punctate, with 
rather broad impressions within the basal angles: apea moderately 
and evenly emarginate, usually distinctly margined, bead mod- 
erate; sides evenly, broadly, and rather strongly arcuate in anterior 
five-sixths, thence to base oblique or more or less sinuate, distinctly 
and rather coarsely beaded; base feebly emarginate to truncate and 
rather coarsely margined; apical angles distinct, not acute nor 
rounded; basal angles rectangular or feebly obtuse. 

Propleure rather finely, thickly, and submuricately punctate and 
rugulose. 

Elytra oval, glabrous, not pubescent, usually narrowing at or some- 
times broadest at the base, the latter truncate and more or less sinuate 
laterally, slightly wider than the contiguous prothoracic base; hu- 
meral angles subrectangular, extending under the basal angles of 
the prothorax; s/des more or less moderately and evenly arcuate, 
apex obtuse and not produced; disc depressed on the dorsum, slightly 
convex, sides rather narrowly rounded and inflexed, sometimes 
slightly carinate at humeri, rather evenly and arcuately declivous 
posteriorly; surface feebly sulcato-striate, intervals more ‘or less 
moderately convex, strize submuricately punctate, punctures irregu- 
larly and more or less densely placed; intervals smooth, irregularly 
and more or less sparsely punctate; inflexed sides inwardly oblique 
and feebly convex to plane, not in the least striate, irregularly, 
rather evenly, finely submuricately punctate and rugulose. Occa- 
sionally the punctures at the sides near the apex become strongly 
muricate or subspiculate. } 

Epipleure slightly wider than in fricosteta and gradually dimin- 
ishing in width to apex, the superior margin beneath the humeri quite 
broadly sinuate; surface irregularly and rather finely submuricately 
punctate. 

Sterna more or less glabrous and somewhat obsoletely or finely pune- 
tate and rugulose. 

Parapleure finely, subasperately and rather densely punctate. 

Abdomen more or less glabrous, finely, rather sparsely and more 
or less obsoletely punctulate and rugulose. 

Legs moderate, femora mutic; anterior tibial spurs dissimilar in the 
sexes, the anterior larger and longer than the posterior; tarsi very 


ee 


REVISION OF ELEODITINI—BLAISDELL. 111 


slightly dissimilar in the sexes, first joint of the anterior tarsi slightly 
produced at tip beneath. 

Male.—Oblong, somewhat elongate and moderately narrow, anten- 
ne reaching to the prothoracic base. Elytra moderately and gradu- 
ally narrowed in apical third; abdomen slightly oblique, moderately 
convex from side to side, broadly and indefinitely flattened along the 
middle; intercoxal process more or less concave. Anterior spur of the 
anterior tibia about a third longer than the posterior, scarcely curved 
and moderately thickened, gradually narrowing from base to apex, 
the latter acute. First joint of the anterior tarsi with a small but 
definite tuft of silken pubescence on the thickened tip beneath. See- 
ond joint simple. 

Female.—Oblong-oval, robust. Antenne reaching to the posterior 
fifth of the prothorax. Elytra broadly oval and arcuately narrowing 
in apical fourth. Abdomen horizontal, strongly convex and _ not 
impressed. Anterior spur of the anterior tibia slightly more than 
one-third longer than the posterior, curved, distinctly broadened, sides 
subparallel and rather suddenly narrowed at apex. First joint of the 
anterior tarsi with a tuft of piceous spinules upon the produced tip 
beneath. 

Two forms may be recognized : 

Forma typica—Glabrous and not pubescent. 

Forma asperata. (LeConte.) — Subscabrous. Antenne scarcely 
reaching to the base of the prothorax; sides of the elytra at apex 
decidedly muricate to subspiculate. Legs sparsely clothed with pale, 
flavate, recumbent hairs. 

Measurements.—Males; Length, 19.5-26 mm.; width, 6.5-10 mm. 
Females: Length, 19-27 mm.; width, 8.5-12 mm. 

Genital characters, male-—Edeagophore more or less elongately 
fusiform and feebly arched. 

Basale more or less elongately suboval, strongly convex and more 
er less gibbous in basal half, smooth and shining; sides evenly arcu- 
ate and converging slightly from apex to base. 

Apicale triangular, slightly elongate, evenly convex above with a 
rather broad median membranous groove in apical half; sides feebly 
arcuate to feebly subsinuate in apical half; apex subacute and very 
narrowly rounded; base feebly and broadly lobed at middle and 
feebly sinuate laterally. 

Sternite more or less parabolic in outline, about as long as wide. 
Each lobe with the external margin more or less evenly arcuate, 
sometimes scarcely subangulate at middle; apex subacute to nar- 
rowly rounded; internal margin more or less arcuate; surface feebly 
convex, shining and glabrous, quite regularly and not densely 
punctate, setose, setae, moderate in length and slightly longer in apical 


TE BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


area. Membrane setose across the bottom of the sinus, the latter 
moderate in width. : 

Female.—Genital segment quadrate and somewhat strongly setose. 

Valvula (Plate 1, fig. 11).—Dorsal plate oblong and slightly con- 
vex antero-posteriorly; external border straight to feebly arcuate, 
apically not defined from the apical lobe, with internal border feebly 
arcuate; surface more or less shining, rather coarsely punctate in 
apical half and along the inner moiety to base, setose, setee rather 
long and more or less flying. Apex with the outer lobe well de- 
veloped, evenly rounded and equal in width to the outer two-thirds 
of the dorsal plate, separated from the small and subacute internal 
lobe by a more or less well-marked sinuation, both lobes clothed with 
jong flying hairs. 

Appendages sometimes visible from above, small and mammilli- 
form. 

Superior pudendal membrane long reaching to the base of the small 
internal lobe, and longitudinally rugulose. 

Ventrolateral surface evenly convex, defined from the apex and 
explanate external border of the dorsal plate, by the submarginal 
groove which passes arcuately inward to the fossa; surface sparsely 
punctate and setose, sete rather short and recumbent. The internal 
margins of the valves are contiguous in basal fifth; genital fissure 
quite broadly fusiform with the inferior pudendal membrane visible 
in the basal half. 

Habitat—Texas (Cameron County, H. F. Wickham; Bosque 
County, G. W. Belfrage). 

Number of specimens studied, 29. 

Type in the LeConte collection. 

Type-locality—* Texas;” collected by Messrs. Schott, Haldeman, 
and Weise. 

Salient type-characters——Somewhat shining. Thorax sparsely 
punctulate, slightly convex; sides strongly rounded, posteriorly 
oblique, scarcely sinuate; base truncate; disc with a small impression 
within the basal angles; apical angles with tips rounded. Elytra 
with the dorsum deplanate, not margined; humeri acute and slightly 
prominent; disc sulcato-striate, striz submuricately punctate, inter- 
vals sparsely punctate, towards apex convex and asperate (LeConte.) 

Diagnostic characters—Quickly separated from ftricostata and 
neomexicana by its shining and glabrous integuments. The elytra 
are feebly sulcate with the intervals more or less moderately convex. 

In the asperate form the surface luster is duller and the punctures 
become more or less muricate, at the sides and apex are acute tubercles 
which terminate in short yellowish bristles; the femora are also 
sparsely clothed with recumbent flavate hairs, that are not observed 
in the typical form. 


REVISION OF ELEODIINI—BLAISDELL. 113 


Forma asperata appears to connect ¢ricostata and pedinoides. 

The mentum, pro- and mesosternum do not differ from those of 
tricostata. 

The metasternal and abdominal processes and abdominal segments 
are also as in the above species, although well developed females 
have the second abdominal segment longer and almost twice as long 
as the fourth. 

The femoral and tibial characters are quite similar to those ob- 
served in tricostata. 

The tarsi are comparatively long and a little stouter than in the 
last-named species, and the narrowing of the joints from the first to 
the fourth or third is less evident. 

In the female of the typical form the protarsi are about two-thirds 
of their length shorter than a mesotarsus. Joints two to four are sub- 
equal in length and width; the fifth is a little longer than the third 
and fourth taken together, while the first in distinctly shorter. 

The mesotarsi are one-fifth of their length shorter than a meta- 
tarsus. The total length of the first joint is about equal to the com- 
bined lengths of the second, third, and fourth; the fifth is scarcely as 
long as the second and third taken together. 

The metatarsi are about two-thirds as long as a metatibia. The 
first joint is distinctly longer than the second and third taken to- 
gether, while the fourth is distinctly shorter. 


ELEODES PEDINOIDES var. NEOMEXICANA, new. 


Closely related to pedinoides, from which it differs as follows: 
Surface dull, subopaque. 

Pronotum evenly and moderately convex, basal impressions obso- 
lete; base broadly and not strongly emarginate at middle. 

Elytra with the disk feebly depressed, moderately convex from side 
to side, laterally somewhat more broadly rounded than in pedinoides, 
inflexed portions less sharply defined from the dorsum and slightly 
convex; base emarginate laterally; Awmeri not carinate; surface 
obsoletely sulcato-striate, irregularly and rather densely, finely, sub- 
granulato-muricately punctate, the sculpturing becoming feebly 
asperate about apex. 

Epipleure with the superior margin rather more strongly sinuate 
beneath the humeri; surface rather densely and muricately punc- 
tate. Legs rather less robust. 

Measurements—Females: Length, 23 mm.; width, 11 mm. 

Genital characters—As in pedinoides, except that the dorsal plates 
of the genital segment in the female are comparatively narrower and 


59780—Bull. 63—09——8 


114 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


the superior pudendal membrane slightly shorter, not quite reach- 
ing the base of the internal apical lobe. 

Habitat—New Mexico (Cloudcroft, August, elevation 9,000 feet, 
W. Knaus). 

Number of specimens studied, 5. Monotype a female in my own 
collection. 

Ty pe-locality—Cloudcroft, New Mexico; collector, Warren Knaus. 

Salient type-characters.—Luster subopaque. Pronotal disk with- 
out basal impressions; elytral sculpturing subgranulato-muricate 
and not sulcato-striate. 

Diagnostic characters.—Kasily recognized by the salient type char- 
acters. . 

Differs from tricostata by the more convex elytra without any evi- 
dence of costze. 

Neomewxicana is a distinct and in all probability an extreme varia- 
tion of pedinoides and in no way related to any other known species. 

The asperate form of pedinoides is evidently a connecting link be- 
tween tricostata and the last-named species. For some time I was 
undecided as to the propriety of considering it a race of tricostata, 
but finally decided that it had greater affinity with pedinoides. As 
a race of tricostata it would be analogous to neomexicana. Its form 
is variable, sometimes that of the one and again that of the other 
species; the sculpturing most usually is that of pedinoides, while the 
sparsely placed flavate pubescence is that of tricostata. 

In all probability a large series would corroborate the opinion 
already expressed—that it is a connective form. 

In neomexicana the prosternum is as in pedinoides, although usu- 
ally more or less vertically subtruncate behind, rarely feebly submu- 
cronate. Other sternal, abdominal, and crural characters are as in 
pedinoides. 

The tarsi are less elongate than in pedinoides and about as stout, 
and the narrowing of the joints from the first to the fourth is only 
distinetly evident in the protarsi; the meso- and metatarsi have their 
respective joints mutually subequal in width. 

In the female the protarsi are about one-half of their length shorter 
than a mesotarsus. 

The mesotarsi are one-ninth of their length shorter than a meta- 
tarsus, and a metatarsus about one-half of its length shorter than its 
metatibia. Otherwise as in pedinoides. 


Subgenus LITHELEODES, new. 


Species under moderate size and ovate in form, the elytral sculptur- 
ing varying from smooth to tuberculate. 

The prothorax is usually subquadrate; the anterior femora of the 
males are more or less dentate in part of the species; first joint of 


a a, ee 


REVISION OF ELEODIINI—BLAISDELL. 115 


the anterior tarsi in the male may bear a minute tuft of pubescence at 
tip beneath; in the female the first joint is more or less thickened and 
slightly more prominent ventrally than the others; in both sexes the 
plantar groove is quite entire, except at the extreme apical margin of 
the first joint; it is nearly entire at this point in some males. 

The pubescent tuft is most evident in extricata; less in the other 
species where it is darker in color and frequently quite obsolete. 

The subgenus is founded upon the female genital characters, whach 
are distinctive of the present aggregate. 

Subgeneric genital characters, male—Apicale of the edeagophore 
triangular, sides sinuate, and the apex more or less attenuate; dorsal 
surface more or less longitudinally grooved at middle. 

Female.—Genital segment quadrate to triangulo-parabolic, rather 
thickly clothed with flying hairs. Dorsal plate of each valve oval- 
oblong in outline, with the internal moiety reflexed and impunctate, 
external one-half broadly and slightly reflexed; external border al- 
most straight to more or less arcuate, passing into the apical margin 
abruptly, forming an angle, or more gradually and arcuately so. 
Apex small and membranous. 

Appendage large, more or less semi-elliptical, sometimes subconical, 
flattened, with external border frequently continuing the line of the 
external margin of the dorsal plate. Fossa narrow and transverse. 

Superior pudendal membrane—Usually scarcely rugulose and 
variable in length, long in extricata and letcheri, short in arcuata 
and granulata. 

Distribution —Extricata has a very extensive Ey occur- 
ring in Texas, New Mexico, Arizona, Nevada, Utah, Colorado, Kansas, 
Nebraska, Wyoming, Idaho, Montana, Dakota, and eastern Oregon. 

Arcuata is only known to me from Arizona. 

Granulata, in Oregon, northern and eastern California, in the desert 
regions; and if determinations are correct, in New Mexico and Colo- 
rado. If this be true, it will be found to occur in the intervening 
States. 

Letcheri in western Nevada and vandykei in northeastern Cali- 
fornia and Oregon. 

Genealogy.—Arcuata and extricata have no doubt arisen from a 
common ancestral ramus, and from the other division of the ancestral 
trunk granulata and letcheri have had their origin. 

Eetricata and granulata are no doubt the older stock, and are at 
present diverging at different parts of their distributional area into a 
considerable number of incipient races, and continued collecting and 
study will bring to light and differentiate many more. 


116 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


ANALYTICAL KEY TO SPECIES OF LITHELEODES. 


Integuments smooth and not at all asperate: > 
Anterior femora smutic in the S@xes. = -__—-.=..- 32 2s 3 eee arcuata. 
Integuments more or less asperate : 
Elytra with approximate rows of punctures, which are feebly muricate 
laterally and on the apex; anterior femora armed in the male; thorax 
finely punctate. 


Mlyita DOC RULCT Cs = ee = See ee ee a eI 

Ply tra, SUlLCHte= oe a eee ees _=-----_..-==_Var, (7igonensis; 
Blytra more. or less distinetly tuberculate:_—_—*=- 5. ee granulata, 
Elytra muricately punctate: 

Blytra pubescent =. 22-325 24 ae be eee letcheri. 

Hiytra: not pubescent=.- =~ == a ee ee eee var. vandykei. 


ELEODES ARCUATA Casey. 


Bleodes arcuata Casry, Contr. to the Descr. and Syst. Col. of N. A., Pt. 1, 
1884, p. 47; Annals N. Y. Acad. Sci., V, Nov., 1890, p. 395. 


Oblong-ovate to ovate, robust; shining, glabrous, intense black, 
and very finely sculptured. 

Head twice as wide as long, broadly and evenly convex, frequently 
slightly impressed laterally and along the frontal suture, the latter 
more or less evident and bisinuate; finely and irregularly punctate, 
punctures very sparse on the vertex, becoming coarser and slightly 
denser laterally and on the epistoma, or dense throughout. Antenne 
moderate, reaching to the prothoracic base, subequal in length in the 
sexes, outer four joints scarcely compressed or dilated, third joint 
equal to the next two combined, fourth to the seventh inclusive sub- 
equal, eighth slightly shorter and about as wide as long, ninth and 
tenth suborbicular, eleventh ovate. 

Pronotum subquadrate, widest at or just in advance of the middle, 
one-eighth to one-fifth wider than long; disc moderately convex on 
the dorsum, more strongly so and arcuately declivous laterally, very 
sparsely and more or less minutely punctulate; apex broadly emar- 
ginate, and quite obsoletely margined; sides rather strongly and 
nearly evenly arcuate, most strongly so just in advance of the middle, 
very feebly and minutely sinuate just in advance of the posterior 
angles, frequently most strongly arcuate at the middle and very 
feebly and more broadly, but briefly sinuate behind, marginal bead 
fine; base evenly rounded, equal to the length, one-third to one-fourth 
wider than the apex, very finely or obsoletely margined; apical angles 
acute, basal obtuse, but not at all rounded. 

Propleure more or less sparsely punctulate and irregularly, more 
or less strongly rugulose. 

Elytra oval, widest at the middle, about one-third longer than wide; 
base slightly and evenly emarginate, adapted to the prothoracic 


— 


REVISION OF ELEODITNI—BLAISDELL. 117 


base, to which it is equal in width; humeri obtuse, angle distinct or 
slightly rounded ; s/des evenly and more or less strongly arcuate, apex 
obtusely rounded; disc moderately convex, sometimes slightly de- 
pressed on the dorsum, quite evenly and strongly rounded at the 
sides, strongly, arcuately, and more or less vertically declivous be- 
hind; punctate, punctures small and arranged in regular series, 
intervals each with a row of irregularly spaced smaller punctures, 
punctures confused laterally, sometimes diffusely punctate through- 
out with series scarcely evident. 

Epipleure somewhat wide, scarcely dilated beneath the humeri, 
thence gradually narrowing to apex, superior margin feebly and 
very broadly sinuate in-basal fifth; surface glabrous, at times nearly 
impunctate, often minutely, sparsely and irregularly punctulate. 

Sterna usually shining, irregularly punctate and rugulose. 

Parapleure rather coarsely punctate. 

Abdomen shining, sparsely and quite regularly punctate, at times 
more or less rugulose; sculpturing coarser at middle of the first seg- 
ment, punctures denser and coarser on the fifth segment. 

Legs moderate in length and stoutness, anterior femora feebly 
sinuate and mutic in the sexes; anterior tibial spurs slightly dis- 
similar in the sexes as well as the anterior tarsi. 

Male.—Moderately narrow, oblong ovate. Elytra somewhat rounded 
behind, arcuately and almost vertically declivous posteriorly. Abdo- 
men moderately oblique, moderately convex, broadly and rather 
slightly impressed on the first segment. Anterior tibize with the 
anterior spur slightly longer and a little stouter than the posterior; 
anterior tarsi with the first joint rather simple at tip beneath, the 
margino-apical tufts of spinules not conspicuous, groove somewhat 
entire. 

Female.—Robust, ovate. Elytra more or less strongly arcuate and 
vertically declivous behind. Abdomen horizontal and more or less 
strongly convex; anterior spur of the anterior tibie slightly broad- 
ened, quite acute, and scarcely longer than the posterior; first joint of 
the anterior tarsi moderately thickened at tip beneath and set with 
short piceous spinules, groove obliterated at tip. 

Measurements.—M ales: Length, 12.2-14 mm; width, 5.2-5.5 mm. 
Females; Length, 14-16 mm; width, 6.8-7.2 mm. 

Genital characters.—M ale.—Edeagophore oblong-oval, pointed at 
the apical extremity, slightly arched. 

Basale oblong, surface moderately and evenly convex; sides feebly 
arcuate. ; 

Apicale triangular; surface moderately convex, more or less de- 
pressed on the dorsum, with a moderately narrow median groove at 
middle third; sides arcuate in basal third, thence broadly and moder- 


118 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


ately sinuate to tip; apex feebly attenuate and subacute; base with 
a rounded lobe at middle third, feebly and broadly sinuate laterally. 

Sternite quadrato-parabolic, each lobe with the outer border 

traight and more or less obliquely truncate apically, angle nar- 

rowly rounded; internal border straight; surface feebly convex, 
sparsely punctate and setose, setee moderately long about apex and 
not dense. 

Female—Genital segment subequilaterally triangulo-parabolic, 
rather short and setose. 

Valvula (Plate 4, fig. 25) —Dorsal plate oval-oblong, distinctly con- 
cave; surface glabous and shining, sparsely punctate, each puncture 
with a long seta, internal moiety in apical half impunctate and more 
or less reflexed, outwardly also somewhat reflexed; external border 
evenly arcuate, angle broadly rounded; apical margin oblique pass- 
ing onto the apex, the latter triangular, acute and rather slender; 
angle and apex clothed with quite long flying hairs; internal border 
arcuate apically, sinuate toward base. 

Appendage conical to depresso-conical, moderate in size with-a 
tuft of rather long hairs. Fossa transverse. 

Basal prominences not evident. 

Superior pudendal membrane scarcely to feebly and distantly longi- 
tudinally rugulose, reaching to about the middle of the dorsal plate, 
opaque; valvular membrane closing the intervalvular cleft apically. 

Ventrolateral surface-—Body moderately convex at base and more 
or less transversely concave before the apex; submarginal groove fine. 
Margins of the fossa not fringed with hairs. Apex appears sub- 
laminate with the internal angle of the fossa. 

Internal margins of the valves contiguous, genital fissure very nar- 
row and closed. Inferior pgs membrane not visible. 

Habitat——Arizona (Casey; H. K. Morrison; Chiricahua Moun- 
tains, July, Fort Grant, uly Santa Rita Mountains, May, all in the 
collection of Hubbard and Schwarz; Santa Rita Mountains, 5,000- 
8,000 feet, June, F. H. Snow). 

Number of specimens studied, 24. 

Type is in the collection of Col. Thomas Casey. 

Ty pe-locality.— Arizona. 

Salient type-characters—Form, robust ; shining and ehh very 
finely sculptured. 

Interocular surface broadly and evenly convex; between the eyes 
the punctures are very fine and irregularly scattered, gradually 
increasing in size and density anteriorly; on the epistoma they are 
very close and much larger. 

Prothorax with base one-fourth longer than the apex; sides strongly 
and nearly evenly arcuate, most strongly so just in advance of the 


REVISION OF ELEODITNI—BLAISDELL. 119 


middle, very feebly and minutely sinuate just in advance of the 
posterior angles; apex broadly emarginate, base very broadly and 
feebly arcuate; anterior angles acute, posterior obtuse, but not at all 
rounded; disk moderately convex, minutely granulate, very sparsely 
and somewhat irregularly punctate, punctures very minute. 

Elytra at base as wide as the contiguous pronotum. Surface mi- 
nutely and regularly creased, very minutely and strongly granulated, 
rather feebly and minutely punctate, punctures arranged in regular 
striw, with a single row of smaller punctures between (Casey). 

Diagnostic characters.—Smooth, very finely sculptured, the elytral 
punctures are arranged in widely spaced rows, with rows of smaller 
punctures between, frequently the serial arrangement is lost and 
the punctures are irregular throughout. The anterior femora are 
mutic in both sexes, 

In evtricata the anterior femora are armed in the male, and the 
elytral sculpturing consists of closely placed series of approximate 
punctures, with an interstitial series of more widely spaced punctures, 
frequently the punctures of both series are subequal in size and spac- 
ing and, as a whole, finer or coarser. The humeri are more prominent 
anteriorly. 

Granulata has the elytra sculptured with rows of more or less 
elevated and rather smooth, shining tubercles, and the interspaces 
are opaque. The males have the anterior femora more or less armed. 

Letcheri and vandykei are more elongate and in form more like 
certain smaller Me/aneleodes; the pronotal apical angles are never 
as prominent anteriorly as in the present species. In Jetcheri the 
elytra are sparsely clothed with flying hairs, and in vandykei as well 
the elytral sculpturing is subasperate. Colonel Casey writes that 
the pronotum and elytra are finely granulate, the latter minutely 
creased as well, but these are microscopical characters only seen under 
moderately high power and are of no diagnostic value. Such micro- 
scopical sculpturing can be seen in numerous other species; it is to 
be observed in extricata. 

The differences of width between the abdominal intercoxal salient 
and that of the metasternal salient, as observed in areuata and 
extricata, are evident, as Casey states, and are good structural dif- 
ferential points, but are not to be always absolutely relied upon. 

In “ extricata the intercoxal salient of the first ventral segment is 
about as wide as the intercoxal salient of the metasternum, while in 
arcuata the first is very much wider than the second.” 

I agree with Colonel Casey in considering arcuata a distinct species, 
although related to extricata. The males of the two species are dif- 
ferent. The elytra are more rounded behind as viewed vertically 


120 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


from above and strongly declivous in the former, much more atten- 
uated and pointed, and gradually declivous posteriorly in the latter ; 
anterior femora mutic in the former, armed in the latter. 

Cognata Haldeman is a form of eatricata, and the males have the 
anterior femora armed and therefore distinct from arcuata. 

The large series of some 400 examples of extricata warrant me in 
drawing the above conclusions. Upon the form of the elytra alone 
I would not separate arcuata, but as far as I have been able to com- 
pare males of the two species, those of arcuata are never armed. 

It is possible—as in armata—that extricata may lose the femoral 
teeth in certain examples, and arcuata to gain them; it is either sup- 
pression in the one or atavistic reversion in the other. If it be found 
a’ common occurrence there would still be sufficient characters to dif- 
ferentiate arcuata; the latter might be considered out of place in the 
present subgenus. 

The mentum is more or less parabolic in outline and rather more 
than moderate size; the lateral lobes are small and not usually notice- 
able; surface longitudinally subcarinate at middle and deeply foveate 
laterally, scabrous and feebly setigerous. 

The prosternum is not suddenly protuberant ventrally with the 
coxe, but moderately so and more or less evenly arcuate from the 
anterior border; the intercoxal portion is subhorizontal or moderately 
arcuate, gradually narrowed behind and quite strongly mucronate. 

The mesosternum is more or less arcuately declivous and concave. 

The metasternum laterally between the coxe is as long as the width 
of a mesotibia at apex. 

The abdominal process is feebly transverse (male) or more strongly 
so (female), and one-fifth (male) to one-third of its own width wider 
than the metasternal salient. 

The post-coxal part of the first ventral segment is equal in length 
to the third; the second is about twice as long (male) or distinctly 
twice as long (female) as the fourth. 

The profemora are moderately clavate, the mesofemora just the 
least thickened outwardly; the metafemora are subequal in width 
throughout. 

The tibial grooves are shallow, the margins not strong. 

The tibize are without distinct tarsal grooves, which are at times 
feebly indicated. Articular cavities closed. Tarsi moderate. Pro- 
tarsi stouter in the female. 

Tarsal formula: 


Pro. Meso. Meta. Metatibia. 
Male.—2 23 3 5 
Female.—2 23 3 4} 


ee ees a | 


jn tnt 2 


REVISION OF ELEODITINI—BLAISDELL. 121 


ELEODES EXTRICATA Say. 


Blaps ertricata Say, Journ. Acad. Nat, Sci. Phila., IIIT, 1823, p. 261. 

Bleodes extricata LeContr, Proc. Acad. Nat. Sci. Phila., 1858S, p. 181.—Horn, 
Trans. Amer, Phil. Soc., XIV, 1870, p. 8309.—Casry, Contr. to the 
Descr. and System. Col. of N. Amer., Pt. 1, 1884, p. 48, 

Bleodes cognata HALpeMAN, Stansbury’s Explor. and Survey, Great Salt 
Lake, Utah, Appendix C, p. 376.—Casgry, Annals N. Y. Acad. Sci., V, 
Noy., 1890, p. 395. 


Fusiform-ovate to ovate, about twice as long as wide, smooth, shin- 
ing to subopaque. Anterjor femora armed in the male; the first 
joint of the anterior tarsi is clothed on the slightly thickened tip 
beneath, with a small subacute tuft of yellowish pubescence (fresh 
examples). 

Head twice as wide as long, moderately convex, more or less im- 
pressed laterally and feebly so along the frontal suture, the latter 
usually more or less vaguely indicated; finely and not very densely 
punctate, punctures denser on the sides and epistoma. Antenne 
quite long, outer four joints very feebly compressed and very slightly 
dilated, third joint subequal to the next two taken together, fourth 
slightly longer than the fifth, the latter and sixth subequal, seventh 
subtriangular, eighth triangular, ninth and tenth suborbicular, elev- 
enth ovate. 

Pronotum subquadrate, widest at or slightly in front of the mid- 
dle, about a fourth wider than long; disc more or less moderately 
convex, rather strongly declivous at the apical angles, finely, evenly, 
and not densely punctulate, not denser nor granulate at the sides; 
apex broadly but not strongly emarginate in circular are, or sub- 
truncate, about equal to the length and more or less obsoletely mar- 
gined ; sides evenly and not strongly arcuate, narrowing rather more 
strongly anteriorly than posteriorly, finely beaded; base feebly 
arcuate, one-fifth to one-fourth wider than the apex and very finely 
margined; apical angles somewhat subacute, with the appearance of 
being slightly prominent anteriorly or subrectangular and _ not 
rounded ; basal angles obtuse, slightly or not rounded. 

Propleure moderately convex, finely and sparsely or obsoletely 
punctulate, quite strongly rugulose. 

Elytra oval, widest at the middle, a little more than twice as long 
as wide ; base feebly emarginate, adapted to the prothoracic base, which 
it scarcely exceeds in width; humeri acute and slightly prominent 
anteriorly, usually more or less adapted to the basal angles of the 
prothorax; stdes evenly arcuate, apex rather narrowly rounded and 
not produced; disc moderately convex on the dorsum, strongly and 
more or less broadly rounded laterally, arcuately declivous pos- 
teriorly; more or less coarsely and rather densely punctate, with a 


122 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


tendency to arrangement in approximate rows of closely placed punc- 
tures, with smaller and more widely spaced puné@tures between, most 
evident in the smoother and less so in the duller and rougher forms; 
punctures confused at the sides and scabrous with minute and slightly 
elevated, appressed points which precede the punctures. 

Epipleure moderately narrow, slightly widened at base beneath 
humeri, thence gradually narrowing to apex, superior margin rather 
strongly and broadly sinuate in basal fourth, scarcely visible from 


above at the humeri; surface smooth, finely, sparsely or obsoletely - 


punctulate. 

Sterna more or less irregularly punctate and rugose. 

Parapleure: Mesosternal pieces quite strongly and rather densely 
punctate; metasternal pieces rather sparsely punctate. 

Abdomen more or less shining, finely and sparsely punctate, more 
strongly so and rugulose between the coxee, with ran denser on 
the fifth segment. 

Legs rather slender and moderately long; anterior tibial spurs 
unequal, the anterior a little longer and slightly stouter than the pos- 
terior, both slender and acute: anterior tarsi slightly dissimilar in 
the sexes. 

Male.—Fusiform-ovate, somewhat elongate and more or less nar- 
row. Antenne reaching slightly beyond the base of the prothorax. 
Elytra distinctly and gradually narrowed behind, more or less gradu- 
ally and arcuately or obliquely declivous posteriorly ; abdomen more 
or less slightly oblique, moderately convex, first segment flattened at 
middle, with the intercoxal region more or less concave. Anterior 
femora armed with a subacute tooth; first joint of the anterior tarsi 
clothed at tip beneath, with a small subtruncate tuft of golden 
pubescence—usually darkly discolored; tip not very noticeably thick- 
ened, groove interrupted. 

Female.—Ovate, rather robust. Antenne reaching to the base of the 
prothorax. Elytra arcuately and somewhat vertically declivous pos- 
teriorly, moderately narrowing in apical fourth; abdomen horizontal, 
strongly and evenly convex, intercoxal process convex rather than 
concave. Anterior femora mutic, or at most but very feebly sinuate 
in outer fourth; first joint of the anterior tarsi narrowly and trans- 
versely thickened at tip beneath, with short piceous spinules, groove 
interrupted at the ventral apical margin. . 

In the material before me several forms should be recognized to 
facilitate the correlation of incipient races: 

Forma typica—Shining. Elytra with the approximate rows of 
moderately coarse punctures more or less evident and very feebly 
scabrous; pronotum finely punctulate. Legs moderate. 


REVISION OF ELEODITNI—BLAISDELL. 123 


Measurements —Males: Length, 12-15 mm.; width, 4.5-5.5 mm. 
Females: Length, 12-15 mm.; width, 5-6 mm. 

Forma convexicollis—Form usually small, shorter, and more ro- 
bust. Pronotal disc quite strongly convex, otherwise as in the typical 
form. 

Measurements.—Males: Length, 11-12 mm.; width, 4.2-4.5 mm. 
Females: Length, 11-12.5 mm.; width, 4.5—5 mm. 

Forma cognata typica—Larger, more robust, smoother. Elytral 
punctures fine and less scabrous. Surface shining to dull. 

Measurements —Males: Length, 15-17 mm.; width, 5-6.5 mm. 
Females: Length, 15-17.5 mm.; width, 6.5-8 mm. 

Forma cognata punctata—Pronotal punctures coarse; elytra 
densely and coarsely punctate, otherwise as in the typical cognata 
form. 

Measurements —Males: Length, 12.5-17 mm.; width, 4.5-7 mm. 
Females: Length, 11-17.5 mm.; width, 4.5-8 mm. 

Forma elongata—Male with body tapering both ways so as to 
be quite evenly fusiform, dorsal median line evenly arcuate from 
head to elytral apex; humeri quite prominent anteriorly; thorax 
quadrate, apical angles acute and anteriorly prominent. Legs long 
and slender. Otherwise as in the typical form. 

Measurements—M ales: Length, 14-15.5 mm.; width, 4.5-5 mm. 

Genital characters, male—Edeagophore of the usual form, mod- 
erate in length, elongately oval-oblong and scarcely arched. 

Basale oblong, evenly convex above, with sides feebly and evenly 
arcuate. 

Apicale slightly elongate and triangular, evenly convex above, 
with a short, shallow, linear groove in median third; sides slightly 
arcuate at basal fourth, thence broadly and feebly sinuate to tip, the 
latter acute and moderately produced; base broadly lobed at middle 
and feebly sinuate laterally. 

Sternite transversely suboblong. Each lobe with the outer border 
straight or feebly sinuate in basal half, thence evenly arcuate to apex, 
the latter rather narrowly rounded; internal border straight or sin- 
uate; surface feebly convex, sparsely punctate and setose, sete longer 
but searcely denser about apical margin. Membrane not setose at 
bottom of the sinus. 

Female.—Genital segment rather short, trapezoidal in outline, and 
setose. 

Valvula (Plate 1, fig. 12).—Dorsal plate somewhat oval-oblong, 
inner fourth reflexed, glabrous and impunctate, outer three-fourths 
more or less obliquely and less strongly reflexed, sparsely punctate and 
setose, setee moderatly long and flying; external border very feebly 


124 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


arcuate; internal border quite strongly arcuate; apical margin trans- 
versely or more or less obliquely truncate and sca¥cely defined from 
the apex at internal fourth. Apex small, triangular, and acute at tip. 
finely setose. 

Appendage quite large, strongly flattened, and semielliptical, with 
its external margin directly continuing the line of the external border 
of the dorsal plate; setose, setee short and flying. Fossa transverse 
and narrow. 

Superior pudendal membrane reaching to the apical fourth of the 
dorsal plate, and more or less irregularly rugulose. 

Habitat—F¥orma typica—lKansas; Nebraska; Colorado (Buena 
Vista, July, elevation 7,900 to 8,000 feet; Robertson, Warren Knaus; 
Denver, October, H. Soltau; Colorado Springs; summit of Veta Pass, 
July, C. V. Riley; Gulnare, Las Animas County, July and August, 
Calif. Acad. Sciences). Texas (El Paso). Oklahoma (Mangum, 
January, H. S. Barber) ; Utah; Arizona (Winslow, July, Barber and 
Schwarz; Chiricahua Mountains, June; Fort Grant; Flagstaff, July; 
Humphreys Peak at the base, F. H. Snow). New Mexico (Las 
Vegas, July). Wyoming (Big Horn Mountains, July, R. P. Currie; 
Dead Indian Creek; Cheyenne). Montana; Dakota; Idaho; Eastern 
Oregon; British America (Medicine Hat). 

Forma convexicollis—Wyoming (Lamarie) ; Montana. 

Forma cognata typica—Arizona (Pinal Mountains, Wickham). 
Colorado (Gulnare); Utah; New Mexico (Cloudcroft, 9,000 feet, 
August, Warren Knaus). 

Forma cognata punctata.—New Mexico; Colorado (Edith, May). 

Forma elongata—Arizona (Pinal Mountains, Wickham). 

Number of specimens studied, 500. 

Type destroyed. 

Type-locality—* Country of the Arkansas and Platte” (Say). 

Salient type-characters.—Oblong-subovate. Head acutely pune- 
tured. Thorax subquadrate, punctures smaller than those of the 
head and more distant; lateral edge regularly arcuate; apex very 
slightly emarginate; base nearly rectilinear, a little arcuate each side 
near the angles, which are obtuse. Elytra destitute of striee, or dis- 
tinctly regular series of punctures, scabrous with minute slightly 
elevated points, each of which precedes a puncture (Say). 

Diagnostic characters—The anterior femora are armed in the 
males, and in both sexes the elytra are sculptured with approximate 
rows of rather course punctures, characters not observed in any 
species of M/elaneleodes. In granulata and its races the elytra are 
sculptured with small, flat tubercles arranged in series, with inter- 
vening scattered acute elevated points. In /etcheri the elytra are 
clothed with very sparsely placed, long flying hairs; in vandykei the 


Se 


REVISION OF ELEODITNI-——BLAISDELL. 125 


elytra are more or less coarsely and muricately punctate at the sides 
and not pubescent. 

In typical extricata the pronotum is very finely punctulate; in a 
large series, however, the punctures will be observed to become 
denser and coarser, until in the punctate form there is an approach 
to granulata and letcheré in that respect. 

Arcuata is smooth and finely sculptured throughout and the an- 
terior femora of the males are not dentate. 

In doubtful instances the female genital characters should be 
studied. 

The mentum is rather large in the female and subparabolic in 
outline; smaller and triangulo-parabolic in the male; surface convex 
at middle and more or less excavated laterally, subscabrously punc- 
tate and not noticeably setigerous. 

The prosternum is not strongly protuberant ventrally with the 
coxw, arcuate antero-posteriorly between the coxa and more or less 
strongly mucronate behind. 

The mesosternum is more or less strongly concave and arcuate to a 
varying degree. 

The metasternum laterally between the coxe is as long as the width 
of the mesotibia at apical third. 

The abdominal process is quadrate and equal in width to the 
metasternal salient; in the most pronounced example of cognata 
before me the process is just a little wider than long and also just the 
least wider than the metasternal salient. 

The abdominal process, post-coxal part of the same segment, and 
the third segment are subequal in length; the second is twice the 
length of the fourth. 

The legs are variable in length and stoutness, and what would be 
expected in a very variable species. 

The profemora are more or less clavate, and the tibie are without 
tarsal grooves. 

The tarsi are variable in length and stoutness, longest in the 
elongate form and distinctly stoutest in the punctate form. But 
slightly longer and stouter in the male. 

Tarsal formula: 


Pro. Meso. Meta. Metatibiz. 
Male.—14 2% 3 48 
Female.—14 24 3 43 


ELEODES EXTRICATA var. ARIZONENSIS, new. 


Ovate, more or less glabrous and shining; elytra suleate and mod- 
erately convex. 

Head twice as wide as long, more or less convex, impressed later- 
ally and along the frontal suture, the latter distinct; surface finely, 


126 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


sparsely and irregularly punctate, punctures larger and denser on 
the epistoma. Antenne rather long, outer three joints feebly com- 
pressed, scarcely dilated, third joint scarcely as lone as the next two 
taken together, fourth slightly longer than the fifth, the latter, sixth, 
seventh, and eighth subequal in length, seventh and eighth as wide as 
long, ninth and tenth rather transversely oval, eleventh subovate. 

Pronotum widest at or slightly in advance of the middle, about a 
third wider than long, length equal to width of apex; dise glabrous, 
finely and rather sparsely punctulate, evenly and moderately convex 
from side to side, rather declivous at the apical angles; apex feebly 
emarginate to subtruncate in circular arc, finely and more or less 
obsoletely margined; stdes feebly to moderately arcuate, slightly 
narrowing anteriorly to apex, rather straight and convergent poste- 
riorly to base, very finely beaded; base shghtly arcuate and finely 
margined, about one-sixth wider than the apex; apical angles dis- 
tinct, slightly prominent and not rounded; basal angles obtuse 
scarcely rounded. 

Propleurw usually obsoletely punctulate, quite smooth and dis- 

tinctly rugulose. 
_ Elytra oval, widest at the middle, usually less than twice as long 
as wide; base feebly emarginate, scarcely wider than the contiguous 
prothoracic base; Awmeri more or less acute and rather prominent 
anteriorly; sides evenly arcuate, apex obtuse and rather narrowly 
rounded; disc evenly and moderately convex, feebly depressed, 
strongly and evenly rounded laterally to the epipleure, rather evenly 
and arcuately declivous posteriorly; surface sulcate, the sulci broad 
biseriately, unisefiately, or irregularly punctate, punctures rather 
coarse, intervals distinctly convex, rather narrow, very sparsely or 
more thickly punctulate; inflexed sides obsoletely suleate and more 
irregularly punctate. 

Epipleure obsoletely punctate, glabrous, and moderately narrow. 

Sterna obsoletely punctate and irregularly rugose, more or less 
shining. 

Parapleure subopaque and rather densely punctate. 

Abdomen horizontal; rather glabrous, sparsely, minutely punctate 
and rugulose, more coarsely so on the first ventral segment between 
the coxe. 

Legs quite slender and moderate in length. Anterior tibial spurs 
unequal, both acute; anterior tarsi slightly dissimilar in the sexes. 

Male.—Body slightly elongated and slender. Antenne reaching 
slightly beyond the base of the prothorax. Abdomen very moderately 
convex, intercoxal process slightly flattened; anterior femora armed 
with an obtuse tooth; first joint of the anterior tarsi clothed on the 
slightly produced tip beneath, with a minute tuft of more or less 
golden pubescence. 


REVISION OF ELEODIINI-——BLAISDELL. 127 


Female.—Rather robust, antenne reaching to the base of prothorax. 
Abdomen strongly convex. Anterior femora mutic; first joint of the 
anterior tarsi with groove entire, without the tuft of apical spinules, 
marginal spinules rather stout. 

Measurements —Male: Length, 13 mm.; width, 5.8 mm. Female: 
Length, 14 mm.; width, 6.5 mm. 

Genital characters as in extricata, except that the genital segment 
of the female is subquadrate. 

Habitat—Arizona (Fort Huachuca). 

Number of specimens studied, 6. 

Sexitypes: Male in H. C. Fall’s collection; female in that of my 
own. 

Type-locality—F ort Huachuca, Arizona. 

Diagnostic characters —Form of a typical extricata, with the elytra 
more or less suleate; otherwise as for that species. 


ELEODES GRANULATA LeConte. 


Eleodes granulata LeContr, Reports of Explor. and Surveys... 47th 
and 49th Parallel, XII, Appendix, No, 1, 1857, p. 50, female.—Horn, 
Trans. Amer. Phil. Soc., XIV, 1870, p. 809. 

Eleodes obtusa LeContr, Proc. Acad. Nat. Sci. Phila., 1861, p. 352, male. 

Eleodes aspera LeConte, Smithson. Miscell. Coll., No. 167, 1865, p. 115. 

(?) Bleodes subaspera Sorrer, Studi Entomol., I1, 1848, p. 246. 

Elongate ovate, more or less opaque, elytra tuberculate. 

Head scarcely twice as wide as long, feebly convex and more or 
less flattened, broadly and very feebly impressed laterally, rather 
coarsely, irregular, and quite densely punctate. Antenne rather long, 
outer three joints very feebly compressed and scarcely dilated, third 
joint as long as the next two taken together, the fourth slightly longer 
than the fifth, the latter, sixth and seventh subequal, eighth slightly 
shorter, sometimes triangular, ninth and tenth slightly transversely 
oval; eleventh ovate. 

Pronotum subquadrate, widest at about the middle, one-fourth to 
one-third wider than long: disc moderately and evenly convex, fre- 
quently quite arcuately declivous laterally, rather coarsely and 
densely punctate, punctures very much crowded along the sides, with 
impunctate areas and denser spots centrally: apex truncate to feebly 
emarginate and more or less obsoletely margined; s//es more or less 
evenly and moderately arcuate anteriorly, rather straight and con- 
verging posteriorly, or obsoletely sinuate before the basal angles, 
finely beaded; base subtruncate, finely margined, equal to or slightly 
longer than the length, two-ninths to one-fourth wider than the apex: 
apical angles obtuse, distinct, not prominent nor rounded; basal 
angles obtuse, at times slightly rounded. 

Propleure quite densely sculptured, punctato-rugulose. 


128 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Elytra oval, widest at the middle, less than twice as long as wide; 
base truncate, equal to or shghtly wider than the contiguous protho- 
racic base; humeri obtuse, not rounded, slightfy prominent ante- 
riorly; sides evenly arcuate, apex subacute; disc moderately convex, 
sometimes more or less deplanate, more or less broadly rounded 
laterally, inflexed sides frequently rather plane, arcuately declivous 
posteriorly; tuberculate, tubercles sometimes flattened, frequently 
evenly rounded and moderately elevated, arranged in rows, alternate 
series at times slightly larger, with intervening scattered, acute, 
elevated punctures, interspaces more or less opaque, frequently the 
punctures along the suture are coarsely submuricate, sculpturing 
more confused at the sides. 

E pipleure narrow, scarcely dilated beneath the humeri, and grad- 
ually narrowing to apex, superior margin broadly upeurving in 
basal fifth, not visible from above; surface more or less glabrous, 
finely punctate. : 

Sterna scabrous, densely punctate and rugose. 

Parapleure quite densely punctate. 

Abdomen sparsely punctulate and more or less densely rugulose. 

Legs usually moderate in length; anterior femora dentate or sin- 
uate; anterior tibial spurs quite similar in the sexes, the anterior a 
little longer and larger than the posterior, both slender and acute; 
anterior tarsi feebly dissimilar in the sexes. 

Male—Elongate ovate, somewhat narrow, antenne reaching a 
little beyond the base of the prothorax. Elytra rather gradually 
and arcuately declivous posteriorly, and somewhat attenuate in apical 
third. Abdomen feebly oblique, moderately convex, broadly and 
slightly or scarcely impressed at middle of the first segment. Ante- 
rior femora with an obtuse tooth at junction of the middle and outer 
third, or more or less feebly sinuate in apical fourth; anterior tarsi 
with the first joint scarcely thickened at tip beneath, groove entire, 
apico-marginal tuft of spinules rather stout, and distant. 

Female.—Ovate, rather robust. Antenne reaching about to the 
base of the prothorax. Elytra rather broadly oval, arcuately and at 
times vertically declivous prosteriorly. Abdomen rather strongly 
convex and horizontal. Anterior femora feebly or not sinuate in 
outer fourth; first point of the anterior tarsi feebly thickened at tip 
beneath, groove entire although shallow, apico-marginal tufts of spin- 
ules somewhat approximate. 

Measurements.—M ales; Length, 14.2-17.2 mm.; width, 5.5-6.5 mm, 
Females: Length, 16-17.5 mm.; width, 7.2-7 mm. 

Genital characters, Male—Edeagophore rather elongate, pointed 
oval, not arched. 

Basale oblong, moderately convex, sides parallel. 


REVISION OF ELEODIINI—BLAISDELL. 129 


Apicale triangular, slightly elongate; surface evenly convex, 
obsoletely grooved at middle third; sides in basal half rather 
strongly arcuate, thence somewhat strongly sinuate to tip, the latter 
acute and rather slender; base broadly and feebly rounded at mid- 
dle, feebly sinuate laterally. 

Sternite transverse. Each lobe with the external border \:ather 
broadly and evenly arcuate, continuously so with the apex, angle 
more narrowly rounded: internal margin oblique and somewhat are- 
uate; surface feebly convex, quite densely punctate and setose in 
apical half, sete moderate in length on the apical border. Mem- 
brane not setose at bottom of the sinus, the latter rather broad. 

Female (Plate 4, fig. 28).—Genital segment quadrate, valves more 
or less reflexed externally and quite conspicuously setose. 

Valeula—Dorsal plate suboval, concave, sides reflexed; surface 
somewhat glabrous, reflexed apical two-thirds of the internal border 
and about base impunctate, elsewhere rather sparsely and quite 
strongly punctate, sete rather long; external border more or less 
arcuate, angle subrectangular; apical margin rather broadly sinuate, 
angle with numerous sete; internal margin more or less broadly 
arcuate. Apex triangular and rather long, finely setose. 

Appendage strongly flattened, large and transversely oval, its outer 
margin more or less continuing the line of the external border of the 
dorsal plate, sparsely clothed with rather long sete. Fossa narrow 
and transverse. 

Superior pudendal membrane not rugulose and reaching to the 
middle of the dorsal plate. 

Basal prominences very slight. 

Ventrolateral surfaces more or less convex across the base and 
transversely concave laterally before the apices; surface sparsely 
punctate and finely setose apically; submarginal groove not clearly 
defined, apparently forming the lateral concavity before the apex. 
Genital fissure very narrow, scarcely open; internal margins of the 
valves contiguous basally and apically. Inferior pudendal membrane 
not visible. 

Habitat.—Oregon (collection of LeConte). California (Siskiyou 
County, collections U. S. National Museum and A. Koebele; Los 
Angeles County, D, W. Coquillett). New Mexico? Colorado (Bow- 
ditch) ? 

Number of specimens studied, 12. 

Type (male) in the LeConte collection. 

Type-locality.— Oregon. 

Salient type-characters.—The following description is by LeConte: 

Elongato-ovata, nigra parum nitida, capite thoraceque subsequaliter punctatis, 


hoe subquadrato, minus convexo, latitudine paulo breviore, lateribus rotundatis, 
ad basin vix sinuatim rotundato, angulis posticis obtusis rotundatis. Elytris 


59780—Bull. 683—09——9 


130 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


thorace latiorbus, ovalibus, apice valde declivibus subacutis, dorso minus con- 
vexis seriatim granulatis et parce muricato-punctatis, gersus suturam sub- 
seriatim punctatis; humeris subproductis; prosterno postice producto; femori- 
bus muticis. Long. .58 (LeConte). 

Diagnostic characters.—As I have been unable to gain a definite 
knowledge of the typical form, I have given LeConte’s description in 
full. He states that in form granulata somewhat resembles 2’. guad- 
ricollis, but less convex, and that the sculpturing of the elytra con- 
sists of small, somewhat shining, flat tubercles, arranged in series, 
with intervening scattered acute elevated punctures. He has also 
described two other forms, obtusa from California and aspera from 
New Mexico, both considered by Doctor Horn as synonyms of the 
present species. There are two specimens before me which have been 
compared with the type of obtvsa LeConte by Mr. Blanchard and 
pronounced by him to be that form. LeConte’s description is as 
follows: 

LE, obtusa.—Hlongata, nigra capite thoraceque confertim punctatis, hoc ovato, 
latitudine paulo breviore, lateribus rotundatis, postice obliquis, angulis posticis 
obtusis, spatio parvo levi utrinque ad medium notato, elytris postice oblique 
attenuatis valde declivibus, basi truncatis humeris obtusis, granulis punctisque 
intermixtis vix seriatim positis; femoribus anticis subtus obtuse augulatis. 
Long. .60. 

He also writes that the elytra are punctured toward the suture, but 
behind and at the sides the punctures are replaced by oval elevations 
of moderate size, which are arranged in rows, though not very dis- 
tinctly, and the humeral angles are not produced. 

The examples from Siskiyou County, California, do not answer 
to the above descriptions. Mr. Blanchard has compared them with 
the type and writes as follows: “ Not like the type of granulata, a 
male from Oregon, which is less rough; and obtusa (male from Cali- 
fornia) has a more convex prothorax than granulata, more rounded 
on the sides, and more coarsely and densely punctate, being in these 
particulars more like your Siskiyou specimen. In granulata the dise 
of the elytra is irregularly punctate and rugose ; in obtusa more rugose 
and less evidently punctate.” In another letter he writes that “* Your 
Siskiyou example is perhaps more nearly aspera.” 

If aspera is a synonym of granulata it will be surprising as found 
in New Mexico. I shall again give a description of LeConte’s in full: 

BH, aspera,—Nigra, capite thoraceque opacis, fortiter sat dense punctatis, hoe 
latitudine paulo breviore, obovato modice convexo, apice subemarginato, basi 
paulo sinuata apica haud angustiore, lateribus tenuiter marginatis rotundatis, 
postice subsinuatis, angulis posticis obtusis haud rotundatis; elytris ovalibus, 
thorace duplo latioribus basi emarginatis humeris subacutis, apice valde de- 
clivibus oblique attenuatis, dorso deplanatis, lateribus subito inflexis, undique 
granulis reclinatis nitidis breviter piliferis dense subseriatim positis, et parce 
transversim rugosis; antennis capite thoraceque vix brevioribus, extrorsum 
paulo incrassatis; femoribus anticis muticis, Long. .60, 


— 


— 


REVISION OF ELEODIINI—BLAISDELL. 131 


. 


After a careful review of the literature and a study of the series of 
examples before me, I can come to only one conclusion, and that is 
that there are four incipient races involved and to be recognized. 

The following forms may therefore be indicated : 

Forma typica—Thorax subquadrate, sides rounded, scarcely sin- 
uate toward the base; basal angles obtusely rounded. Elytra seri- 
ately granulate and sparsely muricato-punctate, on each side of the 
suture subseriately punctate; humeri somewhat produced. 

Jt must be borne in mind that the granulate condition is really 
composed of small flatetubercles, with intervening muricate punctures. 

Forma obtusa.—Thorax ovate, sides rounded, oblique posteriorly, 
basal angles obtuse; disc with smooth spaces each side of the middle. 
Elytra with the humeri obtuse, dise punctate about the suture, later- 
ally and about apex with oval elevations of moderate size, which are 
not very distinctly seriate. 

My specimens are opaque and the pronotal punctures are smaller 
than those of the Siskiyou specimens, and distinctly coarser than in 
the examples from New Mexico. 

Forma aspera—Thorax strongly and densely punctate, sides 
rounded and subsinuate behind, basal angles obtuse and not rounded. 
Eytra with the humeri subacute, dorsum somewhat flattened, sculp- 
tured with bright reclinate granules which are briefly piliferous, the 
granules are densely and subseriately arranged, moderately trans- 
versely rugose. 

Forma tuberculata—Thorax as in aspera. Elytra with the hu- 
meri subacute; dise sculptured with distinct tubercles, which usually 
show a serial arrangement, alternate series larger and not at all 
rugose. 

These Siskiyou specimens are larger and more elongate than 
obtusa, and usually of an intense black and feebly shining. The 
pronotal punctuation is like that of U/pis ceramboides, only that the 
punctures are distinctly separated and not coalescent. The elytral 
tuberculation is almost like that observed in granosa. 

The development of the tubercles shows considerable variation in 
the series before me. The interstitial series are the larger, and in 
the row the tubercles vary in size in different examples, in some sub- 
equal in size and rounded, in another some larger and others smaller, 
rounded or oval, and occasionally they appear to coalesce longitudi- 
nally in twos and threes to form an elongate tubercle of about 2 mm. 
in length, several appearing in the same row: the strial series are 
small granules, these may be of uniform size, or some larger and 
others smaller in the same. series; both series become irregular lat- 
erally and on apex, both as regards size and serial arrangement. 
One example has the majority of tubercles of moderate size and 
equal, with few small granules scattered about; another has a dis- 


132 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


tinctly serial arrangement, the tubercles in each series of uniform 
size, the larger very moderate, the strial quite smalk 

In the specimens of obtusa the arrangement is similar and the 
sculpturing more opaque, somewhat eroded or of a worn appear- 
ance. A large series is needed to fully establish the relationships 
between these different forms, and also between them and /Jetcheri 
and vandyhkei. 

Letcheri is of similar form but with the elytra distinctly pubes- 
cent, the hairs are long, sparse and flying; the sculpturing is muri- 
cato-granulate, but not very asperately so, the granular character is 
distinct and the surface luster is dull. 

In vandykei the luster is more shining and the elytra are not at 
all pubescent, but distinctly granulo-muricate, with short, stiff sete, 
the muricate condition predominates, each eminence is a prickle and 
the granular character obscured. The sculpturing is not at all 
coarse, 

Granosa is recognized by its coarser sculpturing and much more 
coarsely and densely rugoso-punctate pronotum. 

The anterior femora are mutic and scarcely sinuate; forms of 
granulata are more or less muricate, granosa less so, and besides the 
integuments are denser and generally opaque, although Nevada 
examples are shining. 

Solier named a species swbaspera, and it is doubtful whether it is 
the present species or not; Dr. George Horn places it as a doubtful 
synonym of granulata, and writes that if it should prove identical 
the present name must be suppressed and Solier’s name be reinstated. 
I have not been able to settle this point. 

The mentum is moderately large in the females of the tuberculate 
form, smaller in the male; rather small in obtusa. Usually subtri- 
angular, rarely subparabolic and more or less scabrously sculptured, 
scarcely convex, usually somewhat broadly and feebly excavated, 
and not noticeably setigerous. Lateral lobes small and inconspicuous. 

The prosternum is more or less convex between the coxee, feebly 
produced and more or less vertically truncate behind, angle some- 
times mucroid, that is with an angle having the shape of a mucro. In 
the type of granulata the prosternum is produced behind and slightly 
prominent posteriorly in obtusa, 

The mesosternum is more or less obliquely arcuate, broadly and 
more or less deeply concave. 

The metasternum laterally between the coxe is as long as the width 
of a mesotibia at apical fifth. 

The abdominal process is quadrate (male) or slightly transverse 
(female) and equal in width to (male) or one-fourth of its width 
wider (female) than the metasternal salient. 


REVISION OF ELEODIINI—BLAISDELL. 133 


The length of the process is about equal to that of the post-coxal 
part of the first segment; the second is twice as long as the fourth 
(male) or a little longer (female), and about one-third (male) to 
one-half (female) longer than the third. 

The profemora are moderately clavate, and in the tuberculate form 
from Siskiyou County, distinctly dentate, the tooth is short and 
broad and almost subacute; in obtusw obtuse and less developed. 

The tibiew are without tarsal grooves. The mesotibix are at times 
rather strongly grooved and the metatibiwe are simply flattened 
externally. 

The tarsi are variable in stoutness and moderate in length. 

In obtusa the tarsal formula is: 


Pro, Meso. Meta. Metatibix. 
Male.—2 3 3 5h 
Female.—2 3 3 5t 


ELEODES LETCHERI, new species. 


Elongate ovate, subopaque, elytra moderately convex and sparsely 
clothed with flying hairs. 

Head feebly convex, vaguely impressed laterally, somewhat finely, 
quite densely and rather evenly punctate. Anfennw moderately long, 
outer four joints very feebly compressed and scarcely widened, third 
joint equal in length to the next two taken together, fourth slightly 
longer than the fifth, the latter, sixth and seventh subequal, eighth 
triangular and about as wide as long, ninth and tenth transversely 
oval, eleventh oval to ovate. 

Pronotum widest at the middle and a little wider than long; disc 
moderately and evenly convex, finely, rather densely, and more or less 
irregularly punctate, the punctures at the sides frequently coalescing 
to form minute rugule; apex feebly emarginate or truncate and very 
finely beaded; sides evenly and not strongly arcuate in anterior two- 
thirds, thence nearly straight or scarcely sinuate to the base and finely 
beaded; base feebly arcuate or truncate, finely margined, and a little 
wider than the apex; apical and basal angles obtuse, not prominent 
nor rounded. 

Propleure finely and not densely muricato-granulate, finely rug- 
lose. 

Elytra oval, not twice as long as wide; base truncate or very feebly 
emarginate, and slightly wider than the contiguous prothoracic base ; 
humeri subacute and not prominent; s/des evenly arcuate, apex 
obtuse and not produced; disc with the dorsum more or less evenly 
convex, more strongly so laterally, and more or less suddenly, 
obliquely or arcuately declivous posteriorly: very irregularly and 
rather densely punctate, minutely scabrous, the anterior lip of each 


134 BULLETIN 638, UNITED STATES NATIONAL MUSEUM. 


puncture raised into a minute point, which become very small muri- 
cate tubercles at the sides and apex; from the pumetures arise rather 
long, erect, dark-brownish and inconspicuous hairs; the pubescence 
absent on the inflexed sides. 

E'pipleurew feebly dilated beneath the humeri at base, narrow and 
gradually narrowing to apex; surface obsoletely and muricately punce- 
tulate. 

Sterna more or less finely and densely, or obsoletely punctate and 
rugulose. 

Parapleurw quite densely punctate, the punctures more or less 
eroded. 

Abdomen finely and rather thickly punctate and rugulose, the 
first ventral segment coarsely and densely punctate. Horizontal. 

Legs moderate in length and rather slender; anterior femora not 
dentate, but more or less feebly sinuate; anterior spurs of the an- 
terior tibix a little longer and thicker than the posterior, both acute, 
scarcely different in the sexes; anterior tarsi feebly dissimilar in the 
sexes. 

Mate. 
the prothoracic base. Abdomen moderately convex; first segment 
broadly impressed between the coxa; second and third segments 
sometimes obsoletely impressed. First joint of the anterior tarsi 
scarcely thickened at tip beneath, groove quite entire, apico-marginal 
tufts of dark spinules rather approximate. 

Female.—Ovate, somewhat robust, antenne reaching to the pro- 
thoracic base. Abdomen strongly convex from side to side. First 
joint of anterior tarsi slightly. and transversely thickened at tip be- 
neath, with short black spinules, groove entire, although nearly 
obsolete at tip. 

Measurements— Males: Length, 12.5-14.5 mm.; width, 4.8-5 mm. 
Females: Length, 13-15 mm.; width, 5.5-6.5 mm. 

Genital characters, male—Edeagophore of the usual form, rather 
broadly fusiform and scarcely arched. 


Basale rather short, not twice as long as wide; surface evenly 


convex, sides evenly and not strongly arcuate. 

Apicale triangular, evenly convex above, and more or less longi- 
tudinally impressed at middle third; sides feebly arcuate in basal 
half and thence feebly sinuate to tip; apex slightly produced and 
acute; base with a median rounded lobe and sinuate laterally. 

Sternite slightly transverse. Each lobe rather narrow, with the 
external border more or less straight in basal half, thence evenly 
arcuate to apical angle, the latter narrowly rounded; surface quite 
strongly and rather densely punctate and setose, setee not dense and 
moderately long; internal border more or less straight. Membrane 
not setose at bottom of the sinus, the latter broad. 


Shghtly narrow, ovate. Antenne reaching slightly beyond: 


Oe 


REVISION OF ELEODITNI—BLAISDELL. 135 


Female.—Genital segment triangulo-quadrate, rather short, shining 
and sparsely setose. 

Valvula (Plate 3, fig. 5).—Dorsal plate suboval to oblong, inner 
third reflexed and outer two-thirds obliquely reflexed; surface chan- 
neled between the obliquely reflexed sides, glabrous, very sparsely 
punctate and setose, sete moderately long and flying; external border 
straight or feebly arcuate; internal border more or less sinuous; 
apical margin somewhat obliquely truncate, angle more or’ less 
rounded; apex short and broadly subtriangular. 

Appendage short and broad, strongly flattened, somewhat semi- 
elliptical, evenly and broadly rounded or more or less emarginate at 
tip, about equal in length to the apex. Fossa transverse and narrow. 
Apex, appendage, margins of the fossa sparsely set with moderately 
long hairs. 

Superior pudendal membrane more or less longitudinally rugulose, 
reaching to about the apical third of the dorsal plate in the median 
line, laterally extending directly to the bases of the apical lobes, and 
appearing triangularly emarginate when the valves are moderately 
abducted. ; 

Ventrolateral surfaces (Plate 3, fig. 6) moderately tumid in the 
basal two-thirds, transversely concave at base of the apices, finely 
and sparsely punctate, setose; submarginal groove not developed. 
Internal margins of the valves contiguous in basal third, genital 
fissure fusiform and not broad. Inferior pudendal membrane not 
well defined. 

Habitat.—I collected a series of specimens at Verdi, Nevada, eleva- 
tion 6,000 feet, in April. They were taken from beneath stones on the 
level land about the town. 

Number of specimens studied, 15. 

Cotypes in my own and Dr. E. C. Van Dyke’s collection. 

Ty pe-locality.—Verdi, Nevada. 

Salient type-characters.—Thorax subquadrate, sides evenly arcuate 
anteriorly, thence nearly straight to base; disc rather densely and 
irregularly punctured; angles obtuse, and not rounded. 

Elytra with the humeri small and subacute, scarcely prominent 
anteriorly; disc finely muricately punctured and sparsely clothed 
with flying hairs. 

Diagnostic characters—Letcheri is unique in having only the 
elytra set with moderately long hairs, it recalls Cratidus osculans, 
but the pubescence is not nearly so conspicuous as in the latter species. 

At first glance it might be taken for a humeralis, especially if the 
pubescence has been removed, but the small size of the anterior tibial 
spurs will quickly separate it from the female of that species; the 
males would be more difficult of separation under such conditions. 
In /etcheri the sides of the pronotum are scarcely sinuate posteriorly 


136 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


and the base is truncate. The elytral base is truncate and the humeri 
scarcely at all prominent anteriorly. ~ 

In humeralis the sides of the pronotum are generally broadly and 
feebly sinuate behind, with the base more or less rounded; the elytral 
base is distinctly emarginate with the humeri more or less prominent 
anteriorly. The two species are found together. with nigrina and 
tenebrosa at Verdi. 

THumeralis is more asperately sculptured. 

In vandykei the elytra are simply very finely setose and the integu- 
ments are shining, in letcheri they are opaque. 

Variations—There are no special variations in the series before 
me. The males have the pronotum more quadrate and the sides more 
evenly rounded; in the females the pronotal sides are more strongly 
arcuate anteriorly and straighter posteriorly (Plate 11, figs. 25 and 
96), and the elytral disc is frequently more depressed, with the sides 
consequently more strongly rounded. The pronotal marginal bead at 
the sides may be obsolete. 

This species is dedicated to my friend, the late Beverly Letcher in 
recognition of many favors. 

The mentum is moderate, and the middle lobe quite evenly rounded 
anteriorly; surface plane and quite evenly punctate. 

The muricate punctures of the elytra when viewed longitudinally 
appear to be subseriately arranged on the sides. 

The prosternum is more or less feebly convex antero-posteriorly 
between the coxee; compressed and vertically truncate behind, with 
the angle usually rectangular. 

Mesosternum arcuate and more or less vertical, broadly and feebly 
concave. 

The tibial grooves of the femora are distinctly concave, with mar- 
gins well defined and finely asperate. The floors of the grooves are 
quite glabrous. 

In the males the anterior femora have the anterior margins of the 
grooves subangulate, thence to the apex sinuate; although the 
femoral tooth is feeble, the species is no doubt related to species with 
dentate anterior femora, like gvanulata for instance. 

The external borders of the anterior tibize, with those of the middle 
and posterior near base, are obsoletely carinate. 

Tarsal formula : 

Pro. Meso. Meta. Metatibiz. 
Male.—23 3 3 5 


Female.—2 5 


-- 
ie 


ELEODES LETCHERI var. VANDYKEI, new. 


More or less shining, elytra not pubescent. Antenne with the third 
joint scarcely as long as the next two combined, fourth joint a little 


REVISION OF ELEODIINI—BLAISDELL. 137 
longer than the fifth, the latter slightly longer than the sixth, the 
latter and seventh equal. 

Pronotum usually widest at the middle, frequently widest just, 
in front of the middle. 

Elytra irregularly and quite densely muricately punctate, very 
minutely so on the dorsum, coarser on the sides and apex; from each 
puncture arises a rather short, stiff, curved, inconspicuous and semi- 
recumbent seta. These are not evident on the inflexed sides. 

Otherwise as in detcheri, but a little more robust. 

Measurements—Males: Length, 14.5-16 mm.; width, 5-6.5 mm. 
Females: Length, 15-16 mm.; width 7.5 mm. 

Genital characters as in letcheri. 

Habitat—Taken in Modoc County, California, by Dr. FE. C. Van 
Dyke. 

Number of specimens studied, 10. 

Cotypes in my own and Dr. Van Dyke’s collection, 

Ty pe-locality.—Modoe County, California. 

Diagnostic characters.—In form and sculpturing resembles Jetcher/, 
but the surface is smoother and more shining. The elytra are not 
pubescent. 

Prosternum more or less strongly convex antero-posteriorly be- 
tween the cox, compressed and vertically truncate as in /etcherd, or 
with a more or less developed mucro, which is at times feebly deflexed. 

Mesosternum as in /etcheré. 

This form is not in the LeConte collection (Blanchard). 

Tarsal formula: 


Pro. Meso. Meta. Metatibie. 
Male.—24 3 3% 6 
Female.—2 3 34 6 


In the U. S. National Museum collection there is a very interesting 
series of this species taken at The Dalles, Oregon, July, by Hubbard 
and Schwarz; the heterotypes are very much smaller but the series 
lead directly up to the typical form of vandykei. The genital char- 
acters are those of the present subgenus. This small form may be 
characterized as follows: 

Forma parvula.—Small, ovate. Pronotum quadrate, strongly and 
quite semi-circularly convex from side to side; sides feebly and quite 
evenly arcuate; apex truncate in circular arc, with angles rectangular: 
base truncate, angles subrectangular. Elytra distinctly asperate with 
muricate granules, densely placed and in somewhat evident longitu- 
dinal series when viewed antero-posteriorly, more or less transversely 
rugulose laterally; humeri acute and feebly prominent anteriorly. 

Antenne in the male long, reaching beyond the prothoracic base, 
scarcely compressed and somewhat incrassate externally. 


138 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Measurements —Male: Length, 9.5 mm., width, 3.5 mm. Female: 
Length, 9 mm.; width, 3.8 mm. 

This small form would no doubt be referred to humeralis upon 
superficial examination. The anterior spur of the anterior tibize in 
the female is comparatively stouter than in the other species of the 
subgenus. Such an acute observer as Mr. Blanchard referred it to 
“near humeralis.” Without recourse to examination of the genital 
segments it would be very difficult to do otherwise; the sculpturing 
is quite like that of the latter species. 

I take great pleasure in giving the species the name of my col- 
league, Dr. E. C. Van Dyke. 


, 
Subgenus TRICHELEODES, new. 


Pubescent species, body clothed with moderately long and more 
or less erect hairs. The pronotum is coarsely, densely, and more or 
less rugosely punctate; the humeri are scarcely evident and rounded: 
the sculpturing is more or less punctato-muricate, becoming muri- 
cately tuberculate in some heterotypes. 

Subgeneric genital characters, male.—Apical lobe of the edeago- 
phore triangular with a median longitudinal impression. 

Female.—Genital segment subequilaterally triangular. Valvula 
with the dorsal plate arcuate at the external border, apical angle 
obsolete, internal apical lobe well developed. Appendage short mam- 
milliform. Superior pudendal membrane reaching to the middle of 
the dorsal plate. 

The abdomen is horizontal in the male, and in both sexes the first 
ventral segment is about as long as the combined lengths of the next 
two. 

Relationships.—The species of the present subgenus in outward 
habitus most nearly approaches Pseudeleodes, and in the muricately 
tuberculate heterotypes this becomes more evident. 

The female genital characters indicate a common origin with 
Litheleodes and Promus. In the former there is also a tendency to 
hirsuteness (/etcher?). A vast amount of material must yet be care- 
fully studied before the present question can be at all satisfactorily 
settled. Increased effort at collecting in the arid regions of the West 
is a desideratum. 

Distribution —The species prefer the arid regions of Nevada, Utah, 
and eastern California, one species extending northward into Oregon, 
and also, according to Linell, into New Mexico. 

The two species of this subgenus may be separated as follows: 
Larger species more or less opaque, hairs of the elytra distinctly unequal in 

lenecth ‘and: slightly bristhysss- 0-6 =e a eee pilosa, 


Smaller species and Amphidora-like in facies; hairs of the elytra shorter, softer, 
and suequall in ength? S24. 222 ee eo a ere hirsuta, 


— 2 


REVISION OF ELEODITNI—BLAISDELL. 139 


ELEODES HIRSUTA LeConte. 


Bleodes hirsuta LeConrer, Proc, Acad. Nat. Sci. Phila., 1S61, p. 552.—Horn, 
Trans. Amer. Phil. Soc., XIV, 1S70, p. 515. 

Ovate, scarcely elongate, about twice as long as wide, clothed with 
black pubescence, coarsely punctate, and more or less shining. 

Head very feebly convex, frontal suture evident; densely, strongly, 
and rather finely punctate, each puncture bearing a more or less erect 
and rather short hair. Antenne rather long, reaching slightly 
beyond the prothoracic base, scarcely compressed in outer four joints, 
feebly incrassate, third joint equal to the next two taken together, 
fourth just noticeably longer than the fifth, the latter, sixth and 
seventh, subequal, eighth about as wide as long, ninth and tenth feebly 
transversely oval; eleventh short ovate. 

Pronotum subquadrate, one-sixth to one-third wider than long, 
widest at the middle; disc rather strongly convex from side to side, 
laterally arcuately declivous, strongly and densely punctate; inter- 
stices more or less glabrous and shining: laterally the punctures more 
or less coalesce; apex truncate, and quite obsoletely margined, about 
equal to the length; sdes evenly and moderately arcuate anteriorly, 
less so, and nearly straight, and slightly converging posteriorly. 
frequently evenly arcuate from apex to base; distinctly margined. 
bead fine; base scarcely arcuate to truncate, very finely and distinctly 
margined, slightly wider than the apex; apical angles obtuse, not 
prominent anteriorly, scarcely rounded; basal angles obtuse. 

Propleure shining and rather coarsely punctate, more or less 
rugose. 

Elytra oval, more or less shining, widest at the middle; base trun- 
cate; humeri rounded and-angle not evident; sides evenly arcuate; 
apex obtuse and rounded, not in the least produced; disc moderately 
convex on the dorsum, more strongly and arcuately rounded laterally, 


_areuately and more or less vertically declivous posteriorly ; surface 


densely, irregularly punctate, finely muricate, but not asperate, 
pubescent; hairs moderate in length and flying, soft and subequal in 
length. 

Epipleure very narrow, gradually narrowing from base to apex: 
just a little wider beneath the humeri than at a point opposite the 
first abdominal suture; surface finely and sparsely punctate, shining. 

Sterna more or less densely punctate and shining. 

Parapleure coarsely punctate. 

Abdomen glabrous and shining; more or less densely punctate, and 
feebly rugulose. Horizontal. 

Legs moderate, rather strongly but not very densely sculptured, 
clothed with short and semirecumbent hairs; anterior tibial spurs 
slender and subequal, those of the mesotibee subequal: the inner spurs 


140 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


of the metatibia slightly longer than the external. Anterior tarsi 
very slightly dissimilar in the sexes; first joint searcely produced 
ventro-apically ; groove distinct. 

Male.—Somewhat narrow, scarcely robust. Elytra about one-third 
longer than wide. Abdomen moderately convex. Anterior tibial 
spurs quite short; first joint of the anterior tarsi with the spinules of 
the apico-marginal angle not noticeably produced. 

Female—Robust. Elytra rather broadly oval, about one-sixth 
longer than wide. Abdomen strongly convex, anterior tibial spurs 
moderately long and slightly curved. First joint of the anterior 
tarsi with the spinules of the apico-marginal angles noticeably pro- 
duced. 

Measurements —Males: Length, 8.5-10.2 mm.; width, 4-5 mm. 
Females: Length, 9.8-12 mm.; width, 5.1-6 mm. 

Genital characters, male-—Edeagophore of the usual general form, 
elongate, depressed and scarcely arched. 

Basale about five times longer than the apicale, oblong; surface 
nearly plane and narrowly, rather suddenly convex at the sides, the 
latter feebly arcuate. 

Apicale short, equilaterally triangular, convex at the sides and 
apex, dorsal surface nearly plane with a median linear, longitudinal. 
membranous groove at middle two-fourths; sides arcuate in basal 
third, thence moderately sinuate to apex, the latter blunt, the lateral 
sinuations giving it the appearance of being produced: base nearly 
evenly arcuate. 

Sternite slightly transverse. Each lobe with the external border 
nearly straight in basal two-thirds, thence evenly arcuate to the 
internal border, angle obsolete; internal margin straight or feebly 
sinuate; surface shining, rather densely punctate in apical two- 
thirds, setose, setee moderately long about apical margin. Sinus deep, 
narrow and triangular. 

Female.—Genital segment subequilaterally triangular, not strongly 
chitinized, setose. 

Valvula.—Dorsal plate rather narrowly and elongately oblong- 
triangular in outline; surface more or less feebly concave, shining, 
sparsely, very finely punctate and setose in apical half; external bor- 
der more or less arcuate; internal border more or less sinuous; apical 
margin not defined from the apex, angle obsolete; apex short and 
triangular. | 


Appendage short conico-mammilliform, fossa small: sete on apex 
and appendage sparse, somewhat flying and slightly longest at tip 
of the appendage. 

Superior pudendal membrane reaching to the middle of the dorsal 
plates and finely longitudinally rugulose. Basal prominences feeble. 

Ventrolateral surfaces as in pilosa. 


REVISION OF ELEODIINI—BLAISDELL. 141 


Habitat—California (Wickham has taken this species at Bodie in 
July, elevation 8,475 feet.) Nevada (specimens in the U. S. National 
Museum collection simply bear the state label—Hubbard and 
Sehwarz; Wickham has taken it at Reno, Carson City, and Queens 
Station). 

A specimen in the Fuchs collection bears a pencil label * Crescent 
City.” I know of only one such locality in Del Norte County, Cali- 
fornia; on the pin also is a smaller label bearing the letters CC. 1 
do not believe that this species occurs in Del Norte County, and in all 
probability the specimen was collected at Carson City, Nevada. 

Number of specimens studied, 7. 

Type a female in the LeConte collection. 

Type-locality.—_Utah—"Great Salt Lake Desert” (LeConte). 

Salient type-characters.—Thorax with the sides rounded, posterior 
angles obtuse. Elytra closely and irregularly punctate, at the sides 
and apex submuricate (LeConte). 

Diagnostic characters—In form hirsuta resembles Amphidora 
nigropilosa. The body surface is shining and at times quite dull, 
especially on the head and thorax. The hairs of the elytra are nearly 
equal in length, and the sculpturing is quite densely, irregularly and 
not asperately, submuricately punctate. 

This species can only be confused with pilosa, from which it 
especially differs in being smaller and more robust, with more broadly 
oval elytra, and in having the sides of the pronotum distinctly 
margined. 

From Jongipilosa it is quickly recognized by the mutic anterior 
femora and simple elytral apices. 

In hirsuta the pronotum is pubescent, in /etcheré it is not. 

Variations—In two examples in the U. S. National Museum 
collection the punctures of the central area of the pronotal disc are 
more sparsely placed and slightly less strongly impressed than usual. 

The mentum is small and triangular. 

The prosternum is variable as in pélosa, and at times horizontal 
between the cox, mucronate behind. 

The mesosternum is more or less obliquely arcuate and quite broadly 
and deeply concave. 

The anterior tibiw have the external borders more or less feebly 
carinate in basal half and thence more or less finely denticulate to 
apex. These characters are less evident in the middle tibiew and nearly 
or quite obsolete in the posterior. The anterior tibia, and the middle 
to a less extent, are more or less feebly arcuate, and at times more 
evidently so. This character is nearly always more pronounced in 
the males. 

“Two males in the LeConte collection have the front and middle 
tibiw the least bit incurved at tip ~ (Blanchard). 


, 
. 


142 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


The edges of the tibial grooves of the femora are usually not at all 
‘arinate, and not well defined, the marginal ling is more or less 
scabrous; the floors of the grooves are slightly glabrous to punctate, 
shining or opaque. 

First joint of the metatarsi equal in length to the next two taken 
together. 

ELEODES PILOSA Horn. 


Bleodes pilosa Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 314. 


Elongate ovate, black, more or less opaque to feebly shining, 
strongly punctate and sparsely clothed with brownish or black, erect, 
‘ather coarse and slightly stiff, conspicuous hairs, 

Head scarcely convex, feebly impressed laterally, rather coarsely 
and densely punctate, sparsely clothed with rather short and more or 
less erect hairs. Antenne moderate, first eight jomts moderately 
pubescent, outer four joints feebly compressed and scarcely dilated, 
third joint equal in length to the next two combined, joints four to 
eight inclusive subequal, eighth feebly triangular, ninth and tenth 
suborbicular, eleventh subacutely ovate. 

Pronotum widest at or just in front of the middle, one-sixth to 
one-half wider than long; disc moderately and evenly convex, 
very coarsely and densely punctate, the punctures irregularly con- 
fluent, the interstices narrow and convex, more or less bright and 
shining, surface clothed with quite long erect hairs; apex truncate or 
feebly emarginate and obsoletely margined; sides not strongly, but 
evenly arcuate in apical two-thirds, thence somewhat straight and 
feebly converging to the base, or very moderately and evenly arcuate 
from apex to base, margin hardly evident to distinct or obsolete ; 
base truncate and finely margined, about equal to the length and a 
little wider than the apex; apical angles obtuse, not rounded nor 
prominent; basal angles obtuse, sometimes feebly rectangular and not 
prominent. 

Propleure opaque, evenly granulo-muricate, with short semi-erect 
sete. 

Elytra oval and about twice as long as wide; base truncate and 
about as wide as the contiguous prothoracic base; hwmeri indistinct 
and rounded; sides evenly arcuate, apex very slightly prominent 
posteriorly and rounded; disc more or less evenly convex from side 
to side, rather suddenly and more or less arcuately declivous pos- 
teriorly; surface densely and irregularly, finely and muricately punc- 
tured on the dorsum, becoming confusedly muricato-tuberculate on 
the sides and apex; sometimes the surface is faintly substriate, at 
other times the punctures on the dorsum are arranged in rather dis- 
tant series, closely placed and unimpressed, with an interstitial series 


REVISION OF ELEODIINI—-BLAISDELL. 143 


of little more distantly and irregularly placed ones; sparsely clothed 
with rather long, bristling. brownish, or black and conspictiously 
erect flying hairs, those of the interstitial punctures being much 
longer than those of the strial series; inflexed sides much less 
pubescent. 

Epipleure quite narrow at base and gradually narrowing to the 
apex; surface opaque and sparsely, submuricately punctulate. 

Sterna more or less densely and submuricately punctate and rugose, 
sete rather short and semi-recumbent. 

Parapleure not densely nor finely punctate, scarcely setose. 

Abdomen shining and horizontal, rather sparsely punctate; first 
segment densely and rather coarsely punctate and rugulose, pune- 
tures denser on the fifth segment. 

Legs moderate in length and not densely clothed with short semi- 
recumbent hairs, that become longer on the tibial apices and_ last 
tarsal joints; anterior femora mutic in both sexes; anterior tarsi 
quite similar in the sexes, apparently not thickened, without tufts 
of pubescence, nor the apico-marginal spinules of the first joint 
being conspicuous. 

Male—FElongate and somewhat narrow, antenne reaching to the 
prothoracie base. Abdomen not strongly convex, broadly and quite 
strongly impressed between the coxie, less strongly on the second 
and third segments. Anterior tibial spurs subequal, moderately 
short, slender, and acute. 

Female.—Rather robust. Antenne not quite reaching to the pro- 
thoracic base, abdomen strongly convex; tibial spurs longer, the 
anterior tibial slightly curved, subequal in length and _ thickness, 
acute. 

Measurements.—M ales: Length, 11.5-16 mm.: width, 5.5-6 mm. 
Females: Length, 15-17mm.; width, 7-7.5 mm. 

I have recently received from Mr. Warren Knaus a pair of speci- 
mens (male and female) that represent an extreme form of sculptur- 
ing, and which is worthy of being recognized as a form, that may be 
defined as follows: 

Forma ordinata—Elytra with rather distant unimpressed strie of 
rather fine and closely placed punctures, each interval with a distinet 
and regular series of rather distantly placed submuricate tubercles, 
feeble but distinct near the suture, each series increasing in size to the 
sides where both series become irregular and confused. Each interval 
has a few small and sparsely scattered punctures between or about 
the tubercles, otherwise as in pé/osa. 

Measurements.— Male; Length, 15 mm.; width, 6.5 mm, Female: 
Length, 15 mm.:; width, 7 mm. 

Genital characters, male—Edeagophore of the usual form, elon- 
gately oblong-ovate, subdepressed not strongly convex nor arched. 


144 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Basale oblong, rather deeply impressed along the middle in apical 
half;'sides feebly and evenly arcuate. ” 

A picale rather elongately triangular; dorsal surface evenly and not 
strongly convex, vaguely and narrowly impressed at the middle; 
sides very feebly sinuate; apex gradually produced and acute; base 
broadly lobed at middle and feebly sinuate laterally. 

Sternite slightly transversely oblong. Each lobe rather narrow 
and somewhat long: external border quite straight in basal half, 
thence evenly arcuate to the apex, the latter rounded; internal border 
sinuate and more or less membranous; surface feebly convex, shining, 
sparsely punctate and setose, sete rather long and dense at apex. 
Sinus rather broad, membrane not setose. 

Female.—Genital segment subequilaterally triangular, with the 
dorsal surface strongly concave, well chitinized and setose, sete short, 
subequal, and sparsely distributed over the apex, appendage, and 
apical moiety of the dorsal plate. 

Valvula (Plate 4, fig. 14) reflexed externally. Dorsal plate ob- 
long-oval; surface evenly concave, glabrous, finely and sparsely 
punctate; external border not well defined from the lateral surface 
towards base, arcuate apically; internal border arcuate and slightly 
reflexed in apical half; apical border continuously arcuate with the 
external and not defined from the apex, angle and external lobe not 
evident; apex short and triangular. 

Appendage short and mammilliform, fully visible from above, 
directed upwards and backwards; fossa not well defined. 

Superior pudendal membrane reaching to the middle of the dorsal 
plate, finely and longitudinally rugulose. 

Basal prominences feeble. 

Ventrolateral surfaces smooth and glabrous, very sparsely, evenly 
punctate and setose, sete short; surface somewhat semicircularly 
convex from side to side, narrowly and rather deeply impressed later- 
ally before the inferior border of the fossa; internal margins of the 
ralves contiguous in basal third. Genital fissure fusiform oval, with 
the inferior pudendal membrane more or less visible in the basal 
moiety. Submarginal groove fine, not very distinct beneath the 
apical half of the external border of the dorsal plate. 

Habitat.—Forma typica—California (Lassen County, Charles 
Fuchs; Owens Valley, Dr. George Horn; Bodie, elevation 8,475 feet, 
in July, and at Amedee, elevation 4,200 feet, H. F. Wickham) ; Ne- 
rada (western, by Mr. Gabb; Carson, Reno, in July, Humboldt Lake 
in June, H. F. Wickham): Oregon (U. S. National Museum Collec- 
tion, C. V. Riley): New Mexico (Coolidge, U. S. National Museum 
collection, M. L. Linell). 

If Linell’s record is correct, pilosa may be looked for in northern 
Arizona, Utah, and southwestern Colorado. 


REVISION OF ELEODITNI—BLAISDELL. 145 


Forma ordinata.—Nevada (Las Vegas, April and May, W. Knaus). 

Number of specimens studied, 16. 

Type in the Horn collection. 

T ype-locality —Owens Valley. California. 

Salient type-characters.—Thorax subquadrate, feebly narrowed be- 
hind, sides moderately rounded, margin hardly evident; apex and 
base truncate, angles not prominent. Elytra with the humeri indis- 
tinct, surface faintly substriate with densely and irregularly placed 
muricate punctures (Horn). 

Diagnostic characters.—The larger and most elongate species of 
the subgenus. Thorax densely and very coarsely punctate; elytra 
coarsely, muricately and asperately punctured laterally and on apex; 
toward the suture the punctures become quite simple. The hairs of 
the pronotum and elytra are long, quite erect, and somewhat bris- 
tling, the interstitial being much longer than the strial hairs. Sur- 
face opaque, integuments frequently incrusted with dirt. 

The above characters serve to separate it from /A/rsuta, the latter 
being smaller in size, more robust, less coarsely but more densely 
punctate and shining. the elytral hairs are shorter, softer, and quite 
equal in length. 

In Jongipilosa the anterior femora are dentate and the elytra cau- 
date, in pilosa mutic and not caudate. 

In letcheri the thorax is not pubescent and not as strongly punc- 
tured. 

In the ordinate form each elytral interval has a regular series of 
submuricate tubercles, with small and irregularly placed punctures. 

Variations.—The largest examples before me are two females col- 
lected in Oregon, and the smallest a male taken at Carson City, 
Nevada. In several females and a few males the pronotal margin is 
wholly absent and the discal surface passes arcuately into the flanks: 
in the greater number of specimens the margin is more or less evi- 
dent, while in others it is distinct and beaded. 

Pronotal variations (Plate 11, figs. 27, 28, and 29) are noticeable in 
the series at hand. Typically the sides are slightly narrowed be- 
hind and moderately rounded. The males show the greatest amount 
of narrowing, and in one example the sides are very broadly and 
feebly sinuate, the basal angles being slightly prominent; in two 
females the sides are distinctly angulate; in both sexes the sides are 
frequently evenly rounded from apex to the base. 

The anterior tibial spurs, although subequal in length and thick- 
ness, appear to vary slightly so as to obscure the true structural rela- 
tionship. At the most the difference is slight. In males the anterior 
spur may appear just the least bit the longer and thicker; the same 


59780—Bull. 63—09——-10 


146 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


characters are observed in several females, while in others the pos- 
terior spur is just noticeably the thicker. ” 

The variation in length can be accounted for by the wear due to 
age, yet the variation in thickness can not be put aside in that way; 
such might be looked upon as fortuitous, and due to the instability 
of certain inherent morphological tendencies. 

In the series before me the general character of the sculpturing is 
quite constant. 

The mentum is variable and comparatively small, triangular with 
apex subtruncate, or somewhat semicircular, the surface is plane to 
foveate laterally with a more or less well-marked median ridges 

The prosternum is more or less feebly or strongly convex antero- 
posteriorly between the coxe, and moderately produced posteriorly, 
usually vertically truncate with angle deflexed and mucronate; rarely 
rounded behind, with mucro obsolete. 

The mesosternum is arcuately declivous and quite broadly and 
deeply concave. Both sterna are clothed with rather long hairs. 
External borders of the anterior tibie are distinctly carinate nearly 
to the apex, and on the middle and posterior tibize nearly to the mid- 
dle, a character more strongly developed here than in hirsuta. 

The anterior tibiz are not noticeably arcuate, although the external 
borders have a gentle and evenly arcuate outline. The tibial grooves 
of the femora are not usually strongly defined, and most always more 
or less subasperately punctate and opaque. In the specimens from 
Oregon the grooves are rather more strongly defined than in any of 
the other examples before me. 

The first joint of the posterior tarsi is equal to the second and third 
taken together; the latter two are of equal length. 


Subgenus PSEUDELEODES, new. 


This subgenus is erected to receive a single species which is remark- 
ably distinct among its congeners. It is based, as usual, mainly upon 
the female genital characters. Although the general facies of the 
species is quite unique, it is extremely difficult to portray in words. 
The general subgeneric characters may be enumerated as follows: 
Coarsely and tuberculately sculptured, not pubescent, humeri obsolete, 
anterior femora mutic; and the anterior tarsi simple beneath. Abdo- 
men on the same plane as the sterna, that is, horizontal. 

Subgeneric genital characters, male—Apicale of the edeagophore 
triangular, evenly convex above, with a median longitudinal groove. 

Female.—Genital segment short and parabolic in outline. 

Valvula—Dorsal plate subobovate to oval, extending apically upon 
the apex beyond the appendage; sides not reflexed. Apex with the 
internal lobe developed. Appendage short, mammilliform, projecting 
from beneath the external border of the apical margin of the dorsal 


a 


REVISION OF ELEODITNI—BLAISDELL. °147 


plate. Superior pudendal membrane reaching to the middle of the 
dorsal plate. Genital fissure subapical. 

General observations.—The internal spurs of the meso- and meta- 
tibiw are slightly longer than the external. 

The tarsi are somewhat stout and moderate in length. The protarsi 
have the four basal joints short, subequal in length, and slightly 
broader than long. The mesotarsi are about one-fourth of their 
iength longer than a protarsus and slightly shorter than a metatarsus; 
the first four joints are subequal—the first undoubtedly slightly the 
longest—and about as wide as long. The metatarsi have the second 
and third joints subequal, both together slightly longer than the first. 

In the male the intercoxal process of the first abdominal segment 
is subquadrate and just slightly transverse and about equal in length 
to the post-coxal part, the latter quite equal in length to the second: 
the third segment is equal to about three-fourths the length of the 
second, and the fourth is equal to two-thirds the length of the third. 

In the female the post-coxal part of the first segment is almost 
equal in length to the second; the third about two-thirds as long as 
the second, the latter slightly more than twice as long as the fourth. 

The intercoxal process of the abdomen is equal in width to the 
length of the third and fourth segments taken together, and one-fifth 
of its own width greater than that of the metasternal salient. 

The metasternum laterally between the coxe in length equals the 
width of a mesofemur at base. 

The epipleure at apices are feebly dilated and concave beneath the 
elytral apex, the latter being very slightly produced. This character . 
is more noticeable in the females. 

Distribution —Granosa inhabits the more arid and desert regions 
of the West. Thus far it has only been recorded from eastern cen- 
tral and northeastern southern California and western Nevada. 

Relationships.—More closely related to Tricheleodes than to any 
other subgeneric aggregate, this affinity is evident in the resemblance 
of facies and in the tendency to development of tubercles in pilosa. 


ELEODES GRANOSA LeConte. 


Bleodes granosa LeConte, Smithson. Mise. Coll, No. 167, 1865, p. 116.— 
Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 314. 

Ovate, opaque, roughly and coarsely sculptured and_ strongly 
convex. 

Head about twice as wide as long, moderately convex, coarsely and 
densely punctate, the punctures at the sides and on the epistoma more 
or less coalescent, and finely setigerous. Antenne rather short and 
moderately stout, outer four joints scarcely compressed, gradually 
and very feebly dilated, third joint about equal in length to the next 
two taken together, fourth scarcely longer than the fifth, the latter, 


148 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


sixth, and seventh subequal, eighth as wide as long, ninth and tenth 
feebly transverse and oval, eleventh subovate. : 

Pronotum subquadrate, widest at the middle, one-seventh to one- 
fourth wider than long; disc rather strongly and evenly convex, de- 
clivous at the sides, coarsely, densely, and quite evenly punctate, the 
punctures at times more or less confluent and strongly impressed, 
becoming muricato-granulate laterally; apex truncate and finely or 
obsoletely margined; sédes quite evenly and not strongly arcuate to 
subangulate at the middle, thence quite straight to base, obsoletely 
or finely margined; base truncate or very feebly rounded and finely 
margined, shehtly wider than the apex and about equal to the length; 
apical angles rectangular to obtuse and not rounded, occasionally 
minutely prominent; basal angles subrectangular to obtuse, not 
rounded nor prominent. 

Propleurw granulo-scabrous to muricato-granulate. 

Elytra oval, about or less than twice as long as wide; base truncate 
and equal in width to the contiguous prothoracic base; humeral 
angles absent, rounded; sides evenly arcuate, apex feebly produced 
and subacute; disc evenly and strongly convex and almost vertically 
declivous posteriorly; surface tuberculate, the tubercies arranged in 
rows, consisting of series of small, evenly and closely placed ones, 
with interstitial series of much larger and more distantly placed 
tubercles, those of both series becoming subequal in size at the sides 
and smaller on the apex, and more or less irregular. 

Epipleure more or less concave toward the base, and finely, sparsely 
punctate. ; 

Sterna more or less densely granulo-punctate. 

Parapleure sparsely and not coarsely punctate. 

Abdomen horizontal, quite densely punctate, the punctures be- 
coming somewhat granular at the sides. 

Legs moderate. Femora densely and rather finely punctate, the 
profemora mutic, not sinuate. Anterior tibial spurs about equal in 
size and length, acute. Tarsi similar in the sexes; first joint of the 
anterior tarsi not thickened at tip beneath, apico-marginal tufts of 
spinules acute, grooves entire. 

Male.—Somewhat narrow. Antenne scarcely reaching to the pro- 
thoracic base. Abdomen moderately convex, broadly impressed on 
the first three segments. 

Female.—Robust. Antenne reaching to about the posterior fourth 
of the prothorax. Abdomen evenly and strongly convex, feebly ven- 
tricose. 

Measurements—M ales: Length, 9-17.5 mm.; width, 5-7 mm. 
Females; Length, 17-19 mm.: width, 6-8 mm. 

Genital characters, male—Edeagophore of the usual oblong-ovate 
form, rather small. 


Ee 


REVISION OF ELEODIINI—BLAISDELL. 149 


Basale oblong, quite evenly convex above and not arched. 

Apicale triangular and slightly elongate; evenly convex above, 
with a median longitudinal and membranous groove in the middle 
third; apex produced and quite acute; sides arcuate in basal third, 
thence broadly and feebly sinuate to tip; base with a rounded lobe 
at middle and feebly sinuate laterally. 

Sternite slightly transverse and strongly chitinized. Each lobe 
rather narrowly triangular; external border rather evenly areuate, 
apex narrowly rounded, internal border sinuate; surface glabrous 
in basal third, densely punctate and setose in apical two-thirds, 
sete rather long and dense. Sinus broad and quadrate and the mem- 
brane is not setose. 

Female—Genital segment short and robust, subparabolic in out- 
line. 

Valvula (Plate 4, fig. 13).—Dorsal plate subobovate to suboval, 
about twice as long as wide, sides not reflexed, margins accurate; sur- 
face slightly concave and more or less shining, punctate in apical 
two-thirds; apical margin more or less evenly and strongly rounded, 
often not distinctly defined from the apex, the latter very small and 
short, subacute and narrowly rounded. 

Appendage small, conically mammilliform, and projecting from 
beneath the external border of the apical margin of the dorsal plate. 
Fossa not visible from above, and small. Apex, appendage, and 
apical half of the dorsal plate minutely setose. 

Basal prominences obsolete. 

Superior pudendal membrane reaching to the middle of the dorsal 
plate and longitudinally rugulose. 

Ventrolateral surfaces not inflated, evenly convex, surface lines 
straight viewed longitudinally. Surface rather densely punctate 
and the apex finely setose; submarginal groove not developed: inner 
borders of the valves contiguous in basal third, the genital fissure 
subapical and fusiform with the inferior pudendal membrane visible 
in basal half. 

TMabitat—California (Panamint and Coso valleys, A. Keebele; 
Los Angeles County, D. W. Coquillett: Mount Pinos, Kern County. 
Fordyce Grinnell; Colorado Desert, San Bernardino County, and 
Oak Creek, eastern Kern County, Dr. E. C. Van Dyke). Nevada 
(collected by Wm. M. Gabb, Horn). 

Number of specimens studied, 24. 

Type (male) in the LeConte collection. 

T ype-locality.—* California.” 

Salient characters —Uead and thorax strongly and confluently 
punctate. Thorax subquadrate, sides rounded and not’ margined. 


150 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Elytra strongly convex, sculptured with series of small, bright tu- 
bercles, interstices with small granules uniseriately placed (LeConte). 

Diagnostic characters—One of the most distinct species of the 
genus. The sides of the pronotum are slightly but not very distinctly 
sinuate behind the apical angles. The elytral sculpturing consists of 
a series of well-developed tubercles upon each interval, and series of 
small granules represent the stria. The coarseness of the seulpturing 
is unique and is only approached by that of granulata. The latter 
species has the form of guadricollis, the humeri are distinct and some- 
what produced, and the elytra are sculptured with rows of smooth, 
moderately elevated, and somewhat elongate tubercles; also more or 
less subseriately punctate each side of the suture, the anterior femora 
are more or less sinuate. 

Variations —The post-apical sinuations of the pronotal sides are at 
times scarcely evident in the males. 

The moderate series of examples before me is of special interest 
and value, on account of the extreme forms of sculpturing that are 
included. 

Two specimens from the Panamint and Coso valleys are remark- 
able for the extreme coarseness of the sculpturing (Yorma fortis). 
In each specimen the pronotal disc is densely, deeply, and confluently 
punctate, the interstices smooth and convex. The elytral series of 
tubercles are notably regular and distinct, except at the extreme sides 
where confusion and diminution in size occur; elsewhere the inter- 
stitial tubercles are three times as large as in the specimens from Los 
Angeles County, and completely fill the width of the interval; they 
are very glabrous and shining, very convex to subacute at summit. 
The strial granules are regular and distinct between the crowded 
larger tubercles. These insects are less opaque than usual and the 
largest of the series. 

In the Los Angeles County (Mount Wilson) specimens the prono- 
tal punctures are smaller, denser, and shallower, with the interstices 
narrow, scarcely convex, and distinctly opaque.. The elytral series 
of tubercles are regular on the dorsum, at the sides the tubercles and 
granules become confused and quite equal in size. 

The Kern County specimens present a modification of the typical 
form of sculpturing. The pronotal punctures are moderate in size, 
irregularly placed, frequently with impunctate areas; the interstices 
are flat, smooth, and shining. The general sizes of the sculptural fac- 
tors of the elytra are as in the Southern specimens; at base, along the 
suture, and laterally the tubercles become more or less transversely 
confluent, losing to a greater or less extent their tubercular charac- 
ter, producing a transversely rugose and submuricate effect. 


REVISION OF ELEODITNI——BLAISDELL. 151 


The prontal margin is usually obsolete and the sides of the pro- 
notal dise are continuously convex with the flanks. Ordinarily, the 
margin is finely crenulate. In one specimen the margin is distinctly 
developed in the anterior two-thirds of the pronotum, where it is 
acute and minutely reflexed. 

In the LeConte collection along with the type there is placed a male 
and female. without locality labels and of peculiar form as follows: 
The elytra are sculptured as in granosa, the thorax is very opaque 
with rather close, coarse, and shallow punctures, as if the rugosities 
seen in the type of granosa had been worn down. The thorax is 
much broader, with the anterior angles more prominent, the sides 
slightly sinuate behind; elytra not quite so convex, more elongate 
and more attenuate posteriorly. The foregoing applies to the female 
as compared with the female of granosa. 

The male has distinctly longer antennw, joints four to eight in- 
clusive being longer; elytra but little wider at middle than the pro- 
thorax and consequently appear more convex; the anterior and mid- 
dle tibizx scarcely differ from those of the female. The side margin 
of the pronotum is feebly defined anteriorly and obsolete posteriorly. 

I am indebetd to Mr. Blanchard for the above notes, and as I have 
not seen the specimens I can only surmise that they may be a race of 
granosa. A Coso Valley specimen is more elongate than any other of 
the specimens before me and it is decidedly less convex, the elytra 
being somewhat flattened on the dorsum. In one or two specimens 
the elytra are slightly ventricose. 

Specimens recently collected by Mr. F. W. Nunenmacher at Gold- 
fields, Nevada, are deep black and shining, the sculpturing coarse as 
in the Panamint Valley specimens above referred to. 

The mentum exhibits considerable variation, being moderate in size. 
triangular or subparabolic in outline, convex at middle and excea- 
vated laterally. The prosternum is more or less evenly convex antero- 
posteriorly between the coxa, frequently grooved; mucronate behind. 
the mucro moderate, conical or triangular. 

Mesosternum short, very oblique, and broadly concave. 

The external borders of the anterior tibiz are distinctly carinate 
nearly to the apex, and the middle and posterior tibiew to a greater 
or less extent as well. 

The tibial grooves of the femora are well defined, the margins are 
subasperately punctate and the floor of each is more or less punctate 
and opaque. 

The anterior tarsi are subequal in stoutness and length in the sexes. 


Tarsal formula: 


Pro. Meso Meta. Metatibie. 
Male.—24 3 3 6 


Female,—24 3 34 6 


152 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 
Subgenus PROMUS LeConte. 


The first, or the first and second joints of the anterior tarsi of the 
male, are more or less thickened and sometimes flattened beneath, 
and densely clothed with fine pubescence. 

These characters are most strongly developed in goryi, where the 
pubescence is almost spongy and the tufts are distinctly flattened ; 
the first joint of the mesotarsi also have a similar but smaller tuft. 

In fusiformis, striolata, and opaca the tufts are smaller and less 
strongly flattened, and in énsularis and subnitens the pubescent pads 
are smaller and not flattened; in the former the tuft is pointed and 
in the latter subtruncate, in both the pubescence is slightly longer. 

The form is usually more or less fusiform in outline, but in 7nsu- 
laris the greatest width of the elytra is behind the middle, and the 
insect is more ovate than fusiform. Some examples of svbnitens are 
subovate. | 

The sides of the pronotum nearly always converge anteriorly and 
the elytra become gradually broader in the same or nearly the same 
line with the pronotal sides. This is most pronounced in opaca and 
fusiformis; in striolata, goryi, subnitens, and insularis the sides of 
the pronotum converge more or less posteriorly, so that the lateral 
line is interrupted to a greater or less extent. 

Subgeneric genital characters, male.—Apicale of the edeagophore 
distinctly longer than wide, somewhat depressed, dorsal surface 
moderately convex and marked by a more or less well defined mem- 
branous, median groove; apex acute and moderately attenuated; base 
usually broadly lobed at the middle. 

Female.—Genital segment more or less quadrate, internal lobe of 
the apex well developed, the external being rudimentary. Sides of 
the dorsal plate reflexed with the surface concave between, outwardly 
explanate and more or less deflexed at the periphery. The dorsal 
plate has the external border more or less arcuate, the surface strongly 
punctate, conspicuously and sparsely setose, the sete being quite long 
and flying. 

Appendage more or less strongly flattened, conico-ellipsoidal or 
transversely semiellipsoidal, and usually quite large. 

Superior pudendal membrane reaching to the middle or apical 
third of the dorsal plate, 2nd obsoletely or irregularly longitudinally 
rugulose. 

The genital characters of the female are distinctive and definitely 
characterize the subgenus. An examination of the plates (4 and 
5) will elucidate what has already been said. It will be observed 
that the appendage in insularis is the smallest and most conical in 
the series. 


REVISION OF ELEODITNI—BLAISDELL. 153 


General observations.—The head is usually twice as wide as long, 
and moderate in size; comparatively large in subnitens and gory, 
small in striéolata. 

Comparison and direct measurement show the head to be subject 
to considerable variation in size, even among individuals of the same 
species. 

The internal spurs of the meso- and metatibiee are slightly longer 
than the external. 

The tarsi are the stoutest in subnitens, moderately so in gory, and 
slender in striolata, fusiformis, and opaca; less elongate and feebly 
robust in énsularis. 

After careful comparative measurements of the tarsal joints, I 
can not present any useful facts relative to the comparative lengths 
of the several joints of each tarsus in the different species. 

The following are my notes on the tarsi of subnitens: 

In the protarsi the second, third, and fourth joints combined are 
about equal in length to the fifth; the first is slightly longer than the 
second. 

Mesotarsi.—The first joint is subequal to the third and fourth 
taken together; the second, third, and fourth slightly decreasing in 
length in the order named. 

Metatarsi—The first joint is a little longer than the second and 
third taken together; the second very slightly longer than the third; 
second and third together about equal to the fourth. 

The above gives a fair idea of the ratios between the joints of the 
different tars’. In striolata the first joint of the metatarsi is about 
equal to the second and third taken together; the second and third 
are subequal and together slightly longer than the fourth. 

In the male of gory?, the first joint of the protarsi is much thicker 
than the succeeding ones and as long as the next two taken together. 

The differences are so slight that they could not be utilized in the 
determination of species. 
~ In the males the intercoxal process of the abdomen lies in the 
same horizontal plane that the meta- and mesosterna do, and the abdo- 
men behind the metacoxw is deflexed. As a result, its surface is ob- 
lique to that of the sterna. This character is scarcely evident in opaca, 
striolata, very feeble in insularis, and more or less well pronounced 
in the other species. 

In the females the abdomen is always horizontal and in some in- 
stances slightly prominent ventrally with the sterna. 

The relative length of the abdominal segments has to be considered. 

In the male of ‘nsv/aris the intercoxal process of the first abdominal 
segment is quite quadrate, and about equal in length to the second 
segment; the post-coxal portion is also subequal in length to that 


154 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


of the second, the latter being distinctly twice as long as the fourth; 
the third is about two-thirds as long as the second. 

In the female the abdominal salient is transverse, nearly twice 
as wide as long; the post-coxal portion of the first segment is about 
equal to the second in length; the second is nearly two and one-half 
times as long as the fourth; the third nearly two-thirds as long as 
the second. The first three segments are relatively longer than in 
the male. 

Subnitens—The male has the intercoxal process of the first ventral 
segment just noticeably transverse, and the post-coxal part is scarcely 
as long as the second; the third is slightly less than twice as long as 
the fourth. 

The female has the intercoxal process slightly transverse, and the 
post-coxal portion of the first is about equal to the second in length; 
the third is about three-fourths as long as the second and about twice 
as long as the fourth. 

Goryi.—The intercoxal process in the male is distinctly transverse, 
and the post-coxal part of the first segment is scarcely as long as the 
second, the latter being about one-fourth longer than the third; the 
fourth is about one-half as long as the third. 

The female has the intercoxal salient strongly transverse and about 
equal to the second in length: the post-coxal portion is scarcely as 
long as the second; the third is about two-thirds as long as the 
second and about twice as long as the fourth. 

Striolata——lIn this species the male has the intercoxal process trans- 
verse and rather short, about equal to the fourth in length; the post- 
coxal part of the first ventral segment is about as long as the second, 
and the fourth is searcely one-half as long as the latter. 

The female has the intercoxal process moderately transverse and 
scarcely as long as the third segment, and the post-coxal part of the 
first is nearly as long as the second; the third segment is about three- 
fourths as long as the second, and the fourth is nearly two-thirds as 
long as the third. The first three segments are comparatively long. 

Fusiformis.—The intercoxal process of the male is distinctly quad- 
rate and equal in length to the post-coxal part of the same segment, 
which is subequal in length to the third; the second is about twice as 
long as the fourth segment. 

The female has the segments proportioned about as in the male, 
but slightly longer. The intercoxal process is quadrate. 

Opaca.—The male has the intercoxal salient short and slightly 
transverse, the post-coxal portion is equal in length to the third and 
the second is about twice as long as the fourth. 

The abdominal salient of the female is comparatively small and 
nearly quadrate, and the post-coxal part of the same segment is sub- 


REVISION OF ELEODIINI—BLAISDELL. 155 


equal to the second; the third is about two-thirds as long as the pre- 
ceding segment and one-third longer than the fourth, 

Insularis has the abdominal salient about equal in width to the 
length of the third and fourth segments taken together, and about 
one-fourth of its width broader than the metasternal process. The 
metasternum laterally between the coxe is about as long as the width 
of a metafemur at base. 

Subnitens has the intercoxal process equal in width to the length 
of the second segment and about one-fourth of its own width broader 
than the metasternal process. The metasternum laterally between 
the coxw is as long as the width of a mesotibia at apex. 

The intercoxal abdominal salient in goryi is quite wide and equal 
to the combined lengths of the third and fourth segments and about 
one-third of its width greater than that of the metasternal process. 
The metasternum laterally between the cox is equal in length to the 
width of a mesotibia at apex. 

In striolata the intercoxal process of the abdomen is quite equal in 
width to the length of the second segment and about one-third of its 
width greater than that of the metasternal salient. The metasternum 
is quite short laterally between the cox and about equal in length 
to the width of a mesotibia at middle. In the fermale the abdominal 
process is evidently not as wide as the second segment is long. 

Fusiformis has the abdominal process comparatively narrow, being 
about equal to the length of the third segment and subequal in width 
with the metasternal salient. The metasternum laterally is short and 
about equal to the width of the mesotibia at middle. The same re- 
sults of comparison can be applied to opaca that have been obtained 
from fusiformis. 

The above notes on the relative widths and lengths of the sterna 
and abdominal segments can only be considered in a general way as 
being approximately correct, as there is always more or less indi- 
vidual variation in each species. 

In summing up it will be noticed that the coxe are most widely 
separated, and the intercoxal process of the first abdominal segment 
reaches its greatest width in insularis and subnitens; that in fusi- 
formis and opaca they reach their greatest approximation, and that 
the intercoxal salients are quite equal; that gory? and striolata are 
somewhat intermediate, but really more nearly like the first two 
named species than the last two. 

Distribution —Insularis, as far as known, is an insular and peninsu- 
lar species, inhabiting the southern part of Lower California and an 
adjacent island. 

Subnitens inhabits central and southern Arizona. I know of no 
record of its having been taken in Mexico, but it may occur there. 

Gory? is taken in New Mexico, Texas, and Mexico, 


156 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Striolata is found in southern Texas and the adjoining regions of 

Mexico. “ 
Fusiformis is distributed over an extensive area as follows: Texas, 
New Mexico, Kansas, Colorado, Nebraska, and southern Wyoming. 
Opaca also has a wide distribu- 


a 3 tion, being known to me to occur in 
gets EM Fisiforms Texas, Colorado, Kansas, Nebraska, 
EG and southern Dakota. 

Genealogy.—The results of the com- 


Ler ae parative measurements indicate quite 
aris 


clearly upon what structural lines that 
Opaca “ae 3 
divergence has taken place, while the 
relationships of the species among 
themselves have been sufficiently indi- 
cated. I believe that striolata and 
Fic. 2.—GENEALOGICAL DIAGRAM fusiform is have diverged from a com- 
OF THE SUBGENUS PROMUS. * i 
mon stem much later than did goryi 
from the insularis-subnitens ramus. Knowledge of the Mexican 
species may entirely change my views, as it only at present concerns 
the species in the United States. 
Our six species may be separated as follows: 


Subgeneric Trunk 


Species with first joint of the anterior tarsi in the male conspicuously pubes- 
cent beneath. 
Surface shining; thorax distinctly widest at the middle, visibly narrowed 


at base: tarsal pubescent tuft conical. =_2: 2-222 2 eee insularis. 
Surface dull; thorax scarcely wider before the base; tarsal tuft truncate 
2, a ¢ | aM en es See ts Sei ASAE eel epee ey A I ee eS subnitens. 


Species with first two joints conspicuously spongy pubescent beneath. 
Femora (anterior) armed in the males. 


Elytra with distant series of large dents or punctures___-----~--- goryi. 
Elytra with approximate series of rather small and closely placed 
PUNCTULCS = =e Ale ee ee ee striolata, 
Femora mutic in the males. 
Elytral margins rounded, surface glabrous, not pubescent_____ fusiformis. 
Elytral margins acute, surface opaque and pubescent___-_--------~ opaca,. 


ELEODES INSULARIS Linell. 
Bleodes insularis LINELL, Proc. Ent. Soc. Wash., IV., No. 3, p. 180 (author's 
extras published March 7, 1899). 

Elongate, subovate, black, smooth, more or less shining and mod- 
erately convex. 

Head more or less convex, rather finely, more or less evenly, or 
irregularly punctate, punctures not dense, but slightly so on the 
epistoma and sides. Antenna moderate in thickness, long, reaching 
beyond the prothoracic base, very feebly compressed and just in the 
least dilated in the outer three joints, third joint about equal to the 
next two taken together, fourth slightly longer than the fifth, the 
latter, sixth and seventh subequal, eighth slightly shorter, ninth 


REVISION OF ELEODITNI—BLAISDELL, 157 


more or less triangular, tenth scarcely transversely oval, eleventh 
suboval. 

Pronotum subquadrate, rather dull, widest at the middle, one- 
fifth to one-fourth wider than long; disc evenly and moderately con- 
vex, very finely, rather evenly but not densely, and more or less 
obsoletely punctulate; apex more or less emarginate, frequently more 
noticeably so laterally, very finely or more or less obsoletely mar- 
gined; sides broadly, evenly and not strongly arcuate, very finely 
beaded; ase more or less arcuate and finely margined, about one- 
fourth to one-third wider than the apex: apical angles subacute, an- 
teriorly prominent and not everted; basal angles obtuse and not 
rounded nor prominent. 

Propleure finely and irregularly punctulate, more densely and 
strongly so posteriorly, and more or less finely rugulose. 

Elytra oval to subovate, usually widest behind the middle; base 
more or less emarginate and not wider than the contiguous pro- 
thoracic base; humeri obtuse, not rounded nor prominent; sides 
slightly more strongly arcuate behind the middle, apex not in the 
least produced and rounded; disc moderately convex on the dorsum, 
strongly and evenly so laterally, rather suddenly and arcuately de- 
clivous posteriorly; finely punctate, punctures arranged in moder- 
ately distant series, rather closely placed and at times lodged in 
shallow and slightly larger dents, intervals more or less obsoletely 
punctulate; the serial punctures at times more or less obsolete. 

Epipleure moderately narrow, gradually narrowing from base to 
apex and more or less impunctate. 

Sterna more or less moderately punctate and rugulose, finely and 
sparsely pubescent. The prosternum is frequently more strongly 
punctate. 

Parapleure sparsely and irregularly punctate, punctures rather 
shallow. 

Abdomen very finely or obsoletely and rather sparsely punctulate, 
more or less finely rugulose; the intercoxal process more or less 
coarsely sculptured. 

Legs moderate in length and slender; anterior femora more or less 
armed in both sexes; tibial spurs small, the anterior subequal and 
acute; anterior tarsi dissimilar in the sexes. 

Male.—Somewhat narrow. Antenne extending three or four joints 
beyond the prothoracic base. Elytra evenly and strongly arcuately 
declivous posteriorly; abdomen slightly oblique, moderately con- 
vex, flattened at the middle on the first three segments and more 
or less grooved in the middle line. Anterior femora with a distinct 
but obtuse tooth; anterior tarsi with the first joint slightly thickened 
and clothed apically beneath with a dense brush of more or less 
golden pubescence. 


158 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Female.—Rather robust. Antenne with about one or two joints 
reaching beyond the prothoracic base. Elytra aycuately and verti- 
‘ally -declivous posteriorly. Abdomen horizontal, strongly convex 
from side to side. Anterior femora obtusely angulate or simply sin- 
uate; anterior tarsi without pubescence beneath. 

Measurements.—M ales: Length, 14.5-17.6 mm.; width, 5.6-6.8 mm. 
Females: Length, 16-16.5 mm.: width, 6.5-7.0 mm. 

Genital characters, male-—Edeagophore flattened oblong-ovate, and 
slightly arched. 

_ Basale moderately long, oblong, quite evenly convex above, with 
sides nearly parallel to feebly arcuate. 

Apicale triangular, slightly elongate and feebly decurved; surface 
evenly and moderately convex, with a moderately wide membrano- 
chitinous depression at middle third; sides feebly arcuate near the 
base, thence to the tip slightly sinuate; apex somewhat produced and 
subacute; base broadly lobed at middle and slightly sinuate laterally. 

Sternite transverse and setose. Each lobe rather broadly triangular. 
with the external border quite evenly and broadly arcuate; internal 
border more or less arcuate; apex subacute to narrowly rounded: sur- 
face quite evenly convex, rather evenly and not densely punctate in 
apical three-fourths, setee moderately long. Sinus narrow and the 
membrane not setose. 

Female.—Genital segment quadrato-parabolic in outline, quite 
strongly convex antero-posteriorly so that the apical moiety appears 
quite strongly and arcuately declivous apically, setose. 

Valvula (Plate 4, fig. 22).—Dorsal plate obovate, quite strongly 
arcuate antero-posteriorly, broad in apical half and narrowing to 
base, inner third of the apical half quite strongly reflexed; surface 
somewhat shining, rather densely punctate and setose, sete rather 
long and flying, somewhat dense; external border more or less straight 
in basal half, thence broadly arcuate with the apical margin; internal 
border arcuate in apical half, thence outwardly oblique to base and 
more or less arcuate or straight. Apex rather small, subacute at 
tip and more or less finely setose. 

Appendage short mammilliform with few very fine and rather long 
setee about tip. 

Superior pudendal membrane quite broadly and triangularly ex- 
posed and obsoletely longitudinally rugulose. 

Ventrolateral surfaces quite evenly convex, the submarginal groove 
broad and shallow, curving inward beneath the fossa to the base of 
the apex, the external and apical borders of the dorsal plate appear- 
ing quite strongly explanate: surface punctate and setose, sete quite 
long and flying. Internal border of the valves contiguous in basal 
fourth, the genital fissure fusiform and situated at middle two- 
fourths, apparently closed by the inferior pudendal membrane. 


REVISION OF ELEODIINI—BLAISDELL, 159 


In all of the specimens examined the pudendal membrane passes 
directly backward, and therefore immediately continuous with the 
valvular membrane, there being no fornix laterally, the angle formed 
is in the median line and corresponds to margin of the genital orifice. 

Habitat—Island of Santa Margarita (collected by the Alba- 
tross Expedition of 1888, collection of U. S. National Museum) ; 
Lower California (E1 Taste, Gustav Beyer; San Francisquito, Coll. 
Acad. Nat. Sciences of San Francisco). 

Number of specimens studied, 32; 22 from Santa Margarita Island, 
and 10 from the mainland of Lower California. 

Type.—Cat. No. 4169, U. S. National Museum. 

Type-locality.—Santa Margarita Island, Lower California. 

Salient type-characters.—Feebly shining. Thorax subquadrate, 
sides broadly and evenly .rounded, base slightly wider than the apex, 
anterior angles subacute, posterior angles obtuse, dise sparsely and 
finely punctate. Elytra widest at apical third; dise moderately, 
slightly depressed along the suture in the female, with regular distant 
series of fine punctures, the intervals very sparsely and minutely 
punctulate (Linell). 

Diagnostic characters —The species resembles somewhat 4. omissa 
LeConte, from which it can easily be separated by the armed anterior 
femora and by the first joint of the anterior tarsi of the males being 
clothed beneath with a tuft of golden pubescence. 

From subnitens it is recognized by the shining integuments, smalle. 
size, and ovate form. 

From all other members of the subgenus by not having the two 
basal joints of the anterior tarsi in the males with pubescent pads 
beneath. 

All of the specimens that I have seen are quite homogeneous and 
without any tendency to muricate punctuation. Linell’s cotypes are 
before me; most of them are slightly brownish, probably from slight 
immaturity. 

A species found upon the mainland of the peninsula agrees in all 
essential structural characters with the specimens from the island, and 
I therefore consider them to be one and the same. Both have the 
elytra widest behind the middle; the femoral teeth vary in both sexes, 
sometimes acute and again obtuse in the males. In the peninsular 
form the body is rather more convex and the elytral punctuation is 
stronger and the striz more or less impressed, the intervals almost 
feebly convex. The tarsal spinules are ferruginous in the insular 
form and darker in those of the mainland. 

Specimens of the peninsular form, collected at San Francisquito, 
and which were before the great disaster in the collection of the Cali- 
fornia Academy of Sciences, were referred to gentilis by Doctor 


160 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Horn: the latter species is a race of gigantea, and occurs much 
farther north in southern California.* ” 

Some observers have referred this form also to énnocens, which 
does in some respects resemble it, for instance, in being widest at 
the posterior fourth of the elytra, the latter having strie of punctures 
which are more or less impressed and the intervals more or less 
convex. /nnocens is also more elongate, and the genital characters 
present a decided subgeneric difference. Compare in Plate 4, figs. 21 
and 22. 

Relationships.—Insularis is no doubt more closely related to sub- 
nitens than to any other species in the United States fauna. It isa 
true Promus in genital characters, and not intermediate between 
Blapylis and ELleodes proper as suggested by Linell. 

General observations.—The mentum is variable, and moderate in 
size. The middle lobe is subtrapezoidal in outline, sides more or 
less arcuate with the apex or converging with the apex more or less 
rounded. : 

Prosternum not prominent ventrally. In the insular specimens it 
is quite evenly but not strongly rounded, antero-posteriorly between 
the coxe, with a small mucro behind; in the peninsular form it is 
quite horizontal between the cox and terminating behind in a rather 
strong and subtriangular mucro. Between the cox the surface is 
more or less strongly punctato-rugose. 

The mesosternum is quite gradually oblique and more or less con- 
cave. 

The tibial grooves of the femora are usually distinctly evident for 
the whole length of the inferior surface, the floor of each being quite 
plane and glabrous or more or less punctate; the margins are more or 
less acute and cariniform. In the peninsular specimens before me the 
floors are quite glabrous. 

The external borders of the anterior tibiz are distinctly carinate, 
the carina forming the anterior lip of the tarsal groove, which is here 
clearly indicated and not at all glabrous. 

The middle and posterior tibiz are evidently very faintly or not 
in the least carinate externally near the base, and the tarsal grooves 
are indicated in apical three-fourths, margins more or less well in- 
dicated. The grooves are sculptured like the rest of the tibial surface. 

The protarsi are about of equal stoutness in the sexes. 

Tarsal formula: 


Pro. Meso. Meta. Metatibia. 
Male.—24 23 3 54 
Female.—2 23 3 54 


See Proc. California Acad. Sci., 2 ser., IV, Pt. 1, pp. 306 and 349. 


REVISION OF ELEODIINI—BLAISDELL, 161 


ELEODES SUBNITENS LeConte. 


Bleodes subitens LeConte, Ann. Lyceum Nat. Hist. N. Y., V 1851, p. 
134.—Horn, Trans. Amer. Phil, Soc., XIV, 1870, p. 319. 

Fusiform to elongate ovate, noticeably narrowed anteriorly, surface 
subopaque, finely punctate and smooth. 

Head evenly convex, impressions scarcely evident, frontal suture 
distinct; surface finely and evenly punctate, punctures quite indis- 
tinct over the vertex. Antenne long, rather robust, outer four 
joints slightly compressed and feebly widened; third joint equal to 
the next two taken together; fourth just noticeably longer than the 
fifth, the latter to the eighth inclusive subequal in length; ninth and 
tenth feebly and transversely suborbicular; eleventh short oval. 

Pronotum quadrate, widest at the base and slightly narrowing from 
base to apex, rarely widest at the middle and narrowing slightly to 
base as well; dsc evenly convex from side to side, very finely and sub- 
obsoletely punctulate; apex feebly and broadly emarginate, obsoletely 
margined and equal to the length; s#des feebly and quite evenly arcu- 
ate from apex to base, very finely beaded; base broadly and feebly 
arcuate or subtruncate, about one-fourth longer than the length, one- 
sixth to one-fourth wider than the apex: apical angles subacute and 
anteriorly prominent; basal angles obtuse and not rounded. 

Propleure smooth, obsoletely punctulate and rugulose. 

Elytra oval, widest at or just behind the middle, and moderately 
attenuate in apical fourth; base feebly and broadly emarginate or 
subtruncate, slightly wider than the contiguous prothoracic base; 
humeri obtuse or feebly subacute, scarcely or very feebly prominent 
anteriorly; sides evenly arcuate, apex subacute or narrowly rounded ; 
dise moderately convex or slightly depressed on the dorsum, laterally 
regularly rounded, arcuately declivous posteriorly; punctate, punc- 
tures fine and closely placed in rather distant unimpressed series, 
intervals more or less irregularly and very finely punctulate: sculp- 
turing more or less indistinct, not denser nor coarser laterally and 
on apex. 

Epipleure slightly widened at base, thence gradually narrowed to 
apex, not wide, the superior margin broadly sinuate basally and curv- 
ing upward to the humeral angle; surface smooth and obsoletely 
punctulate. 

Sterna smooth and more or less obsoletely sculptured. 

Parapleure smooth and more or less distinctly or obsoletely 
punctate. 

Abdomen smooth, finely punctulate, and more or less indistinctly 
rugulose; fifth segment finely setose about the margin. 

Legs rather long and moderately slender. Anterior femora dentate 
or sinuate; the anterior spurs of the anterior tibie similar in the 

59780—Bull. 683—09——11 


162 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


sexes, and slightly larger and longer than the posterior. Tarsi dis- 
similar in the sexes and rather robust. : 

Male.—Body feebly robust and somewhat narrow. Antenne reach- 
ing beyond the prothoracic base. Elytra not broadly oval, noticeably 
attenuate, and rather obliquely declivous posteriorly; apex not pro- 
duced; abdomen more or less oblique, moderately convex, and the 
first segment is more or less broadly impressed at middle. Anterior 
femora armed with a moderate and acute tooth at outer fifth. Ante- 
rior tarsi with the first joint very feebly thickened, and clothed in 
apical third beneath, with a tuft of silken pubescence; second joint 
simple with groove entire. 

Female.—Robust. Antenne reaching slightly beyond the pro- 
thoracic base. Elytra broadly oval, less attenuate, and arcuately 
declivous posteriorly. Abdomen rather strongly convex, horizontal. 
Anterior femora more or less distinctly sinuate in outer fifth; ante- 
rior tarsi unmodified. 

Measurements.—Males: Length, 21-24.5 mm.; width, 8.2-9 mm. 
Females; Length, 23-26 mm.; width, 10.5-12 mm. 

Genital characters, male-—Edeagophore elongate fusiform, scarcely 
arched. 

Basale oblong-oval, sides subparallel, surface moderately convex. 

Apicale triangular, about one-half longer than wide; Surface mod- 
erately convex with a membranous groove in apical half: sides feebly 
arcuate or straight to the tip, the latter acute; base rather narrowly 
rounded at middle, broadly and feebly sinuate laterally. 

Sternite quadrato-parabolic. Each lobe rather long with the ex- 
ternal border evenly arcuate, apical margin rather narrowly rounded, 
angle not evident; internal border arcuate; surface slightly convex, 
‘ather densely and coarsely punctate, setee long and quite dense. 
Sinus broad, apical half not closed by membrane, the latter not setose. 

Female.—Genital segment subquadrate in outline. Valves expla- 
nate externally, deflexed at apex and reflexed laterally and quite 
densely setose. 

Valvula (Plate 5, fig. 3).—Dorsal plate oblong-oval; internal fifth 
reflexed, glabrous and impunctate, externally upwardly oblique and 
more or less deflexed peripherally; surface longitudinally channeled, 
‘ather densely punctate and setose in outer four-fifths, setee long and 
flying; external border quite arcuate; apical margin arcuate exter- 
nally with the external margin, inwardly sinuate with the internal 
apical lobe and set with sete; inner margin broadly arcuate. Apex 
triangular and rather short, finely setose. 

Appendage large and transversely semielliptical, rather thickly 
set with quite long hairs. Fossa transverse, narrow, and closely 
fitting the base of the appendage. 

Basal prominences not evident. 


a et 


REVISION OF ELEODIINI—BLAISDELL, 163 


Superior pudendal membrane obsoletely rugulose and reaching 
about to the middle of the dorsal plate. 

Ventrolateral surfaces—Body of the segment quite strongly con- 
vex, broadly and quite deeply concave beneath the apex, the con- 
eavity continuous with the submarginal groove, the latter broad and 
shallow from the base beneath the explanate external border of the 
dorsal plate; surface coarsely punctate and setose, sete rather short. 
Genital fissure rather narrowly fusiform, the internal margins of the 
valves being contiguous in basal and apical fourths. Inferior puden- 
dal membrane apparently closing the fissure. 

Habitat—Arizona (region of the Gila River, LeConte and Horn; 
Fort Grant; Oracle in April, Hubbard and Schwarz; Prescott, col- 
lection Charles Fuchs; Nogales, September 22, Albert Koebele and 
F. W. Nunenmacher). 

Number of specimens studied, 20. 

Type in the LeConte collection. 

T ype-locality —* Flumen Gila,” Arizona (LeConte). 

Salient type-characters—Narrowed anteriorly, generally opaque. 
Thorax searcely conspicuously narrowed anteriorly, sides usually 
straight, finely and sparsely punctate. Elytra finely, distinctly, and 
seriately punctate (LeConte). 

Diagnostic characters.—Similar to goryi in general outline, but less 
robust and less inflated, and more fusiform than either the above or 
insularis. The surface is opaque and the elytra sculptured with striz 
of fine punctures. The latter are in contrast to the more or less 
coarse punctures of gory. 

The males have only the basal joint of the anterior tarsi pubescent 
beneath, which distinguishes it from all other members of the sub- 
genus, except énsularés. 

The prothorax appears widest at the base when viewed vertically 
from above, with the sides converging to apex (forma typica) ; but 
these characters depend upon the sides being normally arcuately de- 
flexed, otherwise it would appear more or less widest at the middle. 
The base is always relatively wider, as compared to the apex, than in 
insularis. 

Two males from Prescott, Arizona, have the pronotum widest at 
the middle, and the sides are feebly sinuate before the basal angles, 
the base appearing narrower than normal (forma sinuata). Their 
large size and dull integuments serve to separate them from énsularis. 
Such variations are to be expected; some females exhibit this tend- 
ency, the sides of the pronotum becoming more strongly arcuate. 

The mentum is variable in form as in ‘nsularis. The surface is 
more or less distinctly excavated laterally and as a result the middle 
is more or less longitudinally ridged. 


164 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Prosternum is horizontal or feebly arcuate between the coxe, and 
quite strongly and subtriangularly mucronate behind. The pro- 
sternum gradually slopes from the anterior margin to a point between 
the coxee, so that the coxal region appears less suddenly protuberant 
ventrally than in many other species. 

In a larger series than the one before me there would no doubt be 
greater variation in this respect. 

The mesosternum is moderately oblique and quite broadly concave. 

The tibial grooves of the femora are much less strongly developed 
in this species than in dnsularis. In the anterior femora they are the 
widest and moderately concave, sides converging from the femoral 
teeth, becoming evanescent before reaching the base; the middle 
grooves are less strongly defined and narrower, evanescent before 
reaching the base; the posterior are the least developed, quite narrow 
and evanescent near the middle of the femora. The margins of the 
grooves are obtuse, rising but slightly above the quite flat floors of 
the same. The anterior margins of the profemoral grooves are cari- 
nate, becoming dentately laminate at outer fifth, and thence to apex 
sinuate and more or less obsolete. 

The tibia are not noticeably compressed, and not in the least 
‘arinate externally; the tarsal grooves are rudimentary and more or 
less obsolete. 

The tarsi are about equally stout in the sexes. 

Tarsal formula: 


Pro. Meso. Meta. Metatibe. 
Male.—34 4} 3 _ t( 
Female.——34 43 5 8 


ELEODES GORYI Solier. 


Eleodes goryi Sortrer, Studi Emtomol., 1848, p. 251, pl. x, figs. 14, 15.— 
CHAMPION, Biol. Centr.-Amer., IV, Pt. 1, 1885, p. 93. 

Eleodes seriata LeContr, Proc. Acad. Nat. Sci. Phila., 1858, p. 185.—Horn, 
Trans. Amer. Phil. Soc., XIV, 1870, p. 319. 


Ovate fusiform, more or less shining to subopaque, strongly con- 
vex, elytral punctures usually within larger dents. 

Head \ess than twice as wide as long, feebly convex, rather finely 
and more or less densely punctate. Antenne long, reaching beyond 
the prothoracic base, outer four joints scarcely compressed and feebly 
dilated, third joint not as long as the next two taken together, fourth 
decidedly longer than the fifth, the latter to the eighth inclusive sub- 
equal, eighth triangular, ninth and tenth more or less orbicular, 
eleventh oval. 

Pronotum widest at the middle, where it is but shghtly wider than 
the base; disc moderately and evenly convex, finely, rather densely 
and regularly punctulate; apea moderately deeply emarginate, finely 
or more or less obsoletely beaded, and scarcely equal to the length; 


REVISION OF ELEODIINI—BLAISDELL. 165 


sides more or less converging from base to apex, evenly and more or 
less moderately arcuate and finely beaded: base more or less arcuate 
to truncate and finely margined, two-fifths to one-half wider than 
the apex; apical angles subacute and more or less prominent an- 
teriorly; basal angles obtuse and not rounded. 

Propleurw more or less convex, obsoletely punctulate and more or 
less rugulose. 

Elytra oval, and more or less inflated; base truncate to slightly 
emarginate and scarcely wider than the contiguous prothoracic base; 
humeri obtuse and not in the least rounded to subacute, not promi- 
nent anteriorly; s/des evenly arcuate, apex obtuse and not produced; 
dise very strongly convex and without any semblance of even an ob- 
tuse margin, more or less suddenly and arcuately or vertically de- 
clivous posteriorly; striato-punctate, the punctures small to moderate 
in size and arranged in rather distant series, or more frequently the 
punctures are sunken within larger dents that vary in size in the 
same series, the punctures may be rather evenly and closely to irregu- 
larly spaced in each series, the intervals are finely, irregularly, and 
rather thickly punctulate; the dents are usually larger at the sides, 
scarcely denser, and not evident on the apex. 

Epipleure more strongly widened at base and moderately narrow, 
gradually narrowing to apex; surface obsoletely punctulate. 

Sterna more or less punctate and rugulose. 

Parapleure rather finely and densely punctate. 

Abdomen feebly shining, feebly punctate and more or less obso- 
letely rugulose, punctures denser on the last segment, intercoxal 
process rather broad. 

Legs rather long and more or less moderately stout; anterior fem- 
ora acutely or obtusely dentate: anterior tibial spurs acute and 
unequal, the anterior longer and very slightly thicker. Anterior and 
middle tarsi dissimilar in the sexes. 

Male.—Elongate fusiform, scarcely inflated. Abdomen more or 
less oblique, moderately convex, first segment broadly impressed in 
its entire length. Anterior femora armed with a short, more or less 
acute tooth; first two joints of the anterior tarsi thickened and 
‘dilated, clothed beneath with dense and more or less spongy pubes- 
cence, surface of the pads flat; first joint of the middle tarsi feebly 
thickened and clothed in apical half beneath with a dense oval pad 
of pubescence similar to those of the anterior tarsi. 

Female.—Ovate fusiform. Elytra more or less strongly inflated 
and usually vertically declivous posteriorly. Abdomen horizontal, 
more or less strongly convex, not impressed. Anterior femora with a 
very obtuse tooth or simply sinuate; anterior and middle tarsi un- 
modified. 

Measurements.—Males: Length, 15-20.5 mm.; width, 5.8-9 mm. 
Females: Length, 19-22 mm.; width, 8.2-10 mm. 


166 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Genital characters, male—Edeagophore oblong-oval. rather pro- 
longed and acute at apex, not arched. “ 

Basale oblong, evenly and moderately convex, sides nearly parallel 
and very feebly arcuate. 

Apicale triangular, moderately elongate, more or less arcuately 
decurved at tip; surface evenly and not strongly convex, with a 
median linear groove at middle three-fifths, which is slightly dilated 
in basal third and more or less membranous; sides moderately arcuate 
in basal third and thence broadly and evenly sinuate to apex, the 
latter produced and acute; base subacutely lobed at middle, broadly 
and more or less sinuate laterally. 

Sternite subtriangularly parabolic in outline. Each lobe triangular 
with the external border more or less straight in basal half, thence 
evenly arcuate to apex, the latter moderately rounded; internal border 
only defined near apex, thence to base membranous; surface more or 
less convex, bright and shining and sparsely punctate in the externo- 
basal half, with the interno-apical moiety sloping inwards toward 
the sinus and membrano-chitinous, densely punctate and setose, sete 
quite long. Sinus narrow and nearly closed by membrane, the latter 
more or less setose. 

Female.—Genital segment subquadrate, apical half more or less 
deflexed and conspicuously setose. 

Valvula (Plate 4, fig. 5).—Dorsal plate obovate, with the internal 
fourth of the apical half reflexed, smooth and glabrous; outwardly 
plane or feeble convex, surface shining and sparsely punctate, each 
puncture with a long seta; external border straight to feebly arcuate; 
apical border more or less arcuate to subtruncate and moderately 
rounded at the angle; internal border arcuate in apical two-thirds, 
thence outwardly oblique and more or less sinuate to base; apex 
moderately short, acute and finely setose. 

Appendage flattened, semi-oval, scarcely twice as wide as long, 
sparsely clothed with rather short sete. Fossa large and fringed 
with quite long setz, longest externally. 

Superior pudendal membrane reaching to about the middle of the 
dorsal plate, reflected ‘apically and continuous with the valvular mem- 
brane without fornix, and thereby shortest in the median line. 

Ventrolateral surfaces very moderately convex; surface lines 
straight as viewed longitudinally, rather densely clothed with quite 
long sete; submarginal groove broad and oblique beneath the ex- 
planate external and apical borders of the dorsal plate, terminating 
at the internal margin of the fossa. Valves contiguous in basal third, 
the genital fissure rather broadly fusiform, and closed in the basal 
half by the inferior pudendal membrane. 


. 


REVISION OF ELEODITINI—BLAISDELL. 167 


Habitat—Texas (Brownsville, June, Hubbard and Schwarz, H. F. 
Wickham; Bosque County, G. W. Belfrage; San Diego, May, E. A. 
Schwarz; Beeville, April; San Antonio, June, H. Soltau; Dallas; 
Dimmit County, Hubbard and Schwarz); New Mexico (Dr. George 
Horn); Mexico (Mirador, Sallé; Rinconado, Hoge). 

Number of specimens studied, 45. 

Type is in the Solier collection; of seriata in the LeConte collec- 
tion, 

T ype-locality.—Mexico (Solier); Texas (LeConte). 

Salient type-characters.—Subopaque. Thorax quadrate, slightly 
narrowing anteriorly, sides feebly rounded; apical angles acute; 
basal angles obtuse and not rounded. Elytra with the base broadly 
emarginate, humeri acute, strongly striato-punctate, intervals irregu- 
larly, finely and sparsely punctulate (seriata, LeConte). 

Diagnostic characters.—Distinct by its coarse punctuation, which 
consists of distinct rows of coarse punctures or dents in which the 
punctures are distinctly visible. The elytra are very convex and 
normally without any semblance of even an obtuse margin. Many 
examples have the serial punctures fine, and these are frequently con- 
‘founded with soror (see carbonaria var. soror). 

The basal joints of the anterior tarsi in the males are more strongly 
modified than in any other species of the subgenus. 

Any single locality does not appear to produce any particular form 
of punctuation; the most strongly sculptured specimens before me 
were collected by Belfrage in Bosque County, Texas. 

Specimens from Mexico do not differ from those taken in Texas. 

The mentum is variable; surface convex, coarsely punctate and at 
times more or less excavated laterally. 

Prosternum more or less evenly arcuate from the anterior margin 
to the tip of the intercoxal process; at times more or less horizontal 
between the cox. All of the specimens examined have the prosterna 
strongly and triangulo-conically mucronate behind. 

Mesosternum at times obliquely subhorizontal or feebly arcuately 
oblique, rarely vertically arcuate, always more or less strongly 
concave, 

The grooves of the anterior femora are inwardly attenuate, about 
reaching the base; those of the mesofemora extending inwards to 
about the basal third, while on the metafemora they reach only to 
about the middle. The anterior margins of the anterior grooves are 
scarcely carinate, except in the external sixth, where the teeth are 
developed. 

The tibiz are not compressed and not in the least carinate ex- 
ternally, and the tarsal grooves are obsolete. 


168 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


The protarsi in the male stouter than in the female. 
Tarsal formula: 


> 
Pro. Meso. Meta. Metatibix. 
Male.—s 4 45 G4 
Female.—3 4 43 64 


ELEODES STRIOLATA LeConte. 


Eleodes striolata LeContr, Proc. Acad. Nat. Sci. Phila., 1858, p. 185.— 
Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 319. 

‘Fusiform to ovate, smooth and more or less shining, briefly cau- 
date; elytra with approximate rows of fine punctures. 

Head feebly convex, smooth, finely, not densely and quite indis- 
tinctly punctate, impressions very slight and vague; frontal suture 
evident. Antenne rather long, reaching slightly beyond the pro- 
thoracic base, slender, scarcely compressed and feebly incrassate in 
outer four joints, third joint scarcely as long as the next two taken 
together, fourth very slightly longer than the fifth, the latter to the 
eighth, inclusive, subequal, eighth obconieal, ninth and tenth orbicu- 
lar, eleventh ovate. 

Pronotum subquadrate, widest at about the middle, very slightly 
narrowing at the base, and a little more strongly so anteriorly to 
apex, about one-third wider than long; disc evenly and moderately 
convex from side to side, finely and obsoletely punctulate; apex 
evenly and rather broadly emarginate, finely and more or less obso- 
letely margined; sides evenly arcuate from base to apex, finely 
beaded; dase broadly rounded and very finely margined, one-half 
wider than the apex and about one-third wider than long; apical 
angles subacute and slightly prominent anteriorly; basal angles 
obtuse and more or less rounded. 

Propleurw obsoletely punctate, smooth and more or less feebly 
rugulose. 

Elytra oval, widest at the middle, at times slightly more than twice 
as long as wide; base feebly emarginate and slightly wider than the 
contiguous prothoracic base; humeral angles slightly acute, not pro- 
longed nor clasping the prothoracic base; sides evenly arcuate, apex 
slightly produced; disc feebly to moderately convex on the dorsum, 
laterally rather narrowly rounded, arcuately declivous behind, in- 
flexed sides more or less plane and extending slightly farther than 
usual upon the sides of the abdomen; punctate, punctures fine and 
arranged in approximate series, which are scarcely ever impressed ; 
the punctures of each series are separated by a space that is about 
equal to their diameters; each interval has a single series of more 
distantly spaced and very minute punctules which become more or 
less irregular at times; on the inflexed sides the serial punctures be- 
come coarse and more or less impressed. The apex is slightly pro- 


REVISION OF ELEODIINI—BLAISDELL. 169 


duced, with the tips of the elytra slightly separated and feebly 
deflexed. 

Epipleurw moderately widened beneath the humeri, thence rather 
narrow to the apex; surface smooth, finely and sparsely punctate; 
the superior margin at times more or less obsolete in apical two-thirds, 
being defined by the marginal series of the discal punctures. 

Sterna rather smooth and obsoletely punctate and rugulose. 

Parapleure more or less obsoletely punctate. 

Abdomen smooth and shining, usually obsoletely punctate and more 
or less finely rugulose; horizontal. 

Legs quite long and slender. Anterior femora dentate or sinuate; 
anterior spurs of the anterior tibix about a half longer and slightly 
thicker than the posterior, both acute and quite similar in the sexes. 
Anterior tarsi dissimilar in the sexes. 

Male—Body more or less fusiform, slightly elongate and feebly 
robust. Antenne reaching a short distance beyond the prothoracic 
base. Elytra rather gradually declivous posteriorly. Abdomen 
moderately convex. Anterior femora with a moderately acute tooth 
at outer fifth; anterior tarsi with the first and second joints feebly 
dilated and flattened beneath, densely clothed with a fine golden 
spongy pubescence; mesotarsi unmodified. 

Female.—Body ovate and robust. Antenne reaching slightly be- 
yond the prothoracic base. Elytra arcuately declivous posteriorly. 
Abdomen quite strongly convex. Anterior femora more or less feebly 
sinuate in outer fifth; anterior tarsi unmodified. 

Measurements —Males: Length, 18-21 mm.; width, 7.5-9 mm. 
Females: Length, 20-23 mm.; width, 9.5-10.5 mm. 

Genital characters, male-—Edeagophore  elongately fusiform, 
searcely arched and rather depressed. 

Basale oblong oval, feebly and evenly convex, sides feebly arcuate. 

A picale triangular about one-half longer than wide; surface rather 
feebly and evenly convex, with a narrow membranous groove in 
middle third; sides rather straight or feebly arcuate, apex acute; 
base rather narrowly lobed at the middle, broadly sinuate laterally. 

Sternite transversely parabolic in outline. Each lobe with the 
external border evenly and rather broadly arcuate with the apical 
border, which is narrowly rounded, angle scarcely evident at the 
internal border, the latter short; surface more or less convex, mem- 
branous in the internal half, chitinous externally, sparsely punctate, 
setose; sete rather long, slightly longer on the apical edge, not very 
dense ; surface impunctate in basal third. Sinus broad, nearly closed 
by the membrane, the latter with a few scattered set. 

Female.—Genital segment subquadrate, valves more or less reflexed 
and explanate externally, deflexed at apex and distinctly setose. 


170 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Valvula (Plate 5, fig. 2).—Dorsal plate oblong-oval. Internal 
border slightly reflexed in apical half and impunctate, externally 
more or less reflexed; surface slightly concave longitudinally, finely 
and sparsely punctate, each puncture with a rather long flying seta; 
external border arcuate; apical margin arcuate in the external half 
and sinuate within to the internal lobe of apex, the latter moderate 
and triangular; internal border more or less arcuate apically and 
sinuous toward base. Apical margin set with a few flying hairs, and 
the apical lobe is finely setose. 

Appendage short and broadly semi-elliptical and set with rather 
short sete; its external margin is continuously arcuate with the ex- 
ternal border of the dorsal plate. Fossa narrow and_ transverse. 
Basal prominences not evident. 

Superior pudendal membrane reaching to the middle of the dorsal 
plate, feebly and irregularly longitudinally rugulose. 

Ventrolateral surfaces rather strongly convex, narrowly and trans- 
versely so laterally, glabrous, sparsely and coarsely punctate, finely 
setose about the apex: submarginal groove broad, passing quite 
obliquely to the internal lobe of apex. Genital fissure narrowly 
fusiform; valves quite contiguous basally and apically. Inferior 
pudendal membrane not visible. 

Habitat—Texas (Goliad County, October, J. D. Mitchell; San 
Antonio, Hubbard and Schwarz; Victoria; adjoining regions of 
Mexico, Dr. George Horn). 

Number of specimens studied, 11. 

Type a female in the LeConte collection. 

Type-locality, Laredo (Webb County) to Ringgold Baracks 
(Starr County) in southern Texas along the Rio Grande. 

Salient type-characters.—Shining, thorax quadrate, slightly nar- 
rowed anteriorly and posteriorly; sides moderately rounded, apical 
angles acute, the posterior obtuse. Elytra with the humeri acute, 
slightly caudate at apex, punctures of the disc arranged in approxi- 
mate series, becoming obliterated upon the apex, intervals sparsely 
and finely punctulate (LeConte). : 

Diagnostic characters.—Easily recognized by the character of the 
elytral punctuation, which consists of closely placed series of fine 
punctures, and by the elytral apices being slightly produced and 
feebly divergent. The males are quite fusiform and _ noticeably 
elongate; the females are more ovate; the legs are quite long and 
moderately slender in both sexes. It is most closely related to fus?- 
formis and liable to be confused with some of the larger specimens of 
that species. In strio/ata the anterior femora are armed in the males 
and sinuate in the females, the humeral angles are not prolonged and 


————, 


REVISION OF ELEODITNI—BLAISDELL, 171 


do not clasp the prothoracic base; the prothorax is more or less wider 
at middle than at base. Another distinguishing character is the 
feebly caudate elytra. 

I have seen specimens of strio/ata labeled fusiform/s in collections— 
large examples of the latter may simulate the former by having the 
elytral punctures arranged in closely placed series. The tarsi in the 
two species are longer and more slender than in the other species of 
Promus, opaca excepted. 

At the time of writing the Revision Doctor Horn had but three 
females before him and he had some doubt about placing the species 
in the present subgenus. I can not find any mention in the literature 
of a male ever having been studied. 

I have a series of eight males before me and find them to be true 
Promus. ; 

The mentum is quite triangular, with apex rounded, in the speci- 
mens examined, and the surface is more or less convex at middle and 
foveate laterally. 

LeConte writes that the prosternum is horizontally produced and 
acute. In the series at hand I find it quite homogeneous—rounded 
antero-posteriorly between the cox and gradually, more or less 
evenly subconically mucronate behind. The mesosternum moderately 
oblique, feebly arcuate, broadly and more or less deeply concave. 

The tibial grooves of the femora are not well developed and are 
more or less concave and glabrous, impunctate. Those of the pro- 
femora extend to about the middle and are inwardly attenuate, some- 
times obsoletely indicated to the base; margins obtuse, rarely sub- 
carinate, the anterior margin being usually simply dentately laminate 
to produce the tooth in the male; in the female the anterior grooves 
are generally quite concave and well defined. 

Those of the meso- and metafemora seldom extend inward to the 
middle, rarely more than the outer third and are more or less poorly 
defined; at times the posterior grooves are short and evanescent in 
the females. 

The external surfaces of the tibiae are rounded, and in the exam- 
ples before me not in the least carinate; the tarsal grooves are obso- 
lete, those of the mesotibiwe being frequently indicated, the external 
surface is occasionally flattened. 

The anterior tarsi are distinctly more robust in the male than in 
the female. 

Tarsal formula: 

Pro. Meso. Meta. Metatibia. 
Male.— 3 4] 5 84 
Female.— 2} 4 4] 7 


big BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


ELEODES FUSIFORMIS LeConte. 


Eleodes fusiformis LeContrr, Proc. Acad. Nat.’ Sci. Pifila., 1858, p. 184.— 
Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 318. ; 


Fusiform, body narrowing quite evenly both ways, smooth and 
more or less shining. 

Head more or less feebly convex, smooth, usually without impres- 
sions, frontal suture distinct, finely and not densely punctate, punc- 
tures very sparse over the vertex. Antennw rather long, shghtly 
compressed and incrassate in outer four joints, third joint not quite 
as long as the next two taken together, fourth to the eighth inclusive 
subequal, eighth feebly triangular, ninth and tenth very feebly trans- 
versely orbicular, eleventh short ovate. 

Pronotum trapezoidal widest at the base where it is about one- 
half wider than the apex; disc more or less moderately convex from 
side to side, laterally slightly arcuately declivous, very finely, sparsely 
and indistinctly punctulate; apex moderately and evenly emarginate 
and more or less obsoletely margined; sides converging anteriorly. 
not strongly but evenly arcuate, finely margined; base quite broadly 
rounded, and more or less distinctly margined; apical angles quite 
prominent anteriorly and subacute, sometimes feebly rounded; basal 
angles obtuse and more or less rounded. 

Propleure quite smooth, obsoletely punctate, and more or less 
rugulose. 

Elytra ovate to oval, widest at or a little in advance of the middle; 
base broadly emarginate with the humeral angles clasping the basal 
angles of the prothorax, slightly wider than the contiguous base of 
the same; Awmeri more or less acute and anteriorly prominent; sides 
evenly arcuate, apex not produced but rather narrowly rounded; disc 
more or less feebly convex on the dorsum, rather narrowly rounded at 
the sides but not suddenly inflexed, arcuately declivous posteriorly ; 
inflexed sides quite plane and oblique; surface subseriately punctate, 
the series quite closely placed and at times quite distinct, intervals 
with a single series of more distantly spaced, at times irregularly ar- 
‘ranged punctures, the serial punctures usually slightly larger than 
the interstitial. Often they are subequal and more or less confused, 
and always more or less so laterally and on apex. 

Epipleure on the same plane as the inflexed sides of the elytral 
disc, moderately wide and gradually narrowing from base to apex; 
surface smooth, finely and sparsely punctate. 

Sterna somewhat glabrous, more or less obsoletely punctate and 
rugulose. 

Parapleure quite coarsely and more or less obsoletely punctate. 

Abdomen glabrous and shining, finely and sparsely punctate with 
the intercoxal process more coarsely sculptured. 


fl 
’ 


REVISION OF ELEODIINI—BLAISDELL. 173 


Legs rather long, the anterior comparatively short; anterior femora 
mutic; anterior spurs of the anterior tibia and the anterior tarsi are 
dissimilay in the sexes, 

Male.—Rather robust. Antenne reaching a little beyond the pro- 
thoracic base. Abdomen slightly oblique, moderately convex and 
somewhat broadly impressed on the first two segments. Anterior 
spurs of the protibiw about twice as long and slightly stouter than 
the posterior, slender and acute. Anterior tarsi with the firstsoint 
searcely dilated, slightly compressed so that the vertical diameter is 
slightly greater than the transverse; the first in the apical moiety and 
the second densely clothed beneath with a flattened pad of more or 
less spongy, golden pubescence. 

Female.—Quite robust. Antenne reaching just beyond the pro- 
thoracic base. Abdomen rather strongly convex and horizontal. The 
anterior spurs of the protibiz about one-half longer than the pos- 
terior, feebly thickened and subacute; anterior tarsi unmodified, the 
first joint scarcely thickened at tip beneath, with groove narrow and 
distinct. 

Measurements.—Males: Length, 13.2-17 mm.; width, 6-7.5 mm. 
Females: Length, 11.8-18 mm.; width, 5.8-9 mm. 

Genital characters, male—KEdeagophore small and short oblong- 
ovate, not arched. 

Basale scarcely twice as long as wide, oblong and not strongly con- 
vex, sides feebly arcuate. 

Apicale triangular, about one-half longer than wide, the surface is 
feebly convex, with a feebly developed membranous groove in apical 
half; sides arcuate in basal third; thence to tip broadly and feebly 
sinuate, apex acute; base subacutely lobed at middle and broadly 
sinuate laterally. 

Sternite transversely trapezoidal. Each lobe with the external 
border at base quite straight to feebly arcuate and thence slightly 
converging apically, the apical margin moderately arcuate or trun- 
cate; internal border is scarcely evident: surface scarcely convex, 
chitinous externally and membranous internally, sparsely punctate 
and setose, setee moderately long, with longer ones about the apical 
margin. Sinus broad and closed by the membrane, the latter not 
setose at bottom of the former. 

Female.—Genital segment quadrato-parabolic, valves reflexed later- 
ally and slightly deflexed apically, setose. 

Valvula ( Plate 4, fig. 3).—Dorsal plate oval, with the internal third 
of the apical half reflexed and impunctate, also glabrous; outwardly 
explanate and slightly deflexed, with the surface plane sloping 
obliquely downward and inward. The surface is sparsely punctate 
and setose in outer two-thirds, with the hairs rather long; externa! 


174 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


border moderately arcuate; apical border slightly rounded and feebly 
sinuate just external to the apical lobe, the latterysmall and trian- 
gular; internal border broadly arcuate. The apical margin has a 
few long sete. 

Appendage rather short and transversely semi-elliptical, finely 
setose, and equal in width to the outer two-thirds of the dorsal plate. 
Fossa transverse and narrow. 

Basal prominences not evident. The external border is angulate 
at the base. 

Superior pudendal membrane reaching to the middle of the dorsal 
plate, finely and more or less regularly longitudinally rugulose. 

Ventrolateral surfaces are evenly and moderately convex, glabrous. 
The submarginal grooves are very broad beneath the explanate ex- 
ternal borders of the dorsal plates. Each curves inward around to 
the inner lobe of the apex, so that the apical margin appears explanate 
when viewed from below; surface very sparsely punctate and setose; 
genital fissure very narrowly fusiform. 

Habitat—Texas. Specimens without definite locality labels and 
collected by C. V. Riley, others by Hubbard and Schwarz are the 
largest in the series before me. One specimen from Dallas is also 
large and well developed. A pair collected in Bexar County by 
Charles Drury agree in size with other examples from this State. 
Many years ago I received similar specimens that were collected in 
Bosque County by G. W. Belfrage——Wyoming. Specimens taken at 
Cheyenne by H. Soltau are the smallest specimens before me. Two 
specimens ‘with indistinct labels, but apparently from this locality, 
are larger and moderate in size. Dr. George Horn stated that this 
species is never very abundant, and also as occurring on the plains 
of Nebraska and southward into New Mexico.—Kansas (Wallace— 
W. Knaus). Professor Wickham, in his “ List of the Coleoptera 
of Colorado,” writes that it occurs at Bellevue, and at La Junta on 
authority of Bowditch. 

Number of specimens studied, 24. 

The type in the LeConte collection bears a green dise label, and 
was probably collected in Colorado, in the Platte River Valley, near 
Fort Laramie. 

Salient type-characters—Fusiform. Thorax trapezoidal, narrow- 
ing anteriorly; posterior angles rounded with the base; anterior 
angles acute; disc finely punctate. Elytra with base emarginate, 
clasping the base of the prothorax; humeri anteriorly produced; 
disc rather densely subseriately punctate. Antenne slightly incras- 
sate. Prosternum produced (LeConte). 

Diagnostic characters.—Distinguished from all of the other species 
of the subgenus—except opaca—by the mutic anterior femora. From 


REVISION OF ELEODIINI—BLAISDELL. 175 


opaca it is recognized by the black and shining surface, being neither 
opaque nor pubescent as in that species. 

The elytral punctuation is rather fine and closely subseriate. In 
the large specimens with distinct and closely placed series of pune- 
tures an approach is made toward s¢riolata. (See p. 171.) 

Relationships.—It would be very interesting to determine if, in a 
very large series of specimens collected from all parts of its dis- 
tributional area, any examples of fusiformis could be found with a 
tendency to having the anterior femora sinuate or feebly dentate, 
these characters with any tendency to slight prolongation of the 
elytral apices would show a transition into sfriolata. I expect this. 

General observations.—The mentum is variable. Usually subtri- 
angular with the apex more or less rounded, and at times slightly 
transverse with the anterior margin arcuately rounded from side 
to side; the surface may be longitudinally.convex at middle and more 
or less foveate laterally, or more or less evenly convex with the fovex 
obsolete. 

The prosternum is feebly arcuate between the coxe or subhori- 
zontal, produced and compressed behind into a stout triangular 
mucro; anteriorly as in subnitens. The prosterna in the series at 
hand are quite homogeneous. 

The mesosternum is arcuately oblique and more or less deeply and 
broadly concave. 

The tibial grooves of the femora are distinctly margined, although 
not cariniform, the margins are more or less narrowly rounded; the 
floors are rather concave and glabrous, inwardly and evenly attenu- 
ate, and not quite reaching the bases of the femora. 

The tibiz are more or less cylindrical in transverse section and not 
in the least carinate externally; the tarsal grooves are obsolete, 
although the external surfaces are at times slightly fattened in the 
apical half. 

The anterior tarsi are slightly stouter in the male than in the 
female. 


Tarsal formula: ; 
Pro. Meso. Meta. Metatibix. 


Male.—24 34 43 6 
Female.—24 33 D 6 


ELEODES OPACA Say. 


Blaps opaca Say, Journ. Acad. Nat. Sci. Phila., III, 1828, p. 262. 
Bleodes opaca LeContr, Proc. Acad. Nat. Sci. Phila., 1858, p. 182; Classif. 
Coleopt. North Amer., 1861, p. 226 (Promus).—Horn, Trans. Amer. 
Phil. Soc., XIV, 1870, p. 318. 
Fusiform oval, black, opaque, sparsely clothed with whitish or 
subflavate subsquamiform recumbent hairs, with the dorsum of the 
elytra quite flat. 


176 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Head scarcely twice as wide as long, slightly convex, very feebly 
and quite regularly punctulate. Antenne moderate, scarcely reach- 
ing beyond the prothoracic base, outer four joints very slightly com- 
pressed and dilated, third joint about equal to the next two taken 
together, fourth to the eighth, inclusive, subequal, eighth subtri- 
angular, ninth and tenth suborbicular, eleventh short oval. 

Pronotum widest toward the base, where it is at least one-half 
wider than the apex, and about twice as wide as the head; disc evenly 
and moderately convex, rather densely and very finely punctulate; 
apex emarginate and more or less obsoletely margined; sides feebly 
arcuate and converging from base to apex, very finely beaded; base 
broadly arcuate, frequently very feebly sinuate at middle, finely 
margined; apical angles acute, rather prominent anteriorly, never 
everted ; basal angles broadly and rather obtusely rounded. 

Propleure obsoletely punctulate. 

Elytra pointed oval to ovate, scarcely twice as long as wide; base 
deeply emarginate, humeral angles acute and anteriorly prominent 
clasping the obtuse basal angles of the prothorax; sédes acutely 
margined, evenly arcuate and continuing the lines of the sides of the 
pronotum, apex obtusely rounded; disc flattened and more or less 
feebly convex, sides acutely inflexed and not suddenly arcuately de- 
clivous posteriorly; surface finely, irregularly, and inconspicuously 
punctulate and estriate, rarely striate; inflexed sides with surface 
quite plane, feebly concave at the humeri, and sculptured as on the 
dorsum. 

Epipleure smooth and narrow, very gradually narrowing from 
base to apex; surface obsoletely punctulate. 

Sterna very finely muricately punctured, and more or less obsoletely 
rugulose. 

Parapleure finely punctate. 

Abdomen more or less shining and smooth, finely and quite regu- 
larly punctate, and more or less rugulose. ' 

Legs moderate. Femora mutic in both sexes; anterior tibial spurs 
unequal, the anterior longer and slightly larger than the posterior. 
Tarsi dissimilar in the sexes. 

Male.—Moderately narrow and fusiform. Abdomen moderately 
convex and more or less impressed on the first segment between the 
cox; slightly oblique to the intercoxal process and sterna. First 
two joints of the anterior tarsi slightly widened and clothed beneath 
with dense pads of more or less spongy pubescence, surface of the 
pads quite flat. 

Female.—More or less broadly fusiform oval. Abdomen more or 
less strongly convex; anterior tarsi unmodified. 

Measurements:—M ales: Length, 10.5-12 mm.; width, 4.8—5 mm. 
Females: Length, 10.5-14 mm.; width, 5.5-7 mm. 


REVISION OF ELEODIINI—BLAISDELL, 177 


Genital characters, male.—Edeagophore oblong-ovate as usual and 
not arched, 

Basale short oblong, evenly convex, with the sides more or less 
arcuate. 

Apicale triangular, nearly twice as long as wide; surface evenly 
convex, with a median groove at middle four-sixths, gradually dilat- 
ing from the middle to base; sides feebly arcuate in basal third, 
thence very feebly sinuate to apex, the latter acute; base rather 
broadly lobed at the middle and more or less sinuate laterally. 

Sternite transverse. Each lobe rather short, with the external bor- 
der more or less straight in basal half, thence evenly and broadly 
arcuate to the internal border, angle scarcely evident; internal bor- 
der broadly membrano-chitinous and not defined; surface feebly con- 
vex, sloping inward from the middle to the sinus, glabrous in basal 
and external third, elsewhere punctate and setose; punctures and 
hairs sparsely placed, the latter rather long. Sinus narrow and 
closed by membrane; the latter may be more or less setose. 

Female.—Genital segment quadrate in outline, rather short, setose 
and not deflexed apically. 

Valvula (Plate 4, fig. 4).—Dorsal plate more or less broadly obo- 
vate, with the internal third in apical half reflexed, impunctate, and 
glabrous; surface more or less oblique in the outer two-thirds, and 
consequently rather concave, coarsely and sparsely punctate, setae mod- 
erate in length and more or less flying; external border more or less 
arcuate and continuously so with the arcuate apical margin, angle 
searcely evident; internal border arcuate in apical half, thence to 
base more or less sinuate. Apex small and triangular, subacute, finely 
setose, and continuous with the reflexed internal border of the dorsal 
plate. 

Superior pudendal membrane reaching to about the middle of the 
dorsal plate, and reflexed apically to be directly continuous with 
the valvular membrane, fornix absent, irregularly and obsoletely 
rugulose. : 

Ventrolateral surfaces —Body semicircularly and evenly convex 
and glabrous, rather strongly impressed across the bases of the 
apices; submarginal groove broad and arcuate to the internal 
margin of the fossa, beneath the more or less explanate external and 
apical border of each dorsal plate and fossa, sparsely setose; internal 
border of the valves contiguous in the basal fourth. Genital fissure 
long and narrowly fusiform with the inferior pudendal membrane 
visible in the basal half. 

Habitat—Oklahoma (Fort Supply, Coll. H. S. Barber); Texas 
(Mobeetee, July, H. S. Barber; Coll. Charles Fuchs); Nebraska 
(Sand Hills, September, C. V. Riley; Chadron, August, Lawrence 

59780—Bull. 683—09——12 


178 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Bruner; Dodge County, Coll. F. E. Blaisdell) ; Southern Dakota 
(Alexandria; Volga, Coll. E. C. Van Dyke) ; Kangas (Gove County, 
2.500 feet, F. H. Snow, Warren Knaus, Hubbard and Schwarz; 
Plains, Dr. George Horn) ; Colorado (Dr. George Horn; Trinidad; 
Colorado Springs, F. H. Snow: West Las Animas, H. F. Wickham). 

Number of specimens studied, 38. 

Type not in existence. 

Type-locality as given by Say is Missouri and Arkansas. 

Satient type-characters—Opaque, clothed with whitish hairs; 
thorax with acute anterior and obtusely rounded posterior angles; 
sides hardly arcuate. Elytra with base emarginate to receive the 
prothoracic base; humeral angles acute; lateral edge acute. Lateral 
curve of the body continuous (Say). 

Diagnostic characters.—Distinct from its congeners in haying the 
dorsal surface of the elytra quite flat, with the lateral margins quite 
acute; the surface is opaque and rather sparsely covered with short, 
recumbent, setiform hairs. 

It resembles fusiformis in having the anterior femora unarmed 
and in having the elytral base deeply emarginate with the humeral 
angles embracing the obtusely rounded basal angles of the prothorax. 

Structurally these two species are closely related and undoubtedly 
originated from the same basic stock. In opaca the elytral base more 
strongly clasps the prothoracic base than in fus/formis, and in the 
latter more than in striolata, where this character is at its minimum, 
and entirely absent in goryi, subnitens, and tnsularis. 

It is to be observed that the relations of the prothoracic and elytral 
bases give to the lateral margins of the insect a continuity of outline 
which is most marked in opaca and the least developed in insu/aris, 
these two species constituting the extremes of the subgeneric series. 

Mr. Say, in his description of the species, says that the “ lateral 
edge” of the elytra is not reflected, but in all the specimens before 
me it is slightly and distinctly reflected in basal half of the elytra 
and gradually becomes evanescent as it approaches the apex. 

Variations.—In a series of seven specimens collected at Fort Sup- 
ply, Oklahoma, and Mobeetee, Texas, there is distinct evidence of 
three elytral coste—in other’ words, the sutural and every fourth 
interval is feebly but noticeably convex, with feebly but clearly 
defined strixw of fine and closely placed punctures. Normally the 
“elytra are destitute of striae.” k 

General observations —The mentum is variable, usually triangular 
and rounded at apex, or less frequently subparabolic in outline; sur- 
face feebly convex and more or less foveate laterally. 

Prosternum evenly arcuate antero-posteriorly in the median line, 
sometimes horizontal between and behind the cox; produced pos- 
teriorly into a more or less strong mucro, which is usually compressed 


REVISION OF ELEODITNI—BLAISDELL. 179 


and vertically triangular; rarely tuberculiform and sometimes de- 
flexed. 

The mesosternum is-arcuate in different degrees of obliquity and 
strongly concave. The tibial grooves of the femor are glabrous and 
more or less concave and distinctly defined, with margins narrowly 
rounded. Each groove is inwardly attenuate, the anterior scarcely 
reaching the femoral base, while those of the meso- and metafemora 
searcely attain a slight distance beyond the middle. The external 
surfaces of the tibiw are rounded and more or less obsoletely flattened 
apically. 

Anterior tarsi slightly stouter in the male than in the female. 

Tarsal formula : 


Pro. Meso. Meta. Metatibie. 
Male.—2 24 3 4h 
Fema le.—2 25 33 5 


Subgenus HETEROPROMUS, new. 


Anterior femora mutic. Tarsi slender and spinulose beneath; the 
anterior of the male not dilated nor thickened, and with a small 
transverse tuft of golden pubescence on the ventro-apical margins of 
the first and second joints. Anterior spurs of the anterior tibize 
thicker than the posterior. 

Subgeneric genital characters, male.—Apicale of the edeagophore 
triangular, surface convex and more or less obsoletely grooved along 
the median line. | 

Female—Genital segment triangular in outline, setose, not 
strongly chitinized, dorsal plate not defined from the apex, append- 
age and fossa obsolete. 

Superior pudendal membrane reaching to the apical third of the 
dorsal surface of the valves; internal margins of the valves beneath 
contiguous in basal fourth. Genital fissure long, evenly fusiform, 
and not wide. 

Distribution —Thus far the single species of this subgenus is only 
known from southeastern Texas—from counties bordering on the 
Gulf of Mexico. 

Relationships.—V eterator appears to be quite isolated in the tribe 
Eleodiini, in fact in many respects might be considered as constituting 
a distinct genus. It only resembles Promus (opaca) in external 
facies and no way suggests to my mind any conception of its 
ancestral origin. 

ELEODES VETERATOR Horn. 


EBleodes veterator Horn, Trans. Amer. Ent. Soc., V, 1874, pp. 33. 


Oval, black, opaque, subdepressed above, sparsely clothed with 
short, recumbent yellowish hairs, and slightly more than twice as 
long as wide, 


180 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Head slightly more than twice as wide as long, moderately convex, 
more or less impressed laterally and along the frontal suture, sparsely 
punctulate and pubescent. Antenne rather short, outer four joints 
feebly compressed and very slightly dilated, third joint scarcely as 
iong as the next two taken together, fourth to the seventh inclusive 
subequal, eighth slightly shorter and subtriangular, ninth suborbicu- 
lar, tenth very feebly transversely oval, eleventh rather broadly oval. 

Pronotum nearly twice as wide at base as long, and about one- 
seventh longer than width at apex; disc feebly convex, sparsely 
punctulate and pubescent; ape rather deeply and semi-circularly 
emarginate, obsoletely margined; s/des regularly arcuate and grad- 
ually convergent from base to apex, very finely beaded; base squarely 
truncate, very finely margined and slightly over-lapping the elytral 
base; apical angles slightly prominent anteriorly and subacute; 
basal angles subrectangular and rather narrowly rounded. 

Propleure very finely, sparsely punctulate, pubescent, and rugu- 
lose; broadly and moderately excavated beneath and internally to 
the basal angles. 

Elytra triangularly shield-shaped, squarely truncate at base and 
slightly wider than the contiguous prothoracic base; hwmeri nar- 
rowly rounded; sides evenly arcuate and gradually converging to 
apex, and from the base regularly continuing the curve of the sides 
of the pronotum; apex obtuse and not produced; disc feebly convex, 
quite suddenly deflexed at the sides with margin narrowly rounded, 
but acute in basal third, evenly and strongly arcuately declivous pos- 
teriorly; finely, evenly, and not densely punctulate and pubescent, 
the pubescence arranged so that the surface appears substriate; in- 
flexed sides plane. 

Epipleure rather narrow and feebly dilated at base beneath the 
humeri, very gradually narrowing from base to apex; surface finely 
punctate and-sparsely pubescent. 

Sterna rather densely punctate and feebly shining; prosternum 
obsoletely punctate anteriorly. 

Parapleure very finely and sparsely punctate. 

Abdomen distinctly horizontal in the sexes, glabrous and shining, 
quite coarsely and densely punctate on the first segment, the punc-— 
tures becoming finer and less dense on the succeeding segments, rather 
strongly and evenly convex from side to side. 


Legs rather small and slender; femora mutic, and finely, sparsely 
punctate; tibiew straight; anterior tibial spurs slightly unequal in 
length and stoutness, the internal spurs of the meso- and metatibize 
a little longer than the external; tarsi feebly dissimilar in the sexes. 

Male.—Feebly elongate oval. Antenne reaching to about the ~ 
posterior fifth of the pronotum; elytra widest at the base, sides 


REVISION OF ELEODITNI—BLAISDELL, 181 


noticeably converging to apex. Intercoxal process of the abdo- 
men flattened or more or less slightly concave. Anterior spur of the 
protibiz about one-third longer than the posterior; anterior tarsi 
with joints distinctly grooved beneath, first joint very feebly and 
transversely produced at the ventro-apical margin, with a small 
transverse tuft of golden pubescence upon the same; the second joint 
also has a smaller and similar tuft on the ventro-apical margin. 

Female.—Broadly oval. Antenne reaching to the posterior fourth 
of the pronotum. Elytra widest in basal fourth, sides rather strongly 
arcuate and less noticeably converging to apex. Anterior spur of 
the protibize about a fourth longer than the posterior and slightly 
stouter; first joint of the protarsi not modified beneath. 

Measurements.—Males: Length, 10.5 mm.; width, 5.5 mm. Fe- 
males: Length, 11.5 mm.; width, 6.0 mm. 

Genital characters, male.—Edeagophore of the usual oblong-ovate 
form, elongate and feebly arched. 

Basale short, oblong-oval, evenly convex above with the sides 
evenly arcuate. 

Apicale elongately triangular and very feebly decurved apically; 
surface evenly convex, obsoletely grooved along the median line; sides 
with feeble lobes at basal fourth, thence feebly sinuate to apex, the 
latter moderately produced and acute; base acutely lobed at the mid- 
dle and feebly sinuate laterally. 

Sternite feebly transverse. Each lobe parabolically rounded, with 
the surface feebly convex, sparsely punctate and setose, sete longer 
on the apical margin and not dense. Sinus acutely triangular, mem- 
brane not visible between the lobes. 

Female.—Genital segment equilaterally triangular in outline when 
the valves are adducted, small, setose, glabrous, and shining, not 
strongly chitinized. 

Valvula (Plate 4, fig. 17).—Dorsal plate suboblong, nearly three 
times longer than wide and not defined from the apex, nearly plane, 
finely and sparsely punctulate, punctures denser at apex, each with a 
rather long, delicate, flying hair; external border somewhat straight 
in basal half, evenly arcuate to apex; internal border more or less 
sinuous; apex with the outer lobe obsolete, the internal not defined 
from the general surface. Fossa and appendage obsolete. 

Superior pudendal membrane reaching to the apical third of the 
valves and longitudinaliy rugulose. 

Ventrolateral surfaces evenly convex, not inflated nor excavated, 
finely and sparsely punctate, setose as above; submarginal groove not 
developed; internal margins of the valves contiguous in basal fourth. 
Genital fissure long and evenly fusiform, not wide. Inferior puden- 
dal membrane visible in basal half. 


182 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Habitat—Texas (Alligator Head, Calhoun County, February, 
J. D. Mitchell, collector; Corpus Christi, Nueces @ounty, May). A 
series of nineteen specimens from the Hubbard and Schwarz collec- 
tions (U. S. National Museum) ; others from the collections of Fall, 
Fuchs, and Van Dyke. 

Number of specimens studied, 24. 

Type in the Horn collection. 

Ty pe-locality. Texas.” 

Salient type-characters.—Opaque, subdepressed, sparsely clothed 
with short yellowish hairs. Form elongate (male) or broadly 
(female) oval. Base of the pronotum squarely truncate and slightly 
overlapping the base of the elytra. Pubescence of elytra arranged 
so as to appear substriate. Sides of the body forming continuous 
and evenly arcuate lines from head to the elytral apices (Horn). 

Diagnostic characters —One of the most isolated species of the 
genus. 

The regularly oval form, equally narrowed anteriorly and _ poste- 
riorly, the thorax squarely truncate at base, with the sides regularly 
arcuate and gradually narrowing from base to the apex, are eharac- 
ters which serve to distinguish it from all other species of HJeodes. 

The anterior tarsi are not dilated in the male. There is a wonder- 
ful degree of homogeneity of structure among the individuals of the 
series before me—in fact, one would think that they had been cast 
in the same mold. It bears a superficial resemblance to opaca in its 
opaque integuments and sparse setiform pubescence, but the above 
characters will quickly separate it. 

The genital characters place it in a distinct subgenus. I consider 
it a primitive form that has undergone but little modification in re- 
cent years. 

Mentum variable, triangular, trapezoidal or evenly rounded at 
apex, convex at middle and more or less narrowly concave laterally. 
Prosternum usually horizontal between the coxe and sloping gradu- 
ally upward anteriorly to the anterior margin; rarely feebly arcuate 
between the coxe; feebly dilated behind the equator of the coxe, 
scarcely produced, vertically truncate behind, with angle rectangular, 
or feebly produced and sometimes feebly deflexed at the angle. 

Mesosternum vertically and arcuately declivous, more or less feebly 
concave. The intercoxal process of the abdomen is subquadrate and 
feebly transverse; equal in width to the third and fourth ventral 
segments taken together, the post-coxal part of the first segment is 
equal in length to the second; the third one-fourth of its length 
longer than the fourth. 

The abdominal salient is about one-fourth of its width broader 
than the metacoxal process. The metasternum laterally between the 
cox is a little shorter than the width of a mesotibia at apex. 


REVISION OF ELEODITNI—BLAISDELL. 183 


The tibial grooves of the femora are well defined and concave, 
glabrous and impunctate and prolonged inwardly upon the femora: 
their margins are subcarinate. External borders of the tibiz pune- 
tate and pubescent and not in the least carinate. 

The tarsi are slender and apparently slightly compressed. 

A protarsus is about one-third of its length shorter than a meso- 
tarsus; joints small, the first distinctly larger and longer than, the 
second, the latter about as long as wide; the third just slightly 
shorter, and the fourth still smaller, both slightly wider than long. 
In the female the first joint is less robust. 

The mesotarsi are about one-fourth of their length shorter than a 
metatarsus; the first joint of each is about as long as the third and 
fourth taken together; the second a little longer than wide, and the 
third and fourth about as wide as long, each gradually and slightly 
decreasing in size and length in the order mentioned. 

The metatarsi are about one-fifth of their length shorter than their 
metatibix ; first joint of each is just slightly longer than the next 
two taken together, the second distinctly longer than the third, the 
latter a little more than half as long as the fourth. 


Subgenus ELEODES Eschscholtz. 


In the present subgenus the species have the anterior femora more 
or less dentate in one or both sexes; the teeth are abortive in caudiferu 
and longipilosa. 

The anterior tarsi of both sexes are simply spinous beneath, or at 
the most the first joint may bear a small tuft of modified spinules in 
one or both sexes, and the first one or two joints are never dilated in 
the male, although in a few species the first joint may be thickened 
at apex beneath. 

The internal spurs of the meso- and metatibiz are longer than the 
external. 

For the positive recognition of the constituents of this subgenus 
the female genital characters must be relied upon. 

The male genitalia have undergone but little differentiation along 
the lines of specific specialization and are valueless for the diagnosis 
of species. 

They may be enumerated as follows: 

Subgeneric genital characters, male——Apicale of the edeagophore 
distinctly longer than wide; dorsal surface moderately convex, with 
a more or less complete membranous median groove, which at times 
is strongly developed, at others feebly so; apex acute and not dis- 
tinctly attenuated; base distinctly lobed at the middle. 

Female.—Genital segment more or less trapezoidal in outline and 
robust, conspicuously setose in apical moiety with evident tufts of 


184 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


hairs on the tips of apex and appendage, and also at the external angle 

of the fossa on or beneath the apical angle of the d6rsal plate. 

Internal lobe of the apex developed, the external rudimentary. 

Appendage variable, from conical to depresso-semi-ellipsoidal. 

Dorsal plate oblong and more or less strongly concave, the margins 
being reflexed. 

Submarginal groove more or less visible from above. 

Superior pudendal membrane never reaching beyond the middle 
of the dorsal plate and narrowly exposed. 

The following groups may be defined: 

All those species that have the first joint of the protarsi in both sexes thick- 
ened at apex beneath and bearing a tuft of modified spinules upon the same; 
hummer obtuse.22225 35 3.2. Se er ee aS ae ae OpscuRA Group. 

All those species that do not have the first joint of the protarsi thickened at 
apex beneath and without a tuft of modified spinules; grooves entire. 

EMM eT ODSOleCIe S23 e ee a eee GRANDICOLLIS Group. 
Mivmiert: obiSe a= 3 Set ee ee ee ee DENTIPES Group. 

The Dentipes Group is divided into the following sections: 

Section A—AIl those species which are genetically related by 
having the inferior surface of the cauda, when present, formed by 
the dilated apical extremity of the epipleure. Such species are more 
or less opaque and more or less asperately sculptured. 

Section B.—AlII those species which are genetically related by having 
ihe inferior surface of the cauda, when present, simply margined by 
the undilated apical extremity of the epipleure; the inferior surface 
of the cauda being formed by the general internal surface of the 
elytra. Such species are glabrous and more or less shining and never 
distinctly asperately sculptured. 


; OBSCURA Group. 


This group may be characterized by having the first joint of the 
protarsi distinctly thickened at apex beneath, and bearing a tuft of 
modified spinules upon the same. In the males the protibial spurs 
are somewhat equal in size, although the posterior may be slightly 
longer than the anterior; in the female the posterior spur is more or 
less thickened and distinctly longer, and in both sexes the spurs are 
decidedly divergent. 

The constituents of this group are among the largest of our species. 

The articular cavities of the tibiz are open and continuous with the 
tarsal grooves of the same. The middle lobe of the mentum and the, 
exposed surfaces of the basal divisions of the maxillary palpi are more 
strongly and conspicuously setose than in any other species known 
to me. 

The head appears to be comparatively long, and the abdomen is 
more or less distinctly oblique in the male. 


REVISION OF ELEODITNI—BLAISDELL. 185 


The group genital characters are as follows: 

Male.—Apicale of the edeagophore triangular, dorsal surface mod- 
erately convex, with a median longitudinal linear groove, and the 
base has a rounded lobe at middle. Sternite with an interlobar circu- 
lar membranous area. 

Female.—Dorsal plate canaliculate, sides reflexed and retracted, 
submarginal groove completely visible from above; apex but slightly 
longer than the appendage, the latter short and conic; superior pu- 
dendal membrane not longitudinally rugulose and not reaching pos- 
teriorly beyond the middle of the dorsal plate. 

The three species known to me may be separated as follows: 


Females.—The posterior spur of the protibiw distinctly larger and longer, more 
obtuse, than the anterior. 
Thorax convex and smooth. 
eth RATER STO UMOCd tse te obscura. 
Elytral margins acute at humeri_______~_-_ : ___acuta. 
The posterior spur very slightly thicker or longer than the anterior; both spurs 
gradually narrowing from base alike, both acute. 
Thorax flat or concave; elytral margin acute throughout _____- | suturalis. 
Males.—Anterior spurs of the protibiz quite similar; abdomen oblique. 
Thorax convex and smooth. 


SU ELE RES  POUICIOO <r ee OS CS ee ey obscura. 

Elytral margins acute at humeri______- cos ee thee YT omen: 
Thorax flat or concave. 

Riveral marcin acute’ throughout. 2.2 suturalis. 


The genital characters appear to present the following specific 
differences. The-males as usual present no distinct differential char- 
acters. 

Female.—Internal lobe of apex well developed. Genital segment trapezoidal 


in outline, robust; basal prominences well developed. 
Apical margin of the dorsal plate rounded. 


men i Wiskbein wevenl w POUmied A. ss oe obscura, 

Apical margin rounded externally and oblique within_____- suturalis, 
Apical margin truncate. 

PRRMNCES RTI IES AOTC IID in be se es os ee tt acuta, 


A linear arrangement of our species may be given as follows: 


{ dispersa. 

| deleta. 

| obscura. 
sulcipennis. 


Thorax convex 
arata, 


with its 


acuta two 
: extreme forms. 


suturalis, 


xX concave 
Thorax nea Raha 


186 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Genealogy.—The accompanying diagram will give my idea of the 
relationships of the species. It will be observed"that from the an- 
cestral trunk divergence probably occurred along two lines—(q@) one 
where the prothorax acquired acute margins, and consequently a con- 
cave disc; at the same time the spurs underwent but slight differen- 
tiation, and as a result, in the course of time there were produced 
forms, which being plastic to a moderate extent, became capable of 
being recognized as a series, the heterotypes of which are our suturalis 
and texana. These two races are united by true mesotypes. 

(4) The other line of divergence resulted in a less differentiation 
of the dorsal body wall, but a greater change occurred in the tibial 
spurs. Here the dichotomous changes resulted in the present acuta 
with less calearine development and obscura where the tibial calcars 
are the most differentiated. It appears as if acuta has reacted less 
to modifying factors, while obscura has gradually been the most re- 
active as evidenced by the recog- 


Texana : . 
| eas Suleipemsis nnizable races; the heterotypes being 
united by an unbroken line of 
Suturalis sito = 
< mesoty pes. 
mraccaccare Ngee ex Gone, HPT* So in the lineal arrangement of 
acute margins, spurs ff disparity between anterior the species and their modifications 


tibial spurs ws : . 
two distinct breaks in the chain 


FIG. 3.--GENBALOGICAL piaGRaM or THE are noticeable: First, sutwralis and 

OpscurA GROUP, SUBGENUS ELEODES. acuta are distinctly defined species 
as regards each other; second, acuta and sulcipennis (obscura) while 
less divergent in structure are not united by any known mesotypes, 
the heterotypes of acuta more likely only simulate obscura on the one 
hand and fexana on the other. These are only suggestions, for no one 
can be sufficiently positive to assume the true relationships of forces 
that bring about such interesting forms. 

T also wish to call attention to another fact: In dispersa the tarsal 
srooves of the tibia are the most strongly developed, and the articular 
cavities open the most widely into them, and these characters gradu- 
ally diminish so that in swtwralis they are found at their minimum 
in the present group. 

By the open articular cavities—the floors of which are directly con- 
tinuous with those of the tibial grooves, both being on the same plane 
in the most strongly differentiated examples—greater functional 
power is given to the tarsi, i. e., they must be more strongly extensible 
and capable of being more strongly brought into juxtaposition to the 
tibial surface. 

Distribution—As Known to me the present group is distributed 
through the following 15 States: Texas, New Mexico, Arizona, Kan- 
sas, Colorado, Utah, Nevada, northern California, Oregon, Washing- 
ton, Idaho, Wyoming, Nebraska, southern Dakota, and Oklahoma. 


REVISION OF ELEODITNI—BLAISDELL. 187 


Table of distribution. 


Suturalis. Texana, | Acuta, | Dispersa, Obscura. Sulcipennis, 
ies cbs. | Texas ..... TeXas -- ~~). 2-2 0e cere ee) ee eee scene 
| = | New Mex- |.........-.- New Mex- New Mex- 

1co. } Ico, 1co, 
a SS ee a ee [PATZONe.. . ios sass. e wc) ATIZONA. 
Kansas ..... PRRON a. ToReORES fo 6 fw a cee em alou soos sane e'= 
Pe eC CIA T OM sc eas Bee ceo en eo cen eliw ows ekecews 
Colorado ... Colorado .. Colorado ..| Colorado ..| Colorado -. 
—. Ap AOS ae Mas nk sonal NR ie Bg a pera Fal Db 8 
REET Re ee ee So ea ee, os eae cae vince ecw dae nec Nevada. 
mL ore eee SoC Babe omen ou >'s.sn frauen seme 6b’ Northern Cali- 
fornia. 

Catal oP aes ah ders ano clashes wh Glew ainew aed a's a Oregon ..... Oregon. 
eee ee Wh eo ees cee Se a xed |e few view Washington. 
0 woh a a a eee Pe et eee Dagon Binge) Idaho. 
oe ee BS eee ee Renee ee Wyoming 
NONPANBA 5}... 2.2 =.= prints ake od Sawer bt lemin are wa | 
USES CES a ee dots eae Re ae nes Uae eens | 


| 


It will be seen that sutwra/is inhabits the more central and noril:- 
eastern part of the region west of the Mississippi River; tevana, the 
southern and southern central; acuta, probably the same, like texana,; 
dispersa, central and more southern western; olsevra, central and to 
the northwest, western and northwestern, and probably none are 
found west of the Sierra Nevada Mountains. Su/cipennis extends 
southward into Sonora, Mexico (Morrison), and is probably the most 
widely dispersed member of the group, while déspersa is the least 
widely distributed. 

ELEODES OBSCURA Say. 


Blaps obscura Say, Journ. Acad. Nat. Sci. Phila., IIT, 1828, p. 259; Amer. 
Entomology, I, 1824, p. 32, pl. xvi, fig. 3—LeContTr, Complete Writ- 
ings of Th. Say, I, 1859, p. 32. 

Bleodes obscura Say, Proc. Acad. Nat. Sci. Phila., 1858, p. 181.—Horn, 
Trans. Amer. Phil. Soc., XIV, 1870, p. 306. 

Bleodes dispersa LeContr, Proc. Acad. Nat. Sci. Phila., p. 182.—Horn, 
Trans. Amer. Phil. Soc., XIV, 1870, p. 305. 

Eleodes deleta LeContr, Proc. Acad. Nat. Sci. Phila., 1858, p. 182.— 
Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 305. 

Eleodes sulcipennis MANNERHEIM, Bull. Soc. Nat. Moscow, XVI, 1545, 
p. 266; Mag. Zool., XIII, 1843, No. 128 (fig.).—L&EContTE, Proc. Acad. 
Nat. Sci. Phila., 1858, p. 182.—Horn, Trans. Amer. Phil. Soc., XIV, 
1870, p. 306.—CHaAmpPi0N, Biol. Centr. Amer., IV, Pt. 1, 1884, p. 79. 

Eleodes arata LeContr, Proc. Acad. Nat. Sci. Phila., 1858, p. 182.—Horn, 
Trans. Amer. Phil. Soc., XIV, 1870, p. 306. 


188 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Black or brownish, oblong, elongate. 

Head slightly more than one-half as wide as@the pronotum at 
middle, and slightly less than twice as wide as long; usually more or 
less convex; rather opaque, and coarsely, irregularly punctured, fre- 
quently with impunctate areas at middle; laterally densely punctate. 
Antenne moderate, reaching nearly to the prothoracic base; outer 
four joints just noticeably compressed, gradually and very slightly 
dilated, third joint scarcely as long as the next two taken together; 
fourth slightly longer than the fifth, the latter to the seventh, 
inclusive, subequal and longer than wide; eighth subtriangular and 
about as wide as long; ninth and tenth suborbicular; eleventh ovate. 

Pronotum shining or somewhat opaque, about one-fifth wider than 
long; disc evenly and more or less convex, generally declivous at the 
apical angles, glabrous, finely, sparsely, and irregularly punctate, 
punctures becoming denser and coarse along the sides; apex truncate 
or feebly emarginate, rather coarsely or more or less obsoletely mar- 
gined ; sides evenly and rather broadly arcuate to the posterior fourth, 
thence feebly sinuate or oblique to the basal angles, marginal bead 
moderately coarse and narrowly reflexed; base feebly arcuate, fre- 
quently slightly sinuate at middle, rather coarsely beaded and about 
one-sixth wider than the apex; apical angles obtuse; basal angles dis- 
tinct, more or less rectangular, sometimes very narrowly rounded. 

Propleure impunctate and rugulose; frequently sparsely punctate 
centrally. 

Elytra oblong to oblong-oval about twice as long as wide; base 
scarcely emarginate, scarcely to slightly wider than the contiguous 
prothoracic base; humeri slightly prominent and obtuse; s¢des evenly 
and not strongly arcuate, apes not produced and rounded; diése 
generally widest at or frequently behind the middle, more or less de- 
pressed and convex, laterally always more or less broadly rounded 
throughout, rather suddenly declivous posteriorly; surface more or 
less striate and sulcate, more or less irregularly, muricately punctured. 

Epipleure, sterna, and parapleure more or less obsoletely and 
rather densely punctate, rugulose. The epipleure are moderate in 
width and not more suddenly widened beneath the humeri. 

Abdomen finely and more or less obsoletely punctate, rugulose; 
fifth segment more densely punctate. 

Legs moderately long and quite stout; anterior femora armed in 
the sexes; anterior tarsi with the first joint (Plate 8, figs. 8 and 9) 
thickened and slightly produced at apex beneath, groove obliterated, 
the produced tip bearing a tuft of more or less golden (fresh speci- 
mens) modified spinules; other joints with the plantar groove entire, 
with the usual apico-ventral tufts of spinules separated by the 
groove; similar in the sexes. 


REVISION OF ELEODIINI—BLAISDELL, 189 


Male.—Elongate oblong, somewhat narrow. Antenne reaching to 
basal margins of the prothorax; elytra occasionally with sides par- 
allel or moderately arcuate, dorsum more or less distinctly depressed 
and quite suddenly obliquely declivous posteriorly. Abdomen moder- 
ately convex from side to side, broadly and more or less strongly im- 
pressed on segments one and two, oblique. Anterior femora armed 
with a strong, moderately acute tooth; tibial spurs similar; the an- 
terior about equal, rather stout and convexo-concave in transverse 
section. 

Female.—Elongate oval, robust. Antenne reaching to the pos- 
terior fourth of the pronotum. Elytra rather broadly oval, nearly 
vertically declivous posteriorly ; dorsum moderately convex. Abdomen 
horizontal, evenly and strongly convex from side to side. Anterior 
femora with an obtuse tooth, sometimes reduced to a mere sinuation ; 
protibial spurs dissiniilar (Plate 8, fig. 14), the posterior spur 
broader, more robust, obtuse, and a little longer than the anterior. 

Genital characters, male —Edeagophore elongately oblong-ovate. 

Basale oblong, strongly arched, stout and variable in form. 

Apicale triangular, elongate; sides straight, apex produced and 
acute; surface evenly convex, with a fine linear, membranous groove 
extending from the apex nearly to the base, the latter with a broad, 
rounded lobe at middle and not strongly sinuate laterally. 

Sternite (Plate 4, fig. 15).—Quadrate and strongly chitinized, with 
an interlobar circular membranous area. Lobes elongate, with the 
external borders subparallel in basal half and thence evenly arcuate 
to apex. Apex of each lobe moderately subacute; internal border 
short, straight to feebly arcuate; surface strongly and densely punc- 
tate in apical half, setose, setae quite long and very dense at apex, and 
scattered across the membrane at bottom of the sinus, the latter not 
conspicuously deep. 

Female.—Genital segment subtriangular, robust, strongly, sparsely 
punctate, and setose; setee moderately long, semi-erect, and rather 
evenly distributed. 

Valvula (Plate 1, figs. 138, 15, and 16).—Dorsal plate oblong, with 
the sides rather strongly reflexed, retracted, and subparallel; surface 
concave, sometimes slightly convex within near apex; internal margin 
slightly sinuous to subangulate at basal and middle thirds: apical 
margin more or less evenly rounded, not sharply defined. Sudmar- 
gmal groove entirely visible from above, limited externally by the 
margin of the lateral plate which is visible throughout its entire 
length. Apew short, slightly or scarcely more prominent posteriorly 
than the appendage, coarsely punctate and setose, sete not form- 
ing an apical tuft; external angle of the fossa forming the subapical 
angle, with sete rather condensed, 


190 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Appendage short, stout, and conical, setose, sete at tip forming a 
_ loose tuft. “ 

Superior pudendal membrane reaching to a point about midway be- 
tween the base and apex, not closely longitudinally rugulose. 

Basal prominences well developed. 

Valvular membrane more or less visible from apex to the pudendal 
membrane, and minutely setose. 

Ventrolateral surfaces—Body rather strongly inflated in basal 
half, rather deeply, broadly, and transversely concave before the 
apices, quite strongly setose. Internal margins of the valves con- 
tiguous for a very short distance at base and apex, intervening fissure 
fusiform, and closed in basal half by the inferior pudendal mem- 
brane. 

The present species is very variable, both as regards to form and 
sculpturing. The following three races are founded upon elytral 
sculpturing alone: 


ELEODES OBSCURA var. DISPERSA LeConte. 


Elytra with indistinct strie of closely placed punctures, intervals 
flat and quite densely, very irregularly and muricately punctured. 

Measurements—Males: Length, 25-33 mm.; width, 9.2-11.5 mm. 
Females: Length, 30-35 mm.; width, 12-14 mm. 


ELEODES OBSCURA Say (typical variety). 


Elytra feebly sulcate, intervals not strongly convex, sparsely, 
muricately punctate. Each stria with a series of closely placed 
feebly muricate punctures; intervals with a series of more distantly 
spaced and slightly more muricate punctures. Frequently the 
sculpturing is denser, always more strongly and irregularly so later- 
ally and on apex. 

Measurements —Males: Length, 26-30 mm.; width, 10.2-11.8 mm. 
Females: Length, 25-81 mm.; width, 10.5-13.2 mm. 


ELEODES OBSCURA var. SULCIPENNIS Mannerheim. 


Elytra strongly suleate; intervals quite strongly convex, smooth, 
with a single series of irregularly, distantly spaced, feebly muricate 
punctures, which become decidedly muricate on the apical declivity: 
sulci about equal in width to the intervals, with closely placed 
muricate punctures which become more densely placed toward apex; 
inflexed sides of the elytra obsoletely suleate and irregularly muri- 
cately punctured, 


REVISION OF ELEODIINI—BLAISDELL. 191 


Measurements —Males : Length, 25-31.5 mm.: width, 10.2-11.5 mm. 
Females: Length, 24-33.5 mm.; width, 8.5-13.8 mm. 

The following varietal genital characters may be correlated with 
the above forms of elytral sculpturing, and although they are not 
always as clearly defined, yet they seem to be sufficiently so as to war- 
rant their being recorded. 

- Dispersa.—Genital segment rather elongate, and generally more 
strongly chitinized and sculptured. 

Obscura.—Genital segment subtriangular, sides more convergent, 
apex less truncate; moderately strongly chitinized and sculptured. 

Sulcipennis.—Genital segment rather broadly trapezoidal, slightly 
elongate. Valves broadened apically, and not strongly chitinized nor 
sculptured. 

Habitat —Dispersa: Arizona (C. V. Riley; Coll. Beverly Letcher ; 
Keams District, Navajo Reservation, April, collected by A. W. Bar- 
ber) ; New Mexico (Coll. Charles Fuchs; H. F. Wickham; Coolidge, 
Hubbard and Schwarz; Aztec, April, Coll. C. F. Baker) ; Colorado 
(San Luis Valley, Mr. Bowditch). 

Obscura.—Colorado (Colorado Springs, August, Collector H. Sol- 
tau; Rocky Mountains, Coll. Hubbard, and Schwarz; Greenhorn, 
Trinidad, and San Luis Valley, Hayward); Wyoming (Cheyenne, 
April, Hubbard and Schwarz; May, H. Soltau, National Park, 
August, H. Soltau; Saratoga, Coll. E. C. Van Dyke); New Mexico 
(Santa Fé, Coll. C. V. Riley; Las Vegas, August, Coll. Barber and 
Schwarz) ; State of Washington (Coll. J. B. Smith); Oregon (Coll. 
C. V. Riley). 

Sulcipennis—Arizona (Bright Angel, July, Barber and Schwarz; 
Santa Rita Mountains, May, Hubbard and Schwarz; Pinal Moun- 
tains, Coll. E. C. Van Dyke; Fort Grant, July; Walnut, September, 
H. F. Wickham; Williams, July, Barber and Schwarz: Ash Fork, 
June; Nogales, September, Koebele and Nunenmacher; along the 
Colorado River, Charles Fuchs): Tdaho (Camas Creek, L. Bruner) ; 
Nevada (Hubbard and Schwarz) ; Oregon (Hubbard and Schwarz) ; 
State of Washington (C. V. Riley; Wawawai, Coll. C. V. Piper) ; 
northern California, Nevada (Doctor Horn). 


ELEODES DISPERSA LeConte. 


Number of specimens studied 20. 

Type in the LeConte collection. 

Ty pe-locality — Creek Boundary ;” collected by Doctor Wood- 
house (LeConte). 

Salient type characters.—Oblong, bright. Thorax moderately 
finely punctate; sides strongly rounded, toward base subsinuate; 


192 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


basal angles rectangular. Elytra elongate, densely punctate, punc- 
tures on the dorsum subseriate, laterally and on the apex trans- 
versely confluent and muricate (LeConte). 

Diagnostic characters.—Easily separated from obscura and sulci- 
pennis by its obsolete striz and finely muricate punctuation, by the 
comparatively more elongate and narrower form. Even in a moderate 
series the three races are united by an ample number of examples 
exhibiting all degrees of intermediate sculpturing. 

The mentum has the middle lobe moderate, and about as wide as 
long; sides more or less arcuate; apex also rounded, slightly reflexed, 
truncate or feebly emarginate; surface strongly punctate, slightly 
prominent along the median line, feebly and broadly concave laterally 
and clothed with stiff hairs that curve forward. 

The prosternum is triangularly dilated behind the equator of the 
coxe, and more or less protuberant ventrally with the same; pos- 
teriorly compressed, vertically truncate, and more or less produced; 
at times it is arcuately declivous. 

Mesosternum quite vertically declivous or oblique and more or less 
concave. 

The abdominal process is subquadrate, about one-sixth of its width 
wider than long, its width is also equal to the post-coxal portion of 
the same segment. The second segment is a little more than twice as 
long as the fourth; the third is about equal to the post-coxal part of 
the first; the fourth is about one-third of its length shorter than the 
third. 

The abdominal salient is about a sixth of its width greater than the 
same of the metasternal process. 

Metasternum laterally between the coxe is as long as the width of 
a mesotibia at apex. 

The tibial grooves of the femora are not entire. Those of the pro- 
femora are quite broad, the floors rather feebly concave, smooth and 
impunctate; the margins are strong and gradually converge to meet 
just before attaining the base, the anterior is more or less dentately 
laminate, forming an obtuse (female) or an oblique tooth (male). 

On the mesofemora the grooves become evanescent at internal third, 
but are well defined externally, floors feebly concave and smooth. 

The metafemora have the grooves evanescent at middle without con- 
vergence of the sides, which are well marked externally. 

The tibie are very densely muricate. Each protibia is moderately 
compressed and quite strongly carinate externally in basal half, where 
it is also more or less arcuate; the carina is smooth and at the middle 
of the protibia becomes the anterior boundary of the tarsal groove, 
where it also becomes muricate to the apex. The tarsal groove is 
quite strongly defined and its floor is glabrous and impunctate, open- 


— 


REVISION OF ELEODIINI—BLAISDELL. 193 


ing at apex into the articular cavity, the margins of which become 
continuous with the sides of the groove. 

Each mesotibia may be very briefly carinate at base externally; the 
external surface is quite deeply channeled, to a greater or less length 
by the tarsal groove, the sides of which are quite abrupt and densely 
muricate ; the groove opens into the articular cavity as in the protibia, 
but is narrower and the cavity is less widely open. The floor of the 
groove is quite glabrous. The metatibiwe are obsoletely carinate at 
base externally and also with the tarsal grooves running nearly the 
whole length; each groove is rather shallow and not suddenly 
formed, densely muricate and continuous with the articular cavity. 

The tarsi are robust. 

The protarsi are about a fourth of their length shorter than a 
mesotarsus. The first joint is about one and a half times longer 
than wide; the second, third, and fourth joints are subequal in size 
and length, wider than long, and together about equal to the fifth. 
Each mesotarsus is about a third of its length shorter than a meta- 
tarsus. The first and fifth joints are quite equal in length; the 
second, third, and fourth are subequal, just visibly longer than wide, 
and together longer than the fifth. 

The metatarsi are about a half of their length shorter than a 
metatibia; second and third joints subequal and distinctly longer 
than wide and together about equal to the fourth, the latter about 
equal to the first. 


ELEODES OBSCURA Say. 


Number of specimens studied, 30. 

Type, destroyed. 

Type-locality—* Country bordering the River Platte within a 
hundred miles of the Rocky Mountains.” (Say). 

Salient type-characters—Say’s description is too brief and in- 
definite to give a clear idea of the type: “ Elytra scabrous, grooved, 
margin rounded; thoracic margin not reflected.” 

Diagnostic characters —KEasily recognized from its races by the 
feebly suleate and scabrous elytra. It is also less elongate. 

The prosternum is convex antero-posteriorly, subtruncate behind, 
vertical or oblique and more or less compressed. 

Mesosternum arcuately declivous and more or less concave. 

The tibial grooves of the femora are about as in dispersa. 

The sculpturing of the tibiz is variable, sometimes strongly muri- 
cate and at others subscabrous. Otherwise the tibial characters are 
relatively the same as in dispersa. The articular cavities are really 
less open, as the margin is frequently continued across the confines 

59780—Bull. 63—09——-13 


194 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


between the groove and cavity; the tarsal grooves are usually more 
contracted and more apt to be roughly sculptured 

The relative widths and lengths of the tarsal elements, abdominal 
segments, and sterna are about the same as in the preceding variety, 
and the analysis of its exoskeletal characters indicates its truly inter- 
mediate position. 


ELEODES SULCIPENNIS Mannerheim. 


Number of specimens studied, 54. 

Type in the Mus. Imp. Acad. Scient. St. Petersburg. 

Type-locality is (northern) California. 

Salient type-characters.—Elongate. Thorax subcordate, sides nar- 
rowly reflexo-marginate, finely punctulate, elytra oblong and deeply 
sulcate, posteriorly very slightly dilated, apex attenuate and strongly 
declivous (Mannerheim). 

Diagnostic characters.—Quickly separated by the strongly sulcate 
elytra. 

Some specimens from Colorado are intermediate between obscura 
and sulcipennis, and have the scabrous elytra of the former and the 
distinct sulci of the latter. 

LeConte described a female collected in New Mexico by Captain 
Pope as deleta. It was only an intermediate variation and can be 
placed as forma deleta, connecting dispersa and obscura. It was 
characterized by LeConte as follows: Form of sulcipennis, absence 
of elytral grooves, except at the sides behind the middle, where some 
faint traces are seen; the punctures are submuricate and arranged in 
strie distinct on the back, confused at the sides; between the rows 
are distinct punctures as in obscura; posteriorly abruptly declivous 
and furnished with rows of tubercles, alternately large and small, 
the latter corresponding to the striz of the anterior portion. Ante- 
rior femoral teeth very obtuse. 

Other specimens, male and female, collected in Arizona, which are 
the heterotype of sulcipennis, were also described by him under the 
name arata. 

These strongly developed specimens can be labeled forma arata, 
and may be recognized from the following brief description. 


ELEODES ARATA LeConte. 


Form of sulcipennis, thorax more convex, less flattened and less 
punctured at the sides, and more finely margined; elytral grooves 
deeper, interstices smooth and shining, with much fewer scattered 
punctures. ; 

The student will recognize the five forms, just considered, as ex- 
pressions of variations in the same species, and these forms may be 
arranged in the following morphological sequence: dispersa, deleta, 


EE 


REVISION OF ELEODIINI—BLAISDELL. 195 


obscura, sulcipennis, and arata; which makes the series complete as 
far as known. 

The mentum is more or less distinctly trapezoidal, rather large; 
apex truncate, feebly emarginate or slightly arcuate; surface coarsely 
punctate, fovex more or less obsolete, and rather densely setose ; sete 
black, moderate in length and directed forwards. 

The prosternum is slightly convex between the cox, compressed 
and usually vertically truncate posteriorly, the inferior angle sub- 
mucronate or rounded, sometimes oblique, or more rarely evenly 
rounded antero-posteriorly. 

Mesosternum variable as in dispersa and obscura. 

The tibial, femoral, tarsal, abdominal, and sternal characters are 
practically the same as for the other races, but dispersa appears to 
have the most pronounced form of sculpturing, not including the 
elytral sulci. 

ELEODES ACUTA Say. 


Blaps acuta Say, Journ. Acad. Nat. Sci. Phila., III, 1823, p. 258; Amer. 
Entomology, I, 1824, p. 31, pl. 1, fig. 1—LeEContTEr, Complete Writings 
of Thomas Say, I, 1859, p. 31. 

Bleodes acuta LeContr, Proc. Acad. Nat. Sci. Phila., 1858, p. 181.—Horn, 
Trans. Amer. Phil. Soc., XIV, 1870, p. 306. 


Black, elongate oblong, depressed dorsally, usually more or less 
reddish along the suture. 

Head nearly twice as wide as long, and a little less than one-half 
as wide as the pronotum at middle, feebly convex and more or less 
transversely impressed along the frontal suture, rather opaque, mod- 
erately coarsely, irregularly, and densely punctate. Antenne quite 
stout, not reaching to the prothoracic base, outer four joints scarcely 
dilated or compressed, the third about equal to the next two com- 
bined, fourth slightly longer than the fifth, the latter to the seventh 
inclusive, very slightly decreasing in length in the order named, the 
seventh just a little shorter than the fourth, eighth subtriangular and 
scarcely as wide as long, ninth and tenth subglobular, eleventh short 
ovate. 

Pronotum usually widest at the middle and about three-eighths 
wider than long; disc more or less glabrous and shining, finely and 
sparsely punctate, laterally more or less impressed along the margin ; 
impressed area opaque, irregularly or transversely rugulose and 
finely granulate; apea moderately emarginate and finely margined; 
sides broadly and evenly arcuate to near the base, then more or less 
sinuate and more coarsely margined, margin narrowly reflexed; base 
truncate to feebly arcuate, feebly and broadly sinuate at middle, 
coarsely margined, and about one-third wider than the apex; apical 
angles obtuse; basal angles more or less rectangular, 


196 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Propleure opaque, smooth, very sparsely and finely punctate; 
usually more or less rugulose. ” 

Elytra oblong to elongate oval, nearly twice as long as wide, feebly 
shining, at times more or less opaque; base feebly emarginate or 
sinuate laterally within the humeri beneath basal angles of the 
pronotum, scarcely to slightly wider than the contiguous protho- 
racic base; humeri scarcely prominent, obtuse; sides evenly and not 
strongly arcuate, frequently subparallel at middle, apex not pro- 
duced and rather broadly rounded; disc quite strongly depressed, 
slightly convex, suddenly deflexed laterally with margin narrowly 
rounded but acute at humeri, posteriorly more or less suddenly 
obliquely declivous; surface sulcate, intervals slightly convex with a 
single series of irregularly placed punctures, first four sulci with a 
single series of fine muricate punctures, the remaining outer sulci 
more densely muricately punctured; apical declivity more strongly 
suleate and muricate; inflexed sides obsoletely sulcate and irregu- 
jarly muricately punctate. Scutellum broad and impunctate. 

Epipleure moderate in width, somewhat rapidly but gradually 
and slightly dilated immediately beneath the humeri; surface gla- 
brous and shining, more or less obsoletely or finely and very sparsely 
punctate. 

Sterna and parapleure more or less obsoletely or strongly punc- 
tate and rugose. 

Abdomen finely and more or less obsoletely punctate and rugulose; 
fifth segment more strongly punctate. 

Legs moderately long and stout, and the anterior femora are armed; 
anterior tibial spurs dissimilar in the sexes; first joint of the protarsi 
thickened and feebly produced in apical half beneath, and bearing a 
tuft of golden pubescence, nearly similar in the sexes. 

Male.—Elongate. Antenne reaching to the basal fifth of the 
prothorax. Elytra nearly twice as long as wide, with the sides 
slightly arcuate or subparallel, dorsum strongly depressed, quite rap- — 
idly arcuate and obliquely declivous posteriorly. Abdomen oblique, 
moderately convex, broadly impressed on first two segments. Femora 
much thickened and armed with a strong, oblique, slightly produced 
subacute tooth; protibial spurs similar and about equal; first joint of 
the protarsi with a small, dense, obtusely rounded tuft of yellowish 
pubescence on the produced tip beneath, groove obliterated. 

Female.—Robust. Antenne reaching to about the posterior third 
of the prothorax. Elytra broadly oval, with dorsum moderately de- 
pressed and convex, about one-half longer than wide, and nearly 
vertically declivous posteriorly. Abdomen horizontal, evenly and 
strongly convex. Profemora with a small obtuse tooth; tibial spurs 
long, the posterior spur of the protibixw more robust, slightly longer 
and more obtuse, more strongly divergent than the anterior; first 


REVISION OF ELEODIINI—BLAISDELL, 197 


joint of the protarsi obliquely produced beneath and clothed with 
short pubescence at tip, forming an acute transverse tuft, groove 
apparently obliterated. 

Measurement.—M ales: Length, 28-35 mm.; width, 13-15 mm. 
Females: Length, 30-33 mm.; width, 13.5-16.2 mm. 

Genital characters, male.—KEdeagophore of the usual oblong-ovate 
form. 

Basale oblong, stout, and more or less arched, variable. 

Apicale slightly longer than wide, triangular, sides straight to 
feebly arcuate, becoming slightly sinuate before the apex, the latter 
somewhat produced and acute; base broadly lobed at middle and 
more or less sinuate laterally; surface evenly convex, grooved at mid- 
dle three-fifths, groove more or less linear, with sides impressed, 
forming a fusiform depression. 

Sternite broader than long. Lobes moderate, each with the exter- 
nal border rather broadly arcuate; apex obtuse, and the internal 
border is short and quite straight; surface densely punctate in apical 
half, densely setose in apical third, sete moderately long and not 
extending upon the membrane across the bottom of the sinus, the 
latter nearly closed by the former. The lobes are separated centrally 
at base by a transversely oval membranous area, just cephalad to the 
sinus, the inner borders of the lobes at base are thus rendered semi- 
circularly sinuate. 

Female.—Genital segment subtrapezoidal, robust, rather strongly, 
sparsely punctuate and setose ; sete rather long, flying, semi-erect, and 
quite evenly distributed. 

Valvula (Plate 1, fig. 17).—Dorsal plate oblong, with sides quite 
strongly reflexed and parallel, scarcely retracted; external border 
quite straight; internal slightly angulate and sinuate near base; api- 
cal margin quite truncate, angle evident and set with longer sete ; sur- 
face strongly concave. Submarginal groove shallow and broad at 
base, visible from above but less strongly defined toward apex. Apex 
short, finely setose, and noticeably longer than the appendage. 

Basal prominences strongly developed and setose. 

Appendage short, not prominent, conical, setose, with a loose pencil 
of quite long setz at tip. 

Valvular membrane more or less visible in the intervalvular space, 
caudad to the pudendal membranes. 

Superior pudendal membrane reaching about midway between the 
base and apex, not longitudinally rugulose. 

Ventrolateral surfaces not very strongly inflated, setose in apical 
half, scarcely transversely. concave before the apex; internal margins 
of the valves contiguous in basal half, genital fissure rather widely 
fusiform and the inferior pudendal membrane is not visible. 


198 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Habitat—Texas (Beeville, Coll. Hubbard and Schwarz); Kansas 
(Gove County, elevation 2,500 feet, F. H. Snow) ; @olorado (Denver, 
April, H. Soltau) ; Dakota (Buffalo Gap, Collector A. E. Hill). 

Number of specimens studied, 28. 

Type destroyed. 

T ype-locality —* Near Council Bluffs, Missouri” (Say). 

Salient type-characters.—Elytra scabrous, grooved ; sutural margin 
reddish-brown; external edge narrowly rounded and acute at humeri. 
Thoracic margin not reflected (Say). 

Diagnostic characters—From obscura it is known by the elytra 
being quite suddenly inflexed laterally and therefore very narrowly 
rounded at the sides, and by the margins being acute at the humeri; 
from suturalis by the pronotal disc being convex and not reflexed 
laterally. 

In some specimens the elytral margins are subacute and the dise 
quite flat, so that the passage to suturalis becomes easy. The form 
of the pronotum is quite constant. 

The heterotypes of the series before me resemble obscura on the one 
hand and ¢exana on the other. 

The middle lobe of the mentum is moderate, subtrapezoidal or sub- 
parabolic, apex arcuate or subtruncate with the sides straight and 
converging to apex; surface strongly and rather densely punctate, 
scarcely convex, obsoletely foveate laterally and quite densely setose. 

The prosternum is strongly prominent ventrally with the coxe, 
surface usually nearly horizontal, moderately compressed behind and 
vertically truncate, scarcely produced. These characters are most 
often observed in those specimens which have the elytral dise quite 
flat and margins subacute; frequently the prosternum is convex 
antero-posteriorly and oblique behind, these characters being observed 
in those examples with convex elytral disc and more broadly rounded 
margins; rarely it is continuously convex between the coxe. 

Mesosternum variable. Usually obliquely declivous and more or 
less concave; frequently quite arcuately declivous to somewhat ver- 
tically so and more or less feebly concave. 

The abdominal intercoxal salient is subquadrate, slightly transverse, 
and about as wide as the post-coxal portion of the first segment is 
Jong; third segment about one-third of its length shorter than the 
second; the fourth one-fourth shorter than the third. 

The metasternal process is scarcely one-half narrower than the 
abdominal. Metasternum laterally between the coxe is as long as 
the width of a mesotibia at apex. 

The tibial grooves of the femora are entire on the profemora and 
variable in length on the meso- and metafemora. 

The profemora have the tibial grooves rather broad, with the 
sides slightly arcuate and converging quite to the femoral base; their 


REVISION OF ELEODIINI—BLAISDELL. 199 


floors are glabrous and impunctate; margins scarcely acute, the an- 
terior more or less dentately laminate at about junction of the fourth 
and outer fifth; tibial grooves of the mesofemora are usually short, 
margins evanescent, and the inferior surface rounded to base and 
sculptured as usual; the grooves on the metafemora are similarly 
developed. 

The protibiz are more or less carinate externally, straight or mod- 
erately arcuate, and more or less moderately compressed; the tarsal 
grooves-are not usually well developed, roughly sculptured, and open 
into the articular cavities. 

The mesotibiz are flattened externally and the tarsal grooves are 
usually well developed and variable in length, with floors glabrous 
and sides well defined by muricate edges; they open into the articular 
cavities. 

The metatibe are flattened and more or less broadly marked by a 
shallow groove externally; the floors of the grooves are muricately 
sculptured and the articular cavities are open. 

The tarsi are stout. 

The protarsi are about one-half of their length shorter than a 
mesotarsus. Joints two, three, and four are subequal, short, and 
wider than long; the first is slightly longer and quite abruptly nar- 
rowed at base; the fifth is scarcely longer than the three preceding 
ones combined. 

The mesotarsi are one-sixth of their length shorter than a meta- 
tarsus. The first and fifth joints are about equal in length; the 
second, third, and fourth are subequal and a little longer than wide, 
and together distinctly longer than the fifth. 

The metatarsi are a little more than one-half of their length 
shorter than their metatibia. The first and fourth joints are sub- 
equal in length; the second and third are subequal and distinctly 
longer than wide, and together about equal to the fourth. 


ELEODES SUTURALIS Say. 


Blaps suturalis Say, Journ. Acad. Nat. Sci. Phila., ITI, 1823, p. 257; Amer. 
Entomology, I, 1824, p. 30, pl. xvi, fig. 2—LeContr, Complete Writings 
of Thomas Say, I, 1859, p. 30. 

Bleodes suturalis LeContr, Proc. Acad. Nat. Sci. Phila., 1858, p. 181.— 
Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 306. 

Var. terana LEContTE, Proc. Acad. Nat. Sci. Phila., 1858, p. 182; in 
Thomson’s Arcana Nature, III, 1860, p. 124, pl. xn, fig. 5, female.— 
Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 306; Trans. Amer. Ent. 
Soc., V, 1874, p. 34. 


Oblong, more or less. strongly elongate, dorsum flattened and 
slightly concave, black, frequently with a broad reddish band along 
the elytral suture, epipleure often tinged with the same color. 


200 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Head a little less than twice as wide as long, and scarcely one-half 
as wide as the pronotum; surface plane to slightly gonvex, frequently 
more or less impressed along the frontal suture, sometimes trans- 
versely so between the eyes, and laterally within the moderately 
prominent sides of the frons, opaque, moderately, coarsely, irregu- 
larly, and densely punctate, usually with small impunctate areas. 
Antenne rather stout, scarcely reaching the prothoracie base; outer 
four joints slightly compressed and just perceptibly dilated; third 
joint about equal to the next two combined; fourth, distinctly longer 
than the fifth; the latter to the seventh, inclusive, subequal and 
slightly longer than wide; eighth, triangular and about as long as 
wide; ninth and tenth, suborbicular; eleventh, short ovate. 

Pronotum widest at the middle and about one-half wider than 
long; disc opaque, smooth, slightly convex, finely and sparsely pune- 
tate, with small impunctate areas about the center, frequently with 
irregular impressions; laterally longitudinally impressed from within 
the apical angles to a very short distance in advance of the basal 
angles, terminating in feeble basal impressions, the depressions are 
generally transversely rugulose; apex deeply emarginate and more 
or less obsoletely margined; sides broadly and more or less strongly 
reflexed, evenly arcuate or sometimes very feebly and broadly angulate 
at middle, slightly sinuate in front of the basal angles, marginal 
bead moderately coarse; base truncate and feebly trisinuate, dis- 
tinctly margined, two-fifths to one-half wider than the apex; apical 
angles acute, subacuminate, prominent and more or less everted; 
basal angles rectangular. 

Propleurw opaque and smooth, very finely and sparsely to obso- 
letely, muricately punctate, more or less rugulose at times, and de- 
fined from the reflexed pronotal margin by a longitudinal concavity. 

Elytra oblong, one-third to twice as long as wide and more or less 
opaque; base feebly emarginate, and abont equal to the contiguous 
prothoracie base; huwmeri obtuse and not prominent, rounded be- 
neath the basal angles of the pronotum; sides evenly arcuate to sub- 
parallel, apex scarcely to feebly produced; disc plane to slightly 
convex, very suddenly deflexed laterally, angle of deflexion forming 
an acute and moderately reflexed margin, which becomes obsolete a 
short distance before the apex, more or less suddenly obliquely de- 
clivous posteriorly ; surface suleate, intervals feebly convex, each with 
a single series of rather distantly placed punctures, the four inner 
sulci with but a single series of closely placed submuricate punctures, 
remaining sulci with numerous closely and irregularly placed punc- 
tures, which become denser and rather more strongly muricate toward 
margin; apical declivity somewhat more strongly sulcate and sea- 
brons: inflexed sides not convex, obsoletely sulcate, irregularly and 
muricately punctured. 


Ts 


REVISION OF ELEODIINI—BLAISDELL, 201 


Epipleure moderately narrow, uncinately dilated beneath the 
humeri, and gradually narrowing to apex; surface usually more or 
less obsoletely punctate. 

Sterna and parapleure more or less obsoletely or strongly punctate 
and rugulose. 

Abdomen finely and more or less sparsely, obsoletely punctate and 
rugulose. 

Legs moderate. Anterior femora armed in the sexes; protibial 
spurs and protarsi nearly alike in the sexes, the spurs are quite 
strongly divergent. The first joint of the protarsi is more or less 
thickened and slightly produced at apex beneath, bearing a tuft of 
yellowish pubescence. 

Male.—About twice as long as wide... Antenne scarcely reaching 
to the basal margin of the prothorax. Elytra moderately, suddenly, 
and obliquely declivous posteriorly; apex slightly acuminate. Ab- 
domen slightly oblique, moderately convex, broadly impressed on the 
first two segments. Anterior femora with an acute tooth about one- 
fourth distance from the apex; posterior spur of the protibie ap- 
parently a little longer and slightly stouter than the anterior, fre- 
quently they appear to be quite equal in length, both are rather stout 
and acute; first joint of the protarsi with the produced tip beneath 
rather thick and bearing a small obtuse tuft of modified spinules, 
groove not evident. 

Female—tess than twice as long as wide. Antenne reaching to 
about the posterior fifth of the prothorax. Elytra quite suddenly 
obliquely or vertically declivous posteriorly. Apex obtuse. Abdomen 
horizontal, evenly and strongly convex. Anterior femora with a 
small obtuse tooth, sometimes scarcely more than sinuate in outer 
fourth; posterior spur of the protibi a little longer and stovter than 
the anterior, both are acute, moderately thick, and gradually narrowed 
from the base; first joint of the protarsi slightly and transversely 
produced at tip beneath, bearing a transverse tuft of spinules, which 
is more or less acute, groove more or less obsolete. 

The male genital characters do not apparently show any racial 
differentiation. 

Male—FEdeagophore of the usual oblong-ovate form. 

Basale oblong, scarcely arched, and may be sparsely punctate 

laterally at apex. 
_ Apicale rather broadly triangular, moderately depressed, surface 
more strongly convex apically, with a median-membranous groove in 
apical half; sides rather straight to slightly arcuate; apex scarcely 
produced and more or less deflexed, subacute; base broadly lobed at 
middle, and sinuate laterally. (See Plate 1, fig. 19.) 

Sternite transverse. Each lobe with the external border more or 
less evenly arcuate, and the internal short and straight to feebly 


202 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


arcuate, with apex rounded; surface densely punctate and setose in 
apical ‘two-thirds, setee quite long and not extending upon the mem- 
brane across the sinus; the latter nearly closed by the same. 

The lobes internally at base and cephalad to the sinus are rendered 
semicircularly sinuate by an interlobar transversely oval membranous 
area, the membrane of which is frequently transversely rugose. 

By the form of the elytra two races may be recognized as follows: 


ELEODES SUTURALIS (typical variety). 


Oblong, sides of the elytra evenly arcuate from base to apex, size 
smaller. 

Measurements—Males: Length, 26-26 mm.; width, 11-11.2 mm.; 
Females: Length, 21.5-28 mm.; width, 9.5-12.2 mm. 


ELEODES SUTURALIS var. TEXANA LeConte. 


Elongate oblong. Sides of the elytra more or less distinctly 
parallel, size larger. 

Measurements—Males: Length, 30-35 mm.; width, 11-14 mm.; 
Females: Length, 30-82 mm.; width, 11-13 mm. 

Female genital characters—Suturalis: Genital segment subtrape- 
zoidal, slightly elongate, rather robust and setose, the flying semi- 
erect hairs not very long and somewhat evenly distributed. 

Valvula (Plate 1, fig. 14).—Dorsal plate oblong, with the sides more 
or less reflexed, scarcely retracted, quite parallel; external border 
quite straight, the internal more or less arcuate and the apical margin 
is rounded and prominent externally, becoming inwardly oblique 
internally; surface deeply concave, finely, sparsely punctate and 
setose, seta quite long. Submarginal groove visible from above, 
broad and shallow toward base, fine toward apex, the side of the 
lateral plate visible and limiting it externally. 

Apex short and inconspicuous, scarcely more prominent posteriorly 
than the appendage, finely setose. 

Appendage short, conical, and inconspicuous, with a pencil of rather 
long hairs at tip; margin of the fossa with many sete, especially at 
apical margin of the dorsal plate, setee soft and moderately long. 

Basal prominences moderate. 

Superior pudendal membrane reaching about one-third to one-half 
of the distance from the apex and not rugulose. 

Ventrolateral surfaces—Body rather strongly and _ transversely 
inflated-in basal half, moderately transversely concave laterally be- 
fore the apices; surface finely punctate and setose in apical half. 
Internal margins of the valves contiguous for a short distance at 
base. Genital fissure long and fusiform. 

Female genital characters.—Tewana.—Genital segment subtrape- 
zoidal, rather short, robust, and setose. 


REVISION OF ELEODIINI—BLAISDELL, 2038 


Valvula (Plate 1, fig. 23).—Dorsal plate oblong, rather narrow, 
sides strongly reflexed, retracted externally ; external border more or 
less broadly sinuate, the internal angulate at basal third, and the api- 
eal margin is more or less truncate and rather prominent at angle, the 
latter more or less rounded; surface very deeply concave, finely and 
very sparsely punctate, setose, sete small. 

Apex short, tip very blunt and not prominent, slightly longer than 
the appendage, with sparsely placed and short sete, without pencil 
at tip. 

Submarginal groove not always strongly marked and entirely visi- 
ble from above; sides of the lateral plate largely visible externally. 

Appendage short, conical, and broadly rounded at tip and not 
conspicuous; setose, with apical tuft of moderately short hairs. 

Basal prominences strongly developed. Superior pudendal mem- 
brane reaching scarcely half way from the base, and with two or 
three longitudinal grooves. 

Ventrolateral surfaces——Body quite strongly inflated at base and 
transversely concave before the apices, strongly so laterally, sparsely 
setose; apex rather thick—viewed from below. Internal margins of 
the valves contiguous in about basal third. Genital fissure rather 
broadly fusiform. 

Habitat—Suturalis (var. typica)—Texas (C. V. Riley); Colo- 
rado (Denver, October, H. Soltau; Golden, May, Dyar and Caudell; 
La Junta, Berkeley, Orchard, canyons near Boulder, H. F. Wick- 
ham); Nebraska (Dodge County, collections, C. V. Riley, Hubbard 
and Schwarz, and F. E. Blaisdell) ; Kansas (Riley County, July to 
August, Popenoe; Wallace County, elevation 3,000 feet, F. H. Snow) ; 
Oklahoma (Mangum, September, collection National Museum) ; Da- 
kota (Buffalo Gap, A. E. Hall; Volga, collector, E. C. Van Dyke; 
Southern at Alexandria). 

Var. tevana LeConte-—Oklahoma (Mangum, September, H. 8. 
Barber); Kansas (collections of Snow, Fuchs, Hubbard, and 
Schwarz) ; Colorado (Robinson, coll. E. C. Van Dyke) ; New Mexico 
( Alberquerque, coll. National Museum) ; Texas (San Diego, October, 
Hubbard and Schwarz; Bosque County, G. W. Belfrage). 

Suturalis.—Number of specimens studied, 50. 

Type destroyed. 

Type-locality apparently not mentioned by Say. 

Salient type-characters.—Reddish-brown along the suture. Thorax 
with the edge deeply concave in front, lateral margin dilated and 
reflected; anterior angles with a small excurved point. Elytra 
scabrous, grooved, lateral. edge reflected, slightly elevated and acute 
(Say). 

Var. tevana LeConte.—Number of specimens studied, 20. 


204 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Type in the LeConte collection. 

Type-locality— Ringgold Barracks, Texas. 

Salient type-characters—Thorax with the dise slightly convex, 
sides broadly depressed and slightly reflexed, sides greatly rounded, 
subsinuate behind; anterior angles acutely acuminate; basal angles 
rectangular. Elytra with dorsum plane, sides parallel and margined 
(LeConte). 

Diagnostic characters—Conspicuous among its congeners by the 
acute and more or less reflexed elytral margins, with the pronotal 
margins acute and reflexed and the disc concave. 

The typical race is less elongate with the elytral margins distinctly 
arcuate; var. fevana differs in its elongate and parallel form. 

The form is quite constant in this species. The mentum is of 
moderate size and subtrapezoidal in form; frequently more or less 
arcuate anteriorly, at other times the sides are straight, converging 
anteriorly and the apex more or less truncate, often the apex is 
rounded and narrowly inflexed, giving it a truncated appearance; 
the surface is scarcely convex, coarsely punctured, more or less even 
or ridged along the middle, feebly and broadly foveate laterally 
and setose, setee short, scarcely dense and less conspicuous than in 
the other species of the group. 

Prosternum very variable and as in acuta. In the more strongly 
developed examples of tevana, the intercoxal prosternal surface is 
shghtly oblique, downward and backward so that the. posterior 
angle is prominent and subacutangular to the strongly truncate and 
vertical posterior edge. 

In the smaller specimens of sutwralis the prosternum is strongly 
convex antero-posteriorly. 

The mesosternum is variable as in acuta. The abdominal inter- 
coxal process is nearly quadrate, its width is equal to the length of 
the third segment; the second segment is about equal to the post- 
coxal portion of the first, and twice as long as the fourth; in texana 
it is a little longer; the third is about one-fourth of its length shorter 
than the second segment. 

The abdominal salient is about one-sixth of its width wider than 
the metasternal process (males) or subequal (females). 

The metasternum laterally between the coxe is about as long as 
the width of a mesotibia at apex. 

The tibial grooves of the profemora are nearly entire, their sides 
are well defined and the floors are glabrous; the anterior margin of 
each is dentately laminate at about the outer fourth, it may be simply 
sinuate. 

The grooves of the meso- and metafemora are poorly defined and 
short, scarcely ever extending beyond the middle and generally only 


REVISION OF ELEODIINI—BLAISDELL. 205 


present in the external fourth; their margins are rapidly evanescent 3 
when the grooves reach the middle of the femora the margins are 
about contiguous. 

The protibiz are more or less distinctly carinate externally, slightly 
compressed, and the external surface line is more or less arcuate from 
apex to base; the articular cavities are open, but less so than in the 
other members of the group, and the tarsal grooves are poorly defined, 
usually scabrous, sometimes more or less glabrous. The articular 
savities of the meso- and metatibizw are nearly or quite closed. The 
external surface of the mesotibiw is more or less flattened and more 
or less feebly grooved, scabrous, scarcely ever glabrous except near 
the apex. 

The metatibie are scarcely flattened or grooved externally, 
although they are at times, the surface is scabrous. The mesotibix 
are sometimes feebly and briefly carinate at base. 

The tarsi appear to be variable in stoutness without relation to 
the size of the specimen; they are slightly more elongate than in the 
other species of the group. 

A protarsus is about one-third of its length shorter than a mesotar- 
sus. The first four joints are short and subequal, the first slightly 
the longest. the others wider than long; the fifth is about equal to 
the preceding three taken together. 

A mesotarsus is but slightly shorter than a metatarsus. Joints 
two, three, and four are subequal and a little longer than wide, taken 
together longer than the fifth; the first is scarcely equal to the next 
two combined. 

The metatarsi are just a little more than one-half as long as their 
respective metatibia. Joints two and three are subequal and dis- 
tinctly longer than wide, and together equal in length to the first 
and fourth individually. 


GRANDICOLLIS Group. 


The single species constituting this group has been the only disturb- 
ing element thus far met with in the study of the present subgenus. 

It is without doubt intermediate between the Obscura and Dentipes 
groups, or to be more exact between obscura and dentipes themselves. 
At first I was inclined te treat it as a subsection of section B of the 
Dentipes group. In that position it was completely isolated and far 
from being a contributing member to the harmony exhibited by the 
other species. 

In the smooth and shining integuments it is allied to the Dentipes 
group; its large size, oblong form, broadened tibial spurs, and oblique 
abdomen of the male show strong affinity with the Obscura group, 
while the obsolete humeri make it a unique element in the subgenus, 


206 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


The elytra at base are not margined nor adapted to receive the pro- 
thoracic base, as is the case in the majority of species of the Dentipes 
group; on the other hand the elytral base declines, corresponding 
exactly in structure with obscura. 

In either group it would prove a disturbing element and the only 
course to pursue in the search for taxonomical harmony is to place 
it in a group as here indicated. It is also very evident that it is a dis- 
tinct species, which has undergone practically no differentiation into 
distinct races—in this it is analogous to veterator—although it is 
slowly yielding to the modifying influences of environment. 

It probably represents a very slightly changed branch of the com- 
mon ancestral stem from which the earlier antecedents of the present 
Obscura and Dentipes groups took their origin. In a genealogical 
scheme it must be placed to show the greatest affinity for dispersa. 
Divergent elements can not be arranged in a linear series of mor- 
phological or specific sequence. 

The form of the prothorax is not so very different from that ob- 
served in obscura. 

The tibial spurs are broader and larger than is seen in the Dentipes 
group, and resemble those of obscura. In the male the spurs are of 
the same form, but smaller, shorter, and narrower; in the female the 
protibial spurs are more strongly divergent than in dentipes, and less 
so than in obscura. 

It remains for a critical research into the faunal congeneric con- 
stituents of Mexico to further elucidate the relationships of this in- 
teresting species. 

The group genital characters are as follows: 

Male.—Apicale of the edeagophore elongately triangular; surface 
convex and not grooved; sides nearly straight; base with an obtuse 
lobe at middle and feebly sinuate laterally. Sternite slightly trans- 
verse, each lobe broadly rounded at tip, not separated at base by a 
distinctly oval membranous area, and the membrane is not setose 
across the base of the sinus. 

Female.—Genital segment triangulo-trapezoidal. Valvula: Dorsal 
plate oblong, surface broadly concave, more deeply so internally, in- 
ternal border strongly reflexed, externally not at all retracted; apical 
margin truncate. Apex short and triangular. 

Appendage conical and truncate. Superior pudendal membrane 
not reaching beyond the middle of the dorsal plate and longitudinally 
rugulose. Submarginal groove feeble and not visible from above. 

The genital characters show greater affinity with the Dentipes 
group; the external characters show greater resemblance to the Ob- 
scura group. 

It is undoubtedly intermediate. Compare Plate 1, figs. 8, 15, and 
16; Plate 3, figs. 12 and 13, 


i tt ll 


REVISION OF ELEODIINI—BLAISDELL., 207 
ELEODES GRANDICOLLIS Mannerheim. 


Bleodes grandicollis MANNERHEIM, Bull. Soc. Nat. Moscow, XVI, 1843, p. 
266; Mag. Zool., XIV, 1844, No. 130 (fig.).—Horn, Trans. Amer. Phil. 
Soc., XIV, 1870, p. 312. 

Eleodes valida BourMAn, Freg. Eugen. Resa, Ins., 1858, p. 90. 


Large and robust, oblong oval, black, and shining. 

Head twice as wide as long and more or less feebly convex, quite 
coarsely and rather irregularly, sparsely, and at times densely punc- 
tate; punctures denser at the periphery. Antenna rather short and 
stout, reaching to about the posterior fourth of the prothorax and sub- 
equal in the sexes; outer joints slightly compressed and not noticeably 
dilated; third joint equal in length to the next two taken together ; 
the fourth slightly longer than the fifth; the latter, sixth, and seventh 
subequal in length; eighth and ninth about as long as wide; tenth 
slightly wider than long; eleventh smaller, trapezoidal and longer 
than wide. 

Pronotum widest at about the middle, less than one-third wider 
than long, and more than twice as wide as the head; disc smooth 
and shining, quite strongly convex, and rather strongly declivous at 
the angles; surface finely, evenly, and rather sparsely punctate, 
narrowly opaque, and asperate along the lateral margins; apex trun- 
cate or feebly emarginate and rather finely beaded; sides broadly and 
evenly arcuate, somewhat straight, and converging posteriorly, 
scarcely ever sinuate, finely beaded; base slightly arcuate and rather 
coarsely margined; apical angles distinct and deflexed, at times more 
or less rounded; basal angles obtuse and never prominent. 

Propleure smooth, more or less strongly rugulose and at times 
sparsely punctate. 

Elytra oval, usually estriate, smooth, and shining, about twice as 
long as wide, widest at the middle; not emarginate nor abruptly 
declivous at the base, the latter slightly narrower than the con- 
tiguous prothoracic base; humeral angles absent; sides evenly arcu- 
ate, apex obtuse and rounded; disc very convex, suddenly and rather 
obliquely declivous posteriorly ; surface finely and quite evenly punc- 
tate, punctures arranged in moderately distant, unimpressed series, 
the individual punctures not closely placed; intervals with a series 
of distantly and regularly placed punctures, the punctures becoming 
subasperate at the sides, and more strongly so on the apex. 

Epipleure moderately narrow and very gradually and evenly 
narrowing from the base to the apex; surface smooth and sparsely 
punctate. 

Sterna moderately and quite densely punctate and rugulose. 

Parapleure rather strongly and quite evenly punctate. 


208 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Abdomen sparsely and quite evenly punctate, more or less irregu- 
larly and reticulately rugulose; punctures denser om the fifth segment 
and finely setose. 

Legs moderate in length and very stout. Anterior femora in both 
sexes, armed with an acute tooth near the external fourth; tibial 
spurs slightly broadened, those of the protibize equal and acute. Tarsi 
similar in the sexes and rather strongly spinulose. 

Male.—Robust. Ely tra oblong oval and at least twice as long as 
wide, nearly vertically declivous posteriorly. Abdomen moderately 
oblique, not strongly convex; segments one and two broadly and more 
or less strongly impressed. Tibial spurs feebly broadened. 

Female.—Very robust. Elytra broadly oval and less than twice 
as long as wide, vertically declivous posteriorly. Abdomen hori- 
zontal rather strongly convex, and not impressed. ‘Tibial spurs 
noticeably broadened. 

In the material before me two forms can be recognized, notably 
those from central California, and those from southern California, 
Arizona and Lower California. Mannerheim’s type was undoubtedly 
collected somewhere in the vicinity of San Francisco. 

Forma typica—Thorax narrower with the sides not strongly 
rounded, and the apical angles frequently less strongly marked. 
Elytra less broadly oval, so that the general form is less robust and 
narrower. 

Measurements. : Length, 27-29 mm.; width, 10.5-11.5 mm. 
Females: Length, 29 mm.; width, 11.5-12 mm. 

Forma valida.—Thorax ss oader and more strongly rounded at the 
sides; elytra broadly oval, and the general form much more robust. 

Measureme ; Length, 29.-32. mm.; width, 10.5-11.5 mm. 
Females: Length, 32.5-34. mm.; width, 14.-15.5 mm. 

Genital characters, male—Edeagophore comparatively small and 
elongate oval. 

3asale suboval, widest in basal half, evenly convex from side to 
side, slightly arched; dorsum with a rather strong antero-posterior 
convexity and not gibbous in basal half, apically much narrower 
than at middle and the sides rather strongly arcuate. 

Apicale small and rather elongately triangular; surface evenly 
convex, not grooved; sides nearly straight, scarcely sinuate, the apex 
being Leia produced and very narrowly rounded at tip: base with 
the median third obtuse, not rounded nor acute and feebly sinuate 
laterally. 

Sternite slightly transverse. Each lobe rather long, broadly 
rounded at tip, angle scarcely evident; external border nearly 
straight and rather strongly convergent apically; internal margin 
slightly arcuate or straight; surface quite densely punctate in apical 


+! 


REVISION OF ELEODITINI——BLAISDELL, 209 


two-thirds, setose, setw dense at tip and rather long. Membrane not 
setose across the bottom of the narrow sinus. 

Female.—Genital segment triangulo-trapezoidal, rather robust, 
comparatively small and setose. 

Valvula (Plate 1, fig. 8).—Dorsal plate oblong, glabrous; surface 
broadly concave, more deeply so along the strongly reflexed inner 
side, sparsely punctate, each puncture with a long flying seta; exter- 
nal border rather straight, and the internal somewhat sinuous; 
apical margin truncate, angle quite rectangular and set with quite 
long sete. Apex rather short, semi-chitinous and finely setose, with 
a tuft of rather long sete at tip. Fossa large and fringed with rather 
long setee. 

Appendage conical, truncate, moderate in size and setose, with a 
loose pencil of hairs at tip. 

Basal prominences rather strongly developed. 

Superior pudendal membrane not reaching to the middle of the 
dorsal plate. 

Ventrolateral surfaces rather strongly and transversely prominent 
in the basal moiety, broadly and rather deeply concave before the 
apex; apical region punctate and rather thickly set with quite long 
flying sete. Submarginal-groove feeble. Internal margins of the 
valves contiguous in basal fifth. 

Forma typica—//abitat.—Central California (San Francisco, Van 
Dyke; Marin County, Charles Fuchs; Calaveras County, elevation 
2,000 feet, F. E. Blaisdell; Monterey County; Tulare County, Ralph 
Hopping). 

Forma valida.—Southern California (San Diego, Los Angeles, 
Monterey, San Bernardino, and Kern counties); Arizona (Yuma 
Desert, March; Santa Maria River). 

Lower California (San José del Cabo; Lower Paraiso; San Pedro 
Martir). 

Number of specimens studied, 59. 

Location of type unknown to me. 

Ty pe-locality—* California ” (Mannerheim). 

Salient type-characters.—Elongate, convex, and shining. Head 
deeply punctate. Thorax with sides strongly rounded, margin re- 
flexed, finely punctulate. Elytra oblong, moderately narrowed each 
side of the apex, obsoletely striato-punctate. Anterior femora den- 
tate (Mannerheim). 

Diagnostic characters.—The only species in the subgenus with obso- 
lete humeri. One of our largest and finest species, very distinct and 
with obscure relationships. I believe that a study of the Mexican 
species is necessary before much light can be thrown upon the origin 
of not only grandicollis and dentipes, but armata. 

59780—Bull. 683—-09——_14 


210 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


In the series of examples before me the mentum is variable. 
Although large, it is moderate when taken in ag comparative sense, 
varying from trapezoidal to quadrato- or parabolo-trapezoidal; apex 
truncate to evenly arcuate; surface scarcely convex, strongly punctate 
and almost scabrous, feebly foveate at the basal angles. 

The prosternum is variable and strongly protuberant ventrally 
with the coxe, being almost declivous from the anterior margin. It 
is usually longitudinally grooved between the coxee, rounded antero- 
posteriorly, and not mucronate behind, or with an abruptly developed 
mucro; at times convex between the coxz and subtruncate behind, 
with a deflexed mucro. 

The mesosternum is arcuately or obliquely declivous and more or 
less strongly concave. 

The abdominal process is about a fourth of its width wider than 
the metasternal salient, subquadrate, and equal in length to the post- 
coxal portion of the first segment, which is also equal to the third, the 
latter one-sixth of its length shorter than the second; the fourth is 
one-half of its length shorter than the second. 

Metasternum laterally between the cox is about as long as the 
width of a mesotibia at apex. 

Femora stout. The profemora are thickest at the junction of the 
outer and middle thirds, from that point gradually narrowing each 
way. Tibial grooves moderately large, edges rather thick, subcarini- 
form, very feebly arcuate, and converging to become contiguous a 
short distance before the base; floors very feebly concave, subopaque, 
and extremely, finely, and reticulately rugulose. Anterior margin 
dentately laminate at the outer fourth. 

Meso- and metafemora of nearly the same width throughout; sides 
quite parallel. 

Mesofemora with the grooves moderate in width, margins feebly 
cariniform, converging very gradually to become evanescent a short 
distance from the base; floors of the grooves scarcely concave, opaque 
as on the profemora. 

The metafemora have the grooves about moderate in width, with 
the sides feebly cariniform, gradually converging to become evanes- 
cent at basal third; the floors are as on the profemora. 

The protibiz are slightly compressed, carinate in basal half exter- 
nally, with the tarsal grooves more or less feebly developed, asperate 
and limited posteriorly by a muricato-serrulate margin, articular 
cavities closed. 

Mesotibize more or less rounded, briefly or obsoletely carinate exter- 
nally just before the base; tarsal grooves simply indicated by a 
smooth opaque line, or feebly channeled surrounding surfaces muri- 
cate; articular cavities closed. 


REVISION OF ELEODIINI—BLAISDELL. 211 


The metatibiwe rounded, rather small at base and gradually thick- 
ening to apex; tarsal grooves obsolete or feebly indicated, articular 
cavities closed. 

Tarsi rather long and stout. 

The protarsi are about a fifth of their length shorter than a meso- 
tarsus. Joints two, three, and four equal in length, rather short and 
distinctly wider than long, and together equal to the length of the 
fifth; first about as long as wide. 

Mesotarsi just a little shorter than a metatarsus. Joints two, three, 
and four equal in length, scarcely as long as wide, and together about 
equal to the length of the fifth; first joint a little longer than wide. 

The metatarsi are about half as long as their metatibia. Joints 
two and three equal in length and scarcely longer than wide, and 
together about as long as the fourth; first joint about one-and-a- 
half times longer than wide. 


DENTIPES GROUP, Section A. 


The constituents of the present section have the integuments more 
or less opaque, more or less asperately and muricately sculptured ; 
the surface is never normally entirely glabrous nor shining; further- 
more, the species are undoubtedly genetically related by having the 
inferior surface of the cauda, when present, formed by the dilated 
apical extremity of the epipleure. 

A part of the species are ecaudate, the elytral apices being but 
slightly produced, and in the indifferent stage of development. There 
is also a tendency to hirsuteness. With the development of the 
elytral cauda, the femoral teeth become abortive. 

The protibial spurs are acute and quite similar in the sexes, not 
strongly divergent as in the Obscura Group, and instead have a tend- 
ency to become parallel and backwardly curved. 

The mentum is distinctly bearded in a part of the species, but 
here it occurs as a part of a tendency to general pilosity. The head 
is moderate; the articular cavities of the tibie are nearly closed, and 
as a rule the tarsal grooves of the tibie are scabrous. The abdomen 
is horizontal in both sexes, and the intercoxal salient is always dis- 
tinctly wider than the metasternal process. 

The genital characters of this section are not sufficiently character- 
istic to be of practical use when tabulated. 

The male genital characters present nothing more than those enu- 
merated as subgeneric. The most important character is the presence 
of the membranous groove on the apicale. 

In the female the genital segment is triangular or trapezoidal in 
outline, the dorsal plate of the valves is concave and not retracted, 
the submarginal groove is scarcely visible from above, the appendage 
flattened or conical, the superior pudendal membrane does not reach 


219 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


beyond the middle of the dorsal plates, and the genital fissure is open 
and fusiform. ~ 
Our species may be separated as follows: 


Species ecaudate. 


Elytra sulcate, intervals more or less convex________________=_ hispilabris. 

Elytra: substriate,. intervals flatuo3s=sces22 2 eee sponsa, 
Species caudate. 

Dorsal surface of body not clothed with long, flying hairs________ caudifera. 

Dorsal surface clothed with long, flying hairs____________-_____ longipilosa. 


The female genital characters may be tentatively formulated as 
follows: 


Genital segment more or less quadrato-trapezoidal; appendage flattened and 
subtriangular. 
Dorsal plate oblong, sides parallel; apical margin subtruncate_____ sponsa, 
Dorsal plate irregularly oblong; apical margin more or less arcuate. 
I hispilabris. 
Genital segment more or less triangular; appendage conical. 
Dorsal plate oblong, sides subparallel; apical margin more or less oblique. 


caudifera. 
Dorsal plate oblong-oval, sides arcuate; apical margin more or less arcu- 
SGC oF ee ee ee longipilosa. 


Genealogy—An examination of the accompanying diagram will 
elucidate the apparent relationships and lines of descent. The lines 
of divergence are apparently the ac- 
quisition of the elytral cauda on the 
one hand and the elytral apices re- 
maining indifferent on the other; 
in the former the development of an 
unusual degree of pubescence with 
the disappearance of the femoral 
teeth are very striking morpholog- 
ical coincidents along the lines of 
correlated characters. 

Tracing the divergencies along 
Fic. 4.—GENBEALOGICAL DIAGRAM OF THE the ecaudate ramus of the genea- 

DENTIPES Group, SECTION A, SUBGENUS logical tree, it will appear as {F 

eee sponsa diverged, retaining a com- 
paratively simple form of elytral sculpturing and has apparently 
reacted less than hispilabris to its environment. 

Hispilabris has, indeed, undergone a wonderful amount of differ- 
entiation into a number of incipient races, primarily by the sculp- 
turing remaining comparatively simple with rather fragile integu- 
ments, and, secondarily, by a more decided character of sculpturing 
having been acquired along with an increased thickness of the exo- 
skeletal wall. In the former instance forma typica and nupta were 
evolved, the latter by increased ventricosity of the body; in the latter 


REVISION OF ELEODITINI—BLAISDELL. 213 


case sculptilis took on a stronger form of elytral sculpturing, elongata 
an elongate form, and /evis by robustness and a less pronounced form 
of sculpturing. 

This method of intuitively working out and presenting these gene- 
alogical data—possibly more theoretical than real—may at least lay 
a foundation upon which to erect a more substantial knowledge by 
stimulating biological research and criticism. 

Distribution.—The four species under consideration are distributed 
through the following States: 

Texas, Arizona, New Mexico, Oklahoma, Colorado, Kansas, Ne- 
braska, Nevada, northeastern California, Oregon, Washington, Mon- 
tana, and Wyoming. 

Sponsa is southern in habitat, occurring in Texas, Arizona, New 
Mexico, and Colorado. 

Hispilabris has the most extensive distribution, the typical form 
occurring in Texas, New Mexico, Oklahoma, Kansas, Colorado, Ne- 
braska, and Wyoming; forma nupta has thus far been found in Texas 
and Oklahoma; forma convewa in Texas; forma sculptilis in Arizona, 
Colorado, northeastern California, Wyoming, Montana, and south 
into Mexico; forma elongata in Arizona; forma levis in Utah, Ore- 
gon, and Washington. 

Caudifera from Texas, Arizona, New Mexico, and Colorado. 

Longipilosa thus far from the desert regions of Nevada and north- 
eastern California. 


ELEODES SPONSA LeConte. 


Bleodes sponsa LeConte, Proc. Acad. Nat. Sci. Phila., 1858, p. 184.—Horn, 
Trans. Amer. Phil. Soc., XIV, 1870, p. 313. 

Subovate, elongate, black to nigro-piceous. 

Head twice as wide as long, slightly convex, and more or less im- 
pressed laterally and along the frontal suture; swrface more or less 
evenly punctate, punctures moderate, sometimes with small impunc- 
tate areas. Antenne more or less slender and quite long, reaching 
slightly beyond the base of the prothorax, outer four joints slightly 
compressed, feebly and gradually dilated; third joint about equal to 
the next two taken together, fourth slightly longer than the fifth, the 
latter, with sixth and seventh subequal, eighth about as wide as long 
and triangular in outline, ninth and tenth wider than long and trans- 
versely oval, eleventh subovate. 

Pronotum widest scarcely in advance of the middle, about one- 
fourth wider than long, less than twice as wide as the head, and with 
the length equal to the apex; dise smooth and feebly shining, mod- 
erately and quite evenly convex, more or less declivous at the apical 
angles, frequently very narrowly impressed laterally along the mar- 


914 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


gins in anterior half, scarcely opaque, rather finely and more or less 
evenly and sparsely punctate; apex feebly emarginate or subtruncate, 
generally finely margined laterally and obsoletely so at middle; sédes 
evenly and moderately arcuate anteriorly, less so or rather straight 
and convergent from middle to the base, sometimes feebly sinuate 
in front of the basal angles, marginal bead fine; base feebly arcuate, 
finely beaded, and equal to the apex; apical angles acute, prominent 
anteriorly and frequently everted; basal angles subrectangular or 
shghtly obtuse. 

Propleure smooth, obsoletely punctulate and rugulose, or impunc- 
tate. 

Elytra elongate oval, feebly shining to slightly opaque, about twice 
as long as wide, usually widest at the middle; base very feebly emargi- 
nate and equal to or slightly wider than the contiguous prothoracic 
base; humeri obtuse, sometimes slightly prominent; s¢des evenly arcu- 
ate, sometimes narrowing towards base, apex more or less obtuse, 
never produced; disc more or less evenly convex from side to side, 
or somewhat flattened, more or less rapidly and arcuately or obliquely 
declivous posteriorly ; surface feebly substriate, the striz with rather 
coarse, closely placed, and more or less impressed punctures, the inter- 
vals flat or at times slightly convex and with a more or less uneven 
series of submuricate punctures; sometimes the punctures are in- 
dented, both series becoming more strongly muricate at the sides and 
apex. 

Epipleure moderately narrow, with a broad and gentle upward 
curve at base, thence narrowing gradually to apex; surface smooth 
and obsoletely punctulate. 

Sterna usually more or less feebly punctate and rugulose. 

Parapleure more or less strongly punctate. 

Abdomen horizontal, usually smooth and obsoletely punctulate, 
fifth segment rather more strongly punctate. 

Legs moderate in length and rather slender, profemora more or 
less armed in the sexes; protibial spurs quite equal and acute; tarsi 
similar in the sexes. 

Male—Somewhat narrow. Antenne with about three joints 
reaching beyond the prothoracic base. Elytra rather rapidly, arcu- 
ately, and obliquely declivous posteriorly, apex moderately acute. 
Abdomen moderately convex. Profemora with .an acute or obtuse 
tooth at outer fourth. 

Female—Somewhat robust. Antenne with about one joint ex- 
tending beyond the prothoracic base. Elytra arcuately and nearly 
vertically declivous posteriorly ; apex more or less obtuse. Abdomen 
strongly convex. Profemora mutic or with a very obtuse tooth at 
about the outer fourth. 


REVISION OF ELEODITINI—BLAISDELL. 215 


Two forms are to be recognized, namely: 

Forma typica.—Thorax moderate in width, sides not very strongly 
rounded; elytral punctuation more or less muricate, disc striato- 
punctate, interstitial punctures smaller or at most subequal to the 
strial punctures, and usually not at all indented. 

Forma convexa.—Thorax wider, form more convex, punctuation 
not at all muricate, elytral punctures moderately coarse, the strial 
closely placed, the interstitial larger or subequal and more or less in- 
dented, and more or less irregular in some intervals or laterally. 

Measurements—Males: Length, 22-25 mm.; width, 8-10 mm. 
Females: Length, 22.5-26 mm.; width, 9.5-10 mm. 

Genital characters, male—Edeagophore moderate in size and of 
the usual oblong-ovate form. 

Basale oblong and moderate in length. 

Apicale about twice as long as wide at base, triangular and evenly 
convex above, with a feebly impressed linear depression in middle 
third; sides very feebly arcuate from base to apex, the latter nar- 
rowly rounded and obtuse; base broadly and subacutely lobed at the 
middle and sinuate laterally. 

Sternite transverse. Each lobe triangular with the external bor- 
der evenly arcuate from base to apex; internal margin straight; 
apex subrectangular and narrowly rounded; surface sparsely 
punctate and setose in apical two-thirds, setee moderately long and 
not very dense. Membrane not setose across the bottom of the sinus. 

Female.—Genital segment subquadrato-trapezoidal and moderately 
setose. 

Valvula (Plate 4, fig. 8).—Dorsal plate oblong, with the surface 
more or less broadly concave, margins more or less reflexed and sub- 
parallel; finely and sparsely punctate, setose; external margin feebly 
sinuate to slightly arcuate; internal margin arcuate; apical margin 
subtruncate or feebly rounded, with a few short sete. Angle more or 
less rounded. Apex short and triangular, acute at tip and very finely 
setose. 

Appendage short and depressed, wider at base than long, broadly 
rounded at tip, setose, setse rather short forming a loose tuft; external 
angle of the fossa fringed with a few quite long sete. 

Submarginal groove small and sometimes visible from above. 

Superior pudendal membrane reaching to about the middle of the 
dorsal plate and longitudinal rugulose. 

Ventrolateral surfaces——Body transversely inflated and with the 
basal prominences moderate; quite strongly concave laterally and less 
so centrally in front of the apices; quite strongly punctate in apical 
three-fourths, each puncture with a rather short semi-erect hair. 
Genital fissure moderately narrow. 


216 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Habitat—New Mexico (Santa Fé, Hubbard and Schwarz; Cool- 
idge, Coll. E. C. Van Dyke; Aztec, April, Coll. C.F. Baker; High- 
rolls, June 11); Colorado (southern, George Horn) ; Texas (Marfa, 
elevation 4,600-4,800 feet, and at Alpine, July, 4,400-6,000 feet, 
H. F. Wickham); Arizona (M. L. Linell). 

Number of specimens studied, 20. 

Type in the LeConte collection. 

Type-locality—New Mexico; collected by Mr. Fendler. 

Salient type-characters.—Thorax a little convex and somewhat 
rounded, sparsely punctate, anteriorly broadly emarginate, apical 
angles subacuminate; sides rounded; base subrotundate with the 
angles subrectangular. Elytra at apex obtusely acuminate; dise 
punctato-striate, laterally and at apex muricate, intervals sparsely 
uniseriately punctate (LeConte). 

Diagnostic characters.—S ponsa is to be separated from hispilabris 
by the elytra being feebly substriate and with the intervals flat, rarely 
feebly convex and rather coarsely muricately punctate. In some 
specimens the interstitial punctures are in slightly larger dents and 
the sculpturing is not at all muricate but smooth. The thorax is 
usually less rounded on the sides and nearly quadrate, but there is 
considerable variation. 

The elytral sculpturing sometimes resembles specimens of hispil- 
abris which have the intervals scarcely convex. In a complete series 
the transition from the one species to the other is evident. In sponsa 
the elytral dise is usually flattened. LeConte mentioned a specimen 
collected in New Mexico that had the elytra not at all muricate. 

The mentum is variable. In size it may be comparatively large or 
small, varying in outline from parabolo-trapezoidal, trapezoidal to 
trapezoido-triangular, rarely the apex is subacute; surface coarsely 
punctate to almost impunctate, evenly convex or more or less foveate 
laterally. 

The prosternum and mesosternum as in Aéispilabris; the latter is 
sometimes quite vertical and deeply concave at middle. 

The abdominal process is about as wide as the third segment is 
long, the post-coxal part of the first is shorter than the second seg- 
ment, and the latter is twice as long as the fourth; the third is about 
one-fourth of its length shorter than the second. 

The abdominal salient is but slightly wider than the metasternal 
process. 

The metasternum laterally between the coxw is scarcely as long as 
the width of a mesofemur at base. 

The tibize and femora are about the same as in héspilabris. 

The articular cavities of the tibie are very narrowly open as a 
rule or closed. 


———E——eEe——eE ee 


REVISION OF ELEODITNI—BLAISDELL. 217 


The tarsi are moderate, more slender in the smaller individuals 
and stouter in the larger. 

The protarsi are about three-fifths of their length shorter than 
a mesotarsus. Joints two, three, and four are subequal, and just a 
little wider than long, the three combined about equal to the fifth; 
the first is distinctly longer than wide. 

The mesotarsi are about as long as a metatarsus. Joints two, three, 
and four subequal and about as long as wide; together about equal 
to the fifth; the first joint is scarcely twice as long as wide. 

The metatarsi are about one-half as long as a metatibia. Joints 
two and three are subequal and distinctly longer than wide, together 
subequal to the fourth; the first is scarcely as long as the fifth. 


ELEODES HISPILABRIS Say. 


Blaps hispilabris Say, Journ. Acad. Nat. Sci. Phila., I[1, 1825, p. 259; Amer. 
Entomology, I, 1824, p. 32.—LeContr, Complete Writings of Thomas 
Say, I, 1859, p. 32. 

Bleodes hispilabris LeConre, Complete Writings of Thomas Say, LI, 1859, 
p. 152.—Horn, Trans. Amer. Phil. Soc. x1V, 1870, p. 313.—CHAM- 
PION, Biol. Centr.-Amer., IV, Pt. 1, 1884, p. 79. 

Eleodes sulcata LeContr, Proc. Acad. Nat. Sci. Phila., 1852, p. 67; 1858, 
TewkSL. 

Eleodes lecontei GEMMINGER, Coleopt. Hefte, VI, 1870, p. 122 (new name). 

Eleodes connera LeContr, Reports of Explor. and Survey, ... 47th and 
49th parallel, XII, Pt. 38, 1857, p. 49. 

Bleodes nupta LeConte, Proc. Acad. Nat. Sci. Phila., 1858, p. 1838. 

Bleodes compositus CAsry, Annals N. Y. Acad. Sci., VI, Nov., 1891, p. 58 


a 


Elongate ovate, somewhat shining, more or less convex and sulcate ; 
color black, frequently reddish along the suture. 

Head about twice as wide as long, feebly convex, finely and not 
densely punctate, feebly impressed laterally. Antenne long, mod- 
erate in stoutness, rei aching to the prothoracie base, outer four joints 
slightly compressed and noticeably dilated; third joint equal in 
length to the fourth and fifth combined, fourth a little longer than 
the fifth, the latter, sixth, and seventh subequal, eighth equilaterally 
triangular in outline, ninth and tenth transversely oval, eleventh 
irregularly ovate to subtrapezoidal. 

Pronotum widest at about the middle, about one-fifth to less than 
one-third wider than long, and less than twice as wide as the head; 
dise smooth and more or less shining, quite evenly and moderately 
convex, more or less declivous at the apical angles, finely, sparsely, 
and quite evenly punctate, very narrowly opaque and granulate later- 
ally; apex more or less feebly and broadly emarginate, finely beaded 
laterally and obsoletely so at middle; sides broadly and evenly arcuate 
anteriorly, converging and more or less straight posteriorly, or 
slightly sinuate before the basal angles, finely to rather coarsely 


218 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


beaded, and very narrowly or more or less reflexed; base feebly arcu- 
ate and more strongly margined, a little wider than the apex and 
equal to the length; apical angles acute and more or less everted; 
basal angles obtuse. 

Propleure smooth, more or less rugulose and nearly impunctate. 

Elytra oval, sulcate, usually less than twice as long as wide, widest 
at the middle; base very feebly emarginate, usually very little wider 
than the contiguous prothoracic base; humeri obtuse and not promi- 
nent; sides evenly arcuate, apex more or less acute and usually very 
slightly produced ; disc more or less convex, frequently depressed, and 
more or less suddenly obliquely declivous posteriorly ; surface usually 
deeply suleate, sulci deep and about as wide as the intervals, each with 
a series of more or less evenly and closely placed, small asperate 
punctures; the intervals are more or less strongly convex, smooth and 
shining, each with a single irregular series of distantly placed punc- 
tures. The deflexed sides are more coarsely, irregularly punctate and 
subsulcate. 

Epipleure moderate in width, gradually narrowing from base to 
apex, where they may be feebly dilated beneath the elytral apices and 
sightly excavated at that point; surface smooth, frequently im- 
punctate, at other times very sparsely and finely punctulate. 

Sterna moderately punctate and rugulose. 

Parapleure usually quite smooth and sparsely but not deeply 
punctate. 

Abdomen horizontal, smooth, very finely punctulate and rugulose; 
fifth segment more strongly punctate. 

Legs somewhat slender. Anterior femora armed in the sexes; 
protibial spurs subequal, frequently the posterior appears to be just 
the least longer and stouter than the anterior. Tarsi grooved beneath 
and similar in the sexes. 

Male-—Somewhat narrow and elongate. Antenne reaching slightly 
beyond the base of the prothorax. Length of the pronotum slightly 
greater than the width of base and about one-fifth wider than long. 
Elytra arcuately and obliquely declivous posteriorly. Abdomen 
moderately convex from side to side. Anterior femora with a 
moderate and acute tooth at about the outer third. 

Female.——More or less robust, and rather broad. Antenne reach- 
ing to or slightly beyond the prothorax, the latter with the base 
equal to the length and about one-third wider than long. Elytra 
‘rather suddenly and arcuately declivous posteriorly. Abdomen 
strongly convex. Anterior femora with a more or less obtuse tooth 
at about the outer third. 


Genital characters, male-——Edeagophore of the usual oblong-ovate 
form and small. 


REVISION OF ELEODIINI—BLAISDELL. 219 


Basale rather short, not a great deal longer than the apicale, not 
usually arched nor strongly convex. 

Apicale elongately triangular, quite evenly convex above, with a 
linear impression in middle third; sides feebly arcuate from base to 
apex, the latter subacute; base rather acutely lobed at middle and 
scarcely sinuate laterally. 

Sternite transverse. Lobes rather small; each with the external 
border straight and converging from base to apex, the latter rounded, 
angle introrse and scarcely developed; internal margin straight; 
surface punctate and moderately thickly setose, seta moderate in 
length and not extending upon the membrane across the bottom of the 
sinus, the latter short but rather broad. 

Female—Genital segment quadrato-trapezoidal, width at base 
about equal to the length, surface rather conspicuously setose. 

Valvula (Plate 3, fig. 19).—Dorsal plate more or less irregularly 
oblong, quite deeply concave, with margins reflexed; surface very 
finely and sparsely punctate, setee rather long and flying; external 
margin feebly sinuate to slightly arcuate; internal border rather 
broadly arcuate, sometimes subangulate near base; apical margin 
evenly rounded and set with quite long flying hairs, angle rounded. 
Apex short and triangular, acute at tip, finely setose; external angle 
of the fossa quite thickly set with rather long hairs. Lateral plate 
visible from above external to the dorsal plate, with submarginal 
groove between the two and more or less visible. 

Appendage depressed, more or less triangular and rounded at tip, 
not longer than the apical lobe; surface set with quite long sete. 

Superior pudendal membrane reaching to about the middle of the 
dorsal plate, and longitudinally rugulose. 

Ventrolateral surfaces.—Body rather strongly convex at base, not 
inflated, glabrous, more or less strongly and transversely concave 
before the apices; apical moiety sparsely punctate, with sete rather 
long. Internal margins of the valves contiguous at base for a short 
distance. Genital fissure rather wide and fusiform, closed in basal 
half by the inferior pudendal membrane. 

Salient type-characters—More or less shining. Elytra frequently 
more or less depressed, and somewhat narrowly rounded laterally, 
frequently more or less reddish along the suture; sulci moderate in 
depth, intervals more or less convex. The apical angles of the pro- 
notum are acute and more less everted. Say’s description is too brief 
as to the essential characters. 

The following incipient races or forms should be recognized : 

Forma typica.—See salient type-characters. 

Measurements —Males: Length, 20-25.5 mm.; width, 8.5-10.5 mm. 
Females: Length, 18-28 mm.; width, 8.8-12 mm. 


220 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Number of specimens studied, 150. 

Type destroyed. ~ 

Type-locality—* Inhabits Missouri,” a badly defined geographical 
region in Say’s day. 

Forma nupta LeConte.—Elytra with the dorsum slightly and the 
sides strongly convex (LeConte). The elytra are more or less evenly 
convex from one epipleura to the other, usually somewhat inflated, 
sometimes quite strongly so. Some of the smaller examples are quite 
eychriform. 

Measurements.—Males: Length, 21-27 mm.; width, 9.5-11.2 mm. 
Females: Length, 19-29 mm.; width, 9.8-14 mm. 

Number of specimens studied, 15. 

Type in the LeConte collection. 

Type-locality—Laredo to Ringgold Barracks, Texas. 

Forma sculptilis—Integuments dull and rather thick. Head 
and thorax more or less shining. Elytra more or less convex, sulci 
usually opaque and deep, with the intervals strongly convex, shining 
and conspicuous, quite abruptly limiting the grooves. 

Measurements.—Males: Length, 19-23 mm.; width, 7.8-8.5 mm. 
Females: Length, 19-24 mm.; width, 8-10 mm. 

Number of specimens studied, 40. 

Forma elongata.—General form distinctly elongate and narrower 
as compared to the other forms; black. Elytra are sulecate, but less 
strongly so than in sculptilis; the intervals at times are feebly convex. 
The prothorax is noticeably larger and the legs stout. 

Measurements.—Males: Length, 26.5 mm.; width, 9 mm. Females: 
Length, 29 mm.; width, 10.2 mm. 

Number of specimens studied, 6. 

Forma convexa LeConte.—Elongate, black and bright. Thorax 
moderately punctulate, sides rounded, anterior angles acute and 
slightly prominent; basal angles obtuse. Elytra elongate oval, in- 
tervals subconvex and rugose ; subacute posteriorly. 

Measurements—Length, .9; width of elytra, .32; thorax, .23 (Le- 
Conte). ; 

Specimens not at hand. 

Type is in the LeConte collection. 

Ty pe-locality—Prairie Paso, Texas; collector, Dr. Cooper. 

Forma levis.—Robust, integuments decidedly black, thick and 
alutaceous, also quite smooth. Thorax rather large. Elytra usually 
evenly convex from side to side, sulci very shallow, intervals very 
feebly convex, subobsoletely sculptured. Legs somewhat slender. 

Measurements.—M ales: Length, 23 mm.; width,8 mm. Females: 
Length, 23-26 mm.; width, 9.2-10 mm. 

Number of specimens studied, 8. 


es 


REVISION OF ELEODIINI—BLAISDELL. 921 


The following species described by Colonel Casey evidently belongs 
here: ; 

Forma composita Casey.—Moderately robust, convex, intense black 
throughout, the pronotum dull and strongly alutaceous; elytra shin- 
ing, very feebly alutaceous, glabrous. 

Head densely, finely, rather sparsely punctate and dull; antenne 
rather long and stout, fully as long as the head and thorax, third 
joint between three and four times as long as wide, outer joints 
slightly broader, the ninth and tenth somewhat transversely oval. 

Pronotum about as long as wide; disc evenly convex, very minutely, 
sparsely punctate; apex broadly, very feebly emarginate; s¢des paral- 
lel, broadly arcuate or subangulate just before the middle, very feebly 
sinuate just behind the apical angles, feebly convergent and nearly 
straight to the base; base arcuate and equal to the apex in width; 
apical angles right, slightly blunt, not prominent; basal angles 
broadly obtuse. 

Elytra about two and two-thirds times as long as the prothorax, 
and, in the middle, one-half wider than the latter, about twice as long 
as wide; humeri not at all exposed; sides broadly arcuate, apex 
acutely angulate; disc strongly declivous behind, deeply sulcate, the 
ridges strongly convex and shining, each with a single rather uneven 
series of small, moderately distant punctures, the intervals as wide as 
the ridges, deep, opaque, each with a single series of rather small, 
close-set, asperate punctures. 

Legs slender, the femora shining, rather finely but strongly punc- 
tate, the anterior alone with a strong acute tooth near the apex; 
tibiz strongly rugose except toward base, the spurs of the anterior 
small, acute, and nearly equal. Prosternum with the apex slightly 
reflexed, acute and tuberculiform. 

The above is Casey’s original description. 

Measurements of the type.—J/ale: Length, 16 mm.; width, 6.6 mm. 

Unknown to me. 

Type: a unique in Colonel Casey’s collection. 

Type-locality.—Texas. 

Forma typica.—//abitat—Missouri (Say). Colorado (August, 
C. V. Riley; Denver, April and June, Coll. H. Soltau; May, Colls. 
Dyar and Caudell; Colorado Springs, June, elevation 6,000-7,000 
feet, H. F. Wickham. In his List of the Coleoptera of Colorado, Pro- 
fessor Wickham gives the following localities, the forms are not differ- 
entiated: La Junta, West Las Animas, Canyon City, San Luis Valley, 
Fort Garland, Trinidad, Holly, Fort Collins). Kansas (collections 
of Chas. Fuchs, C. V. Riley, F. H. Snow, and F. E. Blaisdell). New 
Mexico (Las Vegas, August, Colls. Barber and Schwarz; Albuquerque, 


Coll. E. C. Van Dyke). Texas (Mobeetee, October, Coll. H. S. Bar- 


222 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


ber.) Oklahoma (Fort Supply, Coll. H. S. Barber.) Wyoming 
(Cheyenne, June, Colls. Hubbard and Schwarz), Nebraska (Sand 
Hills, September, Coll. C. V. Riley). 

Forma nupta.—Oklahoma (Fort Supply, Coll. H. S. Barker; Texas 
(Alice, E. A. Schwarz; Fort Creek, E. A. Mearns; San Diego, April 
and May, Hubbard and Schwarz; Goliad County, October, J. D. 
Mitchell). 

Forma sculptilis—Arizona (Walnut, July, H. F. Wickham; Oracle, 
July, Coll. Hubbard and Schwarz; Williams, May, Coll. Barber and 
Schwarz; Fort Grant, July, Coll. Hubbard and Schwarz; Ash Fork, 
June; Yuma, Chas. Fuchs). New Mexico (Las Vegas, August, Coll. 
Barber and Schwarz). Colorado (Denver, May, Coil. Dyar and 
Caudell). Utah (Salt Lake City, April, Coll. H. Soltau). Cah- 
fornia (northeastern, Coll. Hubbard and Schwarz). Wyoming 
(Cheyenne, May, H. Soltau; June, Hubbard and Schwarz). Mon- 
tana (Helena, May, Henry Ulke, Hubbard and Schwarz). Mexico 
(Cananea, district of Arizpe, State of Sonora, elevation 4,500-6,000 
feet, Dr. W. C. Alvarez). 

Forma elongata.—Arizona (Coll. Beverly Letcher, Prescott). 

Forma convexa.—Texas (Prairie Paso, Doctor Cooper). 

Forma levis.—Utah (Salt Lake, June, Coll. Hubbard and 
Schwarz). State of Washington (Walla Walla, Charles Fuchs). 

Diagnostic characters—Hispilabris 1s recognized from the mem- 
bers of the Obscura Group by the prominent dentiform apical angles 
of the prothorax and by the simple tarsi; from all species of the 
subgenus //eodes by the more or less deeply sulcate elytra with con- 
vex intervals, and in never being caudate. 

I consider the typical form to have more or less depressed elytra, 
the latter with the intervals feebly to moderately convex and fre- 
quently reddish along the suture; the integuments are generally thin- 
ner than in the other forms. 

The five incipient races which I have recognized are simply ex- 
tremes along particular lines of divergence in the ample series before 
me, and they are all united into one continuous series by an abundance 
of mesotypes. 

They ought to be readily recognized by the brief diagnoses that I 
have given. Their salient characteristics may be epitomized as 
follows: 

Nupta with its strongly convex and more or less ventricose elytra. 
Integuments as in typical hispilabris. 

Sculptilis with the elytral intervals very strongly and prominently 
convex, with the sulci deep and opaque. Denser integuments. 

Elongata by its elongate and moderately convex form, elytral 
intervals more or less moderately convex. Black and denser integu- 
ments. 


REVISION OF ELEODIINI—BLAISDELL. 923 


ww 
Conyewva by its elongate form. Elytra with subconvex and rugose 


intervals. 

Levis by its robust convex form; integuments black and aluta- 
ceous; large prothorax and somewhat eroded sculpturing. 

Composita Casey has evidently the elytral sculpturing of forma 
seulptilis and the general structure of Aispilabris, except that it has 
an aberrant prothorax. But here the difference is no proof that it is 
specifically distinct. Analogous aberrations are observed in dentipes 
in particular, and hispilabris is fully as variable as that species. 

The unique described by me as interrupta in the subgenus Melanc- 
leodes was simply an example of symmetrical fortuitous variation of 
sculpturing, and was an anomalous oméssa. I must hold that the 
present form, which is a unique and undoubtedly related to a very 
variable species, is an aberration, and the collection of others like it 
must give it a greater claim to more than an incipient race. 

The mentum is moderate in size and quite constant in form, and 
more or less trapezoidal; the sides are quite straight and converging 
to the apex, the latter distinctly truncate, or slightly arcuate; the 
sides are not noticeably curved. In fact, I have not yet seen an ex- 
ample with the middle lobe evenly rounded anteriorly. 

The prosternum is variable. Usually moderate in width behind the 
equator of the coxie, evenly convex, or precipitously so behind; fre- 
quently submucronate, vertically truncate, and feebly compressed 
behind in nupta. 

Mesosternum variable; oblique to arcuately declivous and more or 
less concave. 

The metasternum laterally between the coxe is about equal in 
length to the width of a mesofemur at base. 

The abdominal process is subquadrate, slightly transverse, and in 
width about equal to the length of the post-coxal portion of the same 
segment; second segment slightly longer than the post-coxal part of 
the first and about twice as long as the fourth; the third is about 
one-third of its length shorter than the second. 

The abdominal salient is about one-fourth of its width wider than 
the metasternal process. 

It is to be observed that the elytral apices have a tendency to be 
slightly produced, and especially in forma sculptilis, and that the 
epipleure dilate at tip to form the inferior surface of this feeble but 
evidently produced elytral apex. This character becomes of impor- 
tance when taken in relation with the caudal characters of caudifera 
and longipilosa. 

The tibial grooves of the femora are not usually quite entire and 
rather narrow. Those of the profemora may have their margins 
feeble or strongly defined; the floors are glabrous and rather flat or 


224 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


feebly concave, The margins are scarcely arcuate, and converge a 
short distance from the femoral base; the anterior margin is dentately 
laminate at about the junction of the outer fifth and sixth sevenths. 

The grooves of the meso- and metafemora are more or less defined 
by sharp margins, which converge to become contiguous somewhere 
near the inner third, less frequently near the middle. 

The tibiz are variable. Protibiw more or less feebly compressed, 
obsoletely to distinctly carinate externally; the external edge scarcely 
arcuate. The tarsal grooves are sometimes feebly marked and 
asperate, or at other times quite well defined and more or less gla- 
brous. The articular cavities are scarcely open, or feebly so at the 
most. 

The articular cavities of the meso- and metatibiz are usually quite 
closed. 

The tarsal grooves of the mesotibizw are sometimes defined for 
nearly the whole length of the external surface, floor of each quite 
flat and more or less glabrous; usually the grooves are poorly defined 
and asperate. 

The external surface of each metatibia is more or less flattened and 
the groove generally feebly defined. 

The tarsi vary in stoutness according to the general habitus of the 
individual. 

In some examples of the typical form the tarsi appear compara- 
tively longer than in the more robust forma levis. In the typical 
form a protibia is 6 mm. long and its tarsus 3.5 mm. in length; in 
forma levis the protibia is 6 mm. and the tarsus 3 mm. in length. 
The actual lengths of parts are of no value, but only when taken 
relatively. 

It appears as if the shortening is done at the expense of the length 
alone. In the typical form joints two, three, and four are evidently 
longer than wide; in forma levis the same joints are distinctly wider 
than long. In each instance they are subequal to each other, and 
combined equal to the length of the fifth. The first joint is about 
one-half longer than wide. 

In the typical form the protarsi are three-sevenths of their length 
shorter than a mesotarsus and in the forma Jwvis one-third shorter. 

The mesotarsi in the typical form are about one-ninth of their 
length shorter than a metatarsus—to the unaided eye they appear 
subequal. Joints two, three, and four are longer than wide and 
slightly decreasing in length in the order named; together they are a 
little longer than the fifth; first joint about equal in length to the 
second and third taken together. 

In forma levis the meso- and metatarsi are subequal, and the meso- 
tarsi have joints two to four, inclusive, quite equal. 


REVISION OF ELEODIINI—BLAISDELL, 225 


Forma. typica has a metatarsus half as long as a metatibia; in 
forma levis it is distinctly less than one-half as long. In the former 
type form joints two and three are subequal in length and about one 
and one-half times longer than wide, and together equal to the fourth ; 
the first joint is a little shorter than the fourth. In forma levis the 
relative lengths are practically the same. 

It appears that with each slight variation in size the relatiye pro- 
portions between the tarsi and their joints also correspondingly vary, 
and at the same time the essential specific characters were homoge- 
neous in the examples tested. 

Of course, these variations must be within normal specific limits. 
These facts become taxonomic pabulum for those students who recog- 
nize a new species every time certain structural elements present 
slight differences in their relative proportions. There are times, of 
course, when these characters are of great value, but specific instabil- 
ity must determine their value in a given case, and normal specific 
variations must be recognized. 


ELEODES CAUDIFERA LeConte. 


Bleodes caudifera LeConrr, Proc. Acad. Nat. Sci. Phila., 1858, p. 184.— 
Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 314. 


Elongate, ovate, caudate. 

Head about twice as wide as long, feebly convex and more or less 
impressed laterally along the frontal suture; surface irregularly and 
rather finely punctate, sparsely so with impunctate areas at center, 
and densely punctate laterally. Antennw moderate, scarcely com- 
pressed, last four joints very slightly and gradually dilated, third 
joint very slightly longer than the fourth and fifth combined; fourth 
to the seventh, inclusive, subequal; eighth slightly shorter and 
broader; tenth and eleventh about as wide as long, the latter ovate. 

Pronotum widest in front of the middle, about one-third wider 
than long and about twice as wide as the head; disc rather shining, 
quite smooth, moderately convex, and slightly declivous at apex and 
basal angles, more or less impressed and opaque laterally along the 
margins in the anterior two-thirds, finely and very irregularly punc- 
tate with impunctate areas at middle; laterally the opaque area is 
more or less transversely rugulose and granulate; apex feebly 
emarginate or truncate, very finely or obsoletely margined; sides 
evenly and quite strongly arcuate in anterior one-half, quite straight 
or feebly sinuate and convergent from middle to base, marginal bead 
fine or obsolete; base slightly arcuate and very finely margined, 
wider than apex; apical angles rounded, obtuse or minutely denti- 
form and everted; basal angles obtuse, 


59780—Bull. 683—09——15 


226 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Propleure smooth and opaque, rugulose, with few scattered 
granules, especially upon the acetabular convexitigs. 

Elytra oval, shining, and more or less suleate, less than twice as 
long as wide—apical prolongation not included—widest at middle 
and truncate at base, where they are slightly wider than the contigu- 
ous prothoracic base; Awmeri obtuse and more or less rounded; 
sides evenly arcuate, apex horizontally produced; dise moderately 
convex, slightly flattened and arcuately rounded laterally, obliquely 
declivous posteriorly passing into the cauda; surface striate, strie 
more or less impressed and strongly punctate, the punctures muricate 
and closely placed, intervals more or less convex but not strongly so, 
uniseriately punctate, the punctures distantly placed and muricate, 
those of the outer intervals more or less confused and at times rather 
densely and muricately subrugoso-punctate; inflexed sides obsoletely 
striate and quite densely and irregularly muricately punctate and 
at times slightly concave at base of apex. 

Cauda moderately long, granulo-muricate, above with the suture 
impressed, tip emarginate; beneath concave, sparsely muricato- 
granulate, the surface of each lateral half being a dilatation of the 
apical epipleural surface of the corresponding side; in the median 
line the two inferior epipleural margins are contiguous. 

Epipleure rather narrow and gradually narrowing from base to 
apex, where they dilate to become the inferior surface of the cauda; 
surface opaque, smooth, and sparsely muricately punctulate, except 
at base. 

Sterna more or less irregularly, submuricately punctate and 
rugulose. 

Parapleure more or less irregularly punctate. 

Abdomen horizontal, rather finely, sparsely, and evenly submuri- 
cately punctate. 

Legs moderately long and more or less slender. Femora rather 
thickly clothed with moderately long appressed setae, with an occa- 
sional long flying hair on the external surface. Tibiz densely muri- 
cate and rather sparsely set with long flying hairs; anterior spurs 
quite equal in length and size and more or less backwardly curved. 
Tarsi similar in the sexes and clothed with rather long hair-like 
setee, those on the last joint of each tarsus projecting noticeably be- 
yond the ungues. 

Male—Somewhat narrow. Antenne reaching slightly beyond the 
prothoracic base. Elytra rather gradually and obliquely declivous 
posteriorly, becoming sinuate in passing into the horizontal cauda; 
the latter may be equal to about one-sixth of the entire elytral length. 
Abdomen feebly ventricose to the fourth segment. Anterior femora 
with a broad and very obtuse tooth or mutic. 


REVISION OF ELEODIINI—BLAISDELL. 227 


Female.—Rather robust. Antenne reaching scarcely to the protho- 
racic base. Elytra somewhat suddenly obliquely declivous posteriorly 
and sinuate at base of apex. Cauda equal to about one-eleventh of 
the entire elytral length. Abdomen quite strongly ventricose to the 
fourth segment. Anterior femora quite mutic. 

Measurements without and with the cauda—Males: Length, 21 
and 24; 25 and 29.5 mm.; width, 8-10 mm. Females: Length, 22 and 
94; 26.5 and 29.5 mm.; width, 9.5-12 mm. 

Genital characters, male-—Edeagophore of the usual oblong-ovate 
form. 

Basale short and oblong, subequal in length to the apicale. 

Apicale elongate and suboblong-triangular, moderately convex and 
slightly flattened, with a linear membranous groove in middle third; 
sides parallel in basal half, thence arcuate, becoming slightly sinuate 
at apex, the latter acute and slightly produced; base arcuate, scarcely 
sinuate laterally. 

Sternite subquadrate. Each lobe about twice as long as wide, 
broadly rounded at apex; external margin nearly straight; internal 
margin rather arcuate; surface glabrous, not very thickly punctate 
and setose, setee moderately long. 

Female——Genital segment triangular, subtruncate, length about 
equal to the width at base. 

Valvula (Plate 4, fig. 7)—Dorsal plate oblong with the surface 
broadly concave, margins reflexed and nearly parallel; surface finely 
punctured in apical two-thirds, each puncture bearing a short semi- 
recumbent seta; external border nearly straight; internal margin 
slightly sinuous: apical margin more or less oblique, asperate, set 
with moderately long and evenly distributed sete, which are not 
longer at the angle, the latter more or less rounded and never promi- 
nent. Apex membrano-chitinous, scarcely longer than the append- 
age, acute; surface set with fine sete, apical tuft rather short. 

Appendage chitinous, conical, quite long and nearly as long as 
the apex, blunt, and asperate in apical half, set with rather long sete, 
tuft at tip quite long. 

Superior pudendal membrane scarcely reaching one-third from the 
base. 

Ventrolateral surfaces —Body moderately convex but not notice- 
ably inflated, slightly and transversely concave before the apices, 
finely punctate and set with very fine, short sete in apical two-thirds. 
Submarginal groove obsolete. External margin of the dorsal plate 
not prominent laterally. Genital fissure long and fusiform, with the 
inferior pudendal membrane visible in basal half. 

Habitat—New Mexico (Aztec, April, C. F. Baker; Coolidge, Coll. 
E, ©. Van Dyke and Hubbard and Schwarz; Santa Fé, C. V. Riley; 


228 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Luna, Hubbard and Schwarz) ; Arizona (Adamana, May, H. S. Bar- 
ber; Winslow, July, Coll. Barber and Schwarz; Hglbrook, Coll. E. C. 
Van Dyke, Charles Fuchs, and Beverly Letcher); Colorado (Henry 
Ulke; Eastern, George Horn); Texas (El Paso, March, Coll. H. 
.Soltau). 

Number of specimens studied, 40. 

Type in the LeConte collection. 

Type-locality—New Mexico: collector, Mr. Fendler. 

Salient type-characters.—Thorax scarcely convex, slightly punc- 
tate, laterally a little concave and transversely rugose; anteriorly 
broadly emarginate, apical angles not acuminate; sides strongly 
rounded, narrowed posteriorly; posterior angles obtuse. Elytra_ 
slightly flattened, substriate with seriate punctures, intervals uni- 
seriately and sparsely punctate, at the sides and apex muricate, pos- 
teriorly obliquely declivous and prolonged. Anterior femora mutic 
(female) (LeConte). 

The following forms are before me: 

Forma glabra.—More or less shining and glabrous, not at all muri- 
‘at, except on the cauda. Elytra striate, strize impressed, with quite 
coarse and closely placed punctures; intervals moderately convex, 
with a single series of small, distantly placed punctures. Legs quite 
slender. Cauda gradually narrowing from base to apex. 

Measurements—Male: Length, 25 mm.; with cauda, 31 mm.; 
width, 9.5 mm. Female: Length, 20 mm.; with cauda, 22.5 mm.; 
width, 9 mm. 

Habitat.—Texas (El Paso), Arizona. 

Forma sublevis.—Subopaque to feebly shining. Antenn slender 
and rather long. Elytra with series of unimpressed punctures: in- 
tervals flat with a series of distantly spaced punctures, which are as 
large as those of the striz. Sculpturing submuricate. Legs rather 
slender. 

Measurements —Female: Length, 25 and 28 mm.; width, 10 mm. 

Habitat—Arizona. 

Forma typica.—See salient type-characters. 

Forma scabra.—More or less shining. Elytra with unimpressed 
strie of very closely placed and minute muricate tubercles; intervals 
with a single series of distantly placed and larger muricate tubercles, 
which become distinctly larger laterally and more tuberculiform. In- 
flexed sides of the elytra rugoso-muricate. The general surface is 
very minutely rugulose. Size of forma typica. Legs rather short. 

Habitat—New Mexico (Aztec). 

Diagnostic characters—Both sexes are caudate. Thorax widest 
just in advance of the middle; body not notieeably pubescent, tibiz 
and tarsi with long flying hairs. These characters serve to differen- 
tiate caudifera from sponsa, hispilabris, and longipilosa, 


REVISION OF ELEODITNI—BLAISDELL. 229 


Doctor Horn in the Revision failed to mention that the female was 
caudate. 

‘Forma glabra is a rather striking variation, on account of the 
smooth integuments and in having the cauda longer than usual in the 
male, and moderately arcuately decurved in its apical third, being 
gradually narrowed from base to apex. 

Forma sublevis has the sculpturing of sponsa, and forma scabra 
has the elytra tuberculo-scabrous. The mentum is comparatively 
rather small and triangulo-trapezoidal in outline. The sides con- 
verge anteriorly and are scarcely at all arcuate; the apex is more or 
less rounded and the surface is more or less foveate as well as coarsely 
punctured. 

The mentum and sides of the epistoma are set with rather long 
black hairs. 

The prosternum is variable, moderately wide between the cox 
and somewhat triangularly dilated behind the equator of the coxx; 
more or less convex antero-posteriorly, or feebly convex to subhori- 
zontal between the acetabula; more or less moderately produced and 
compressed, oblique to vertically truncate behind, at times sub- 
mucronate. 

The mesosternum is variable as to the degree of obliquity and 
more or less concave. 

The abdominal process is subquadrate, slightly wider than long 
and about as wide as the post-coxal portion is long, the latter 
equal in length to the second segment; the third is twice as long as 
the fourth, and about one-third of its length shorter than the second. 

The abdominal salient is about one-seventh of its width wider than 
the metasternal process. 

The metasternum laterally between the cox is about as long as 
a mesofemur is wide at base. 

The tibial grooves of the femora are not entire, the floors are 
glabrous, quite plane, and limited by rather well defined edges. 
Those of the profemora are moderate in width; the anterior margin 
is scarcely dentately laminate at the outer third in the male and less 
so in the female; the posterior margin is obsolete internally before 
becoming contiguous with the anterior, neither one reaching the 
femoral base. 

The mesofemoral grooves are obsolete at the internal third of the 
femora and the margins do not become contiguous; on the meta- 
femora they scarcely extend basally beyond the middle, where they 
become evanescent before becoming contiguous. 

The protibie are somewhat compressed and distinctly carinate 
externally; the tarsal grooves are scarcely developed and muricate. 


230 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


The mesotibize are feebly compressed and the external surface is 
ridge-like, and the tarsal grooves are evident, although more or less 
feeble and muricate. 

The metatibiz are flattened externally, obsoletely grooved, an 
muricate. Both the meso- and metatibiz are more or less carinate 
in basal third externally. 

The tarsi are quite stout. 

The protarsi are scarcely one-third of their length shorter than a 
mesotarsus. Joints two, three, and four are subequal, just a little 
wider than long, and together about equal to the fifth; the first is a 
little longer than wide. 

The mesotarsi are about one-seventh of their length shorter than a 
metatarsus. Joints two, three, and four are subequal, a little longer 
than wide, and together just the least longer than the fifth; the first 
is about one-half longer than wide. 

The metatarsi are scarcely one-half as long as a metatibia. Joints 
two and three are subequal, distinctly longer than wide, and together 
scarcely as long as the fourth; the latter is a little longer than the 
first. 

ELEODES LONGIPILOSA Horn. 


Eleodes longipilosa Horn, Trans. Amer. Ent. Soc., XVIII, Feb., 1891, p. 42, 
DL i, fis. 13: 


Elongate oval, moderately shining, surface sparsely clothed with 
long, black, flying hairs; caudate. 

Head twice as wide as long, more or less moderately convex, 
coarsely and more or less irregularly punctate, hairs long and flying. 
Antenne moderate in length and thickness, scarcely compressed or 
dilated in outer four joints, third joint equal to the next two taken 
together, fourth to the eighth inclusive subequal, seventh conico- 
triangular, eighth triangulo-quadrate, ninth and tenth suborbicular, 
eleventh ovate and pointed; clothed with rather long hairs. 

Pronotum widest at or slightly in advance of the middle, about one- 
fourth wider than long, and twice as wide as the head; dise mod- 
erately convex, coarsely, irregularly, and more or less sparsely punc- 
tate, rather narrowly. opaque and granulate along the sides, fre- 
quently with impunctate areas at center; apex slightly emarginate, 
finely or more or less obsoletely margined; sides arcuate in front 
(occasionally subangulate) and obliquely converging posteriorly, 
marginal bead fine or obsolete and replaced by minute granules; base 
more or less arcuate, finely margined, one-tenth to one-fourth wider 
than apex and about equal to the length; apical angles small, sub- 
acute, not everted or subdentiform and feebly everted; basal angles 
obtuse and scarcely rounded. 

Propleure sparsely muricate and rugulose. ’ 


. 


REVISION OF ELEODIINI—BLAISDELL. 231 


Elytra elongate oval, widest at middle and more attenuated pos- 
teriorly; base truncate or feebly emarginate, slightly wider than the 
contiguous prothoracic base; Awmeri obtuse and more or less rounded ; 
sides evenly arcuate and slightly sinuate at apex, the latter pro- 
duced, forming a moderately long, horizontal, or slightly down- 
wardly oblique cauda; disc feebly convex on the dorsum, rather 
strongly rounded laterally, and somewhat gradually, arcuately de- 
clivous posteriorly ; surface more or less striato-punctate, very vagtely 
suleate, punctures near the suture scarcely muricate, more strongly 
so laterally and on the apex, serial punctures of the striz moderate 
in size and closely placed, the interstitial series similar and more dis- 
tantly spaced, both series becoming irregular laterally and on the 
apex; suture impressed on the cauda, the latter emarginate at tip; 
each puncture with a long flying hair. 

Epipleure moderate in width and gradually narrowing from base 
to apex, where they dilate to form the inferior surface of the cauda; 
surface opaque and obsoletely punctulate. 

Sterna more or less punctate and rugose, sparsely clothed with 
long brownish hairs. 

Parapleure coarsely punctate and not noticeably pubescent. 

Abdomen horizontal and more or less ventricose in both sexes, quite 
closely punctate, most densely so on the first and fifth segments. 

Legs moderate in length and thickness, densely sculptured and 
clothed with long, flying, sparsely distributed hairs. Anterior 
femora mutic in both sexes; anterior tibial spurs acute, subequal, 
moderate in length, and more or less curved. Tarsi similar in the 
sexes. 

Male.—Somewhat narrower than the female. Antenne reaching 
to the prothoracic base. Elytra slightly narrowly oval. Abdomen 
very feebly flattened on first two segments. 

Female—Rather robust. Antenne scarcely attaining the pro- 
thoracic base. Elytra quite broadly oval. Abdomen strongly 
convex. 

Measurements without and with the cauda—Males: Length, 19 
and 21.5; 19.5 and 22.5 mm.; width, 7.2 mm. Females: Length, 19 
and 21; 20 and 22.2 mm.; width, 8.2 mm. 

Genital characters, male——Edeagophore of the usual oblong-ovate 
form, rather elongate, and scarcely arched. 

Basale rather short oblong, evenly convex; sides feebly arcuate. 

Apicale elongate, nearly as long as the basale, and triangular; 
surface quite evenly and moderately convex, with a narrow median 
and more or less obsolete groove in basal two-fifths; sides moderately 
arcuate in basal half, thence more or less sinuate to apex, the latter 
produced and acute, not deflexed; base broadly rounded at middle, 
and more or less sinuate laterally. 


232 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Sternite quadrato-parabolic and slightly transverse. Lobes slightly 
elongate, each with the external border feebly singate in basal half, 
thence evenly and rather broadly arcuate to apex, the latter rounded ; 
internal border membrano-chitinous and not well defined in basal 
half; surface more or less convex, from the middle gradually sloping 
to the sinus, rather coarsely punctate and setose; sete rather long at 
apex and moderately dense; membrane not setose across the bottom 
of the sinus; the latter broad. 

Female.—Genital segment somewhat equilaterally triangular, 
slightly flattened, and setose. 

Valvula (Plate 4, fig. 23).—Dorsal plate oblong oval; sides more or 
less reflexed; surface more or less concave, smooth, and shining- 
finely and sparsely punctate, each puncture with a fine and rather 
short. seta ; external margin more or less feebly arcuate; apical margin 
feebly arcuate, angle scarcely evident; internal margin more or iess 
arcuate, feebly sinuate in basal half; apex short, triangular, subacute 
at tip, and finely setose. Valvular membrane visible in apical half. 

Appendage about as long as the apex, conical, with a loose tuft of 
fine and not very long hairs at tip. 

Superior pudendal membrane finely and not closely longitudinally 
rugulose, not quite attaining the middle of the dorsal plate. 

Ventrolateral surfaces —Body evenly but not strongly convex, and 
not transversely concave before the apex, shining and glabrous, 
finely, sparsely punctate and setose; submarginal groove obsolete; 
internal margins of the valves contiguous in basal fifth. Genital 
fissure long, fusiform, and rather wide, closed by the inferior puden- 
dal membrane in basal half, apically by the valvular membranes. 

Habitat—Nevada (Humboldt Valley, Mr. Krause), California 
(Amedee, Lassen County, July, elevation 4,200 feet., H. F. Wickham). 

Number of specimens studied, 16. 

Type in the Horn collection. 

Type-locality—Humboldt Valley, Nevada; collector, Mr. Krause. 

Salient type-characters.—Clothed with long, black, flying hairs. 
Thorax about a fourth wider than long; sides arcuate in front. 
obliquely narrowed posteriorly; disc moderately convex, coarsely, 
sparsely, irregularly punctate; apex slightly emarginate, anterior 
angles subacute, but not everted. Elytra with disc feebly convex, 
very vaguely sulcate; surface irregularly, sparsely muricate, the two 
grooves nearest the suture coarsely punctate. Elytra prolonged at 
tip in a cauda. (Male.) (Horn.) 

Diagnostic characters —Remarkably distinct among its congeners 
on account of being clothed throughout with long black or brownish 
flying hairs; caudate in both sexes with the anterior femora obso- 
letely dentate. 


Ee ar TE EN 


REVISION OF ELEODITNI—BLAISDELL, 233 


The mentum is quite trapezoidal in the females, and also rather 
broad; triangulo-trapezoidal and smaller in the males. The sides 
are more or less straight, and converge to apex, the latter arcuate or 
truncate; surface more or less strongly punctate, scarcely foveate, and 
set with long hairs. 

The prosternum is somewhat variable, usually feebly convex and 
longitudinally grooved between the coxe, compressed and vertically 
truncate behind, with the angle more or less prominent and sub- 
mucronate; sometimes oblique and feebly mucronate. 

Mesosternum scarcely to strongly concave, and more or less arcu- 
ately oblique. 

The abdominal process is about equal in width to the length of the 
post-coxal part of the same segment, and subquadrate. The second 
segment is longer than the post-coxal portion of the first; the third 
is half of its length shorter than the second, and about twice as 
long as the fourth. 

The abdominal salient is a little wider than the metasternal inter- 
coxal process. 

The metasternum laterally between the cox is as long as the width 


of a metafemur at base. 
The tibial grooves of the femora are not entire, their margins are 


distinct and granular, their floors flat and very sparsely granulate 
at the sides. 

On the profemora the sides of the grooves become contiguous a 
short distance from the femoral base; on the meso- and metafemora 
they are somewhat contiguous near the middle, thence to base the 
femora are rounded beneath. 

On the profemora the anterior margins of the grooves are not 
visibly laminate. 

The protibiz are quite distinctly carinate externally; the meso- and 
metatibizw are also in basal fourth externally; tarsal grooves on the 
protibie evident, but feebly and coarsely sculptured; the grooves are 
less evident on the mesotibiz and densely sculptured ; metatibize quite 
circular in transverse section and not grooved, coarsely sculptured. 
The protibix are slightly compressed, with the articulac cavities more 
or less feebly open; on the meso- and metatibiz the cavities are closed. 

The tarsi are moderate in length and rather stout, with joints rather 
closely articulated. 

The protarsi are about a fifth of their length shorter than a meso- 
tarsus. Joints two, three, and four rather short and subequal, dis- 
tinctly wider than long, and together about equal in length to the 
fifth ; the first about as long as wide. 

The mesotarsi are about equal in length to a metatarsus; joints 
two, three, and four about as long as wide and subequal, together 


234 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


about equal to the length of the fifth; first joint just a little longer 
than wide. ; 

The metatarsi are about one-half as long as a metatibia; joints 
two and three subequal in length, a little longer than wide, and 
together shorter than the fourth; the first about a half longer than 
wide. 

DENTIPES GROUP, Section B. 


The present section contains those species that have the integu- 
ments glabrous and more or less shining, and never distinctly as- 
perately sculptured. The caudal characters, as already defined, show, 
without doubt, a genetic relationship not heretofore recognized. As 
in Section A some of the species are ecaudate, but unlike those in that 
section, there is here no tendency to hirsuteness among its constituent 
species; on the other hand there is a tendency to complete non-devel- 
opment of such epidermal structures; spines and spinules are de- 
veloped here as in the species of the preceding Section. 

The anterior tibial spurs are similar in the sexes, acute and mod- 
erately divergent, at times feebly curved backward. 

The abdomen is horizontal and the intercoxal salient of the first 
segment is always wider than the metasternal process. 

The genital characters are the same in this section as in the pre- 
ceding one. 

The species may be tabulated as follows: 


All of the femora armed. 
Femoral teeth strongly developed, long and acute; elytra more or less 


verticglly declivous behind: =—- 2.2 = ee eee armata. 
Femoral teeth rudimentary but distinct on the mesofemora and obsolete 
on ‘the: metafemora- 2-2 2.2222 =e ee var. impotens. 


Femoral teeth quite equilaterally triangular, shorter, broader, and less 
acute; elytra evenly and arcuately declivous behind and more attenuate. 
militaris. 
Femoral teeth smaller and obtuse, otherwise as in militaris__var. femorata, 
Anterior femora alone armed. 
Thorax strongly constricted at base. 
Thorax strongly transverse and very strongly rounded at the sides. 

Dise with sides narrowly impressed and granulate, more or less evenly 
and moderately convex; elytral apex more or less attenuately pro- 
duced or 'caudate: $244.28 eee a eee eee acuticauda, 

Disc with sides broadly impressed and granulate, less convex; punc- 
tuation finely muricate; elytra more or less attenuate and not 
CAhUGEtC. 2a ee ae ee ee eee var. laticollis. 

Thorax moderately strongly and evenly rounded at the sides; elytra not 
attenuate behind with apex obtusely rounded, never acute nor pro- 
GUCEGd: 22232 ee Se ee ts Se ee dentipes. 

Thorax feebly constricted at base. 

Thorax transverse; elytra striato-punctate, series quite closely placed 
and impressed, intervals more or less convex____-~------ subpinguis. 


REVISION OF ELEODIINI—BLAISDELL. 235 


Thorax subquadrate, sides not strongly rounded; elytra smooth, not at 
all muricately punctate. 

Divine) -eipiey water closely placed... eee gracilis. 
bog TUR a EC ch en ee ee 9 iP 

Thorax subcylindrical, sides evenly and feebly arcuate throughout; 
elytra with unimpressed distant rows of nearly simple punctures. 

subcylindrica. 
Thorax not constricted at base; males more or less caudate. 

Elytra strongly ventricose; apical angles of prothorax moderately 
prominent anteriorly; first joint of the protarsi (male) clothed at 
apex beneath with a transverse tuft of golden modified spinules. 

ventricosa, 

Elytra feebly ventricose; apical angles of prothorax larger, more prom- 
inent anteriorly, acute and more or less reflexed; first joint of ante- 
rior tarsi without transverse tuft of spinules_____________ var. falli. 

Elytra rarely ever ventricose and distinctly striato-punctate. 

Apical angles of prothorax feeble and not prominent anteriorly. 
eschscholtzii. 
Apical angles usually well developed, acute and anteriorly prom- 


PUTT gcd eel ar TUE an See a Ee Be tel a RP ee var. luce. 
Elytra not distinctly striato-punctate and elongate oval. 

Anpieal- angles of prothorax everted_=_.___ 2... -=...--~—_ tenuipes. 

Aen aoe lem NOU CVCrbCU os- 26 oe eee ee wickhami. 


The student must bear in mind that in all probability he will find 
the above table inadequate as an aid in the determination of all 
specimens. If the examples be more or less typical there should be 
no trouble, but among species so polymorphous, puzzling forms will 
constantly be found. 

In the text I have given attention to the most marked incipient 
races as well as to uniques and aberrations when at hand. is 

Genealogy.—To speculate upon lines of descent of the species con- 
stituting the present section involves great doubt as to the correctness 
of the assumptions, but may be a useful working hypothesis. Never- 
theless I must present my convictions; for there is no doubt in my 
mind but that the ancestral stock had the femora strongly armed as 
in armata, and that this species is a direct and somewhat modified 
modern exponent of that stock. 

A typical armata does not have the sides of the prothorax sinuate 
or constricted before the base. Now, I have before me examples of 
armata which have the sides of the prothorax as strongly sinuate as 
in any dentipes, and all of the femora are well armed. 

I advance the theory that all of the species now constituting the 
present section of the Dentipes Group in our fauna—which includes 
Lower California as well as all of that territory north of Mexico— 
have had their origin from the ancestral armata stock. It may be 
asked when did this divergence take place? I would answer, first, 
that which took place in early times, and, second, that which has 
taken place in more recent times. 


236 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


To the first class belong those forms which have had origin from 
the primitive eschscholtzii, gracilis, and acuticaud@ rami. 

The accompanying genealogical diagram will elucidate my meaning. 

The more recent derivatives of the eschscholtzii ramus are our pres- 
ent eschscholtzii, luca, and tenuipes-wickhami,; of the gracilis ramus, 
our present gracilis, distans, and subcylindrica (*%); of the acuti- 
cauda ramus, the present acuticauda, and laticollis. Around each of 
these more recent forms are the present incipient species and races 
which render our species so polymorphous. 


Armata 
Impotens 


Femorata 


Eschscholtzi 


Acuticauda_ » 


Ancestralfl Trunk 


Fic. 5.—GENEALOGICAL DIAGRAM OF THE DENTIPES GROUP, SECTION B, SUBGENUS ELEODES. 


The recent derivatives of armata, are militarts and femorata by 
the modification of general form and slow atrophy of the femoral 
teeth; there are edentate forms derived from these species at the 
present time in Lower California, and two of them are before me. 

An example of this loss of the femoral teeth is our émpotens. I 
have studied very instructive series of armata from Fresno and 
Merced counties. These series show all intermediate stages between 
a typical armata and a typical dentipes. 

These remarks are sufficient to indicate my ideas on this subject. 

Distribution —The species of Section B are found inhabiting six 
States or regions: Mexico, Texas, New Mexico, Arizona, California, 
and Lower California. 


REVISION OF ELEODITNI—BLAISDELL. 


237 


Table of Distribution. 


: ) : 
Arizona. .| New Mex- | Texas..| Mexico .|........-. 


| errr ee ; 
1co, 
etre tee Se On eno atin twa waatal> oa5 cuaiee California 
; 
subcylindrica .......| Arizona. 7 PS eet odes te tie ake dk baa scle «we | 
SE Peres Pre wesaos i ab te | Cees See | 
8 eS, ee rotation ta sicenktecelrens scat California | 
2 | tad ; 
ee a LAG) tt 3m Fete geile eas tae RO er California | 
Ste eS <a, <a California) 
militaris ..........- MPa ik. a Mc ee ce cin dt ase dlpee eee ae Lower Cal- 
ifornia. 
ON ee ES ee eee a a. See Southern 
Califor- 
nia. 
EE SES ae ee De ete Bk ee Bs eee | California 
: F | ; “ 
SEE SBS ECE re! ARS REUSE Ce eee ee mee ee California 
eschscholtzii ....---- 1 Ee See Texas. -| ee eet EEE 
RIE rere get oe rte ke Pe Soe Ce Le ee so elwalbeue cect LowerCal- 
ifornia. 
ON ida Sool e enh henlv anda das = UNM ia Gon anieties sh ome 
mcmnamt ....---.-< a Lg: og een Cee Oe as 
ico. 
i | ae ee epee ore Texas .:)Mexiceod <1... f... 
os Os SS ee ee Texas. .| 


ELEODES GRACILIS LeConte. 


Bleodes gracilis LEContTr, Proc. Acad. Nat. Sci. Phila., 1858, p. 184.—Horn, 
Trans. Amer. Phil. Soc., XIV, 1870, p. 312.—CuHAmpPpion, Biol. Centr. 
Amer., IV, Pt. 1, 1884, p. 79. 


Elongate oval, subfusiform, black, sometimes slightly piceous, 
smooth, shining and moderately convex, never caudate. 

Head twice as wide as long, slightly convex, more or less feebly 
impressed laterally, sometimes along the frontal suture and less often 
transversely so before the vertex; surface finely, irregularly, and not 
densely punctate. Antenne rather long, reaching a little beyond the 
prothoracie base, slightly incrassate and feebly compressed in outer 
four joints, third joint subequal to the next two combined, fourth 
longer than the fifth, the latter, sixth and seventh subequal, eighth 
triangular, ninth and tenth transversely oval, eleventh ovate. 

Pronotum widest at or slightly in, front of the middle, about one- 
fifth wider than long and more or less subquadrate ; disc more or less 
moderately convex, finely and more or less sparsely punctulate, 
scarcely granulate along the sides; apea subtruncate or feebly emar- 
ginate, very finely and more or less obsoletely margined; sides very 
moderately and evenly arcuate in anterior two-thirds, rather more 
feebly arcuate, straight or subsinuate and converging in basal third, 


238 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


marginal head fine; base more or less arcuate and finely margined, 
slightly wider than the apex and equal to the length ; apical angles 
acutely dentate or acuminate and more or less everted; basal angles 
obtuse, not rounded nor prominent. 

Propleure more or less convex, obsoletely punctulate and more or 
less rugulose. 

Elytra oval, rather elongate, widest at the middle, smooth, more or 
less shining and not in the least muricate; base feebly emarginate and 
scarcely wider than the contiguous prothoracic base; humeri obtuse 
not rounded nor prominent; sides evenly arcuate, apex subacute and 
not produced; disc rather feebly convex on the dorsum, strongly and 
evenly so laterally, more or less gradually arcuately declivous poste- 
riorly; surface striato-punctate, punctures fine and not impressed, 
those of the striz sometimes larger, closely placed, the intervals flat, 
with a series of more or less smaller and quite distantly placed punc- 
tures; the punctures become but slightly irregular at the sides and on 
the apex. 

Epipleure moderate in width beneath the humeri and gradually 
narrowing to apex; surface smooth and obsoletely punctate. 

Sterna usually not strongly punctate and more or less irregularly 
rugose. 

Parapleure sparsely punctate. 

Abdomen horizontal, very sparsely, finely, and obsoletely punctu- 
late, more or less finely and obsoletely rugulose; most distinctly 
punctate on the fifth segment. 

Legs moderate in length, rather slender but variable. Anterior 
femora armed in the sexes, with a moderate more or less acute tooth. 
Anterior tibial spurs equal and acute in the sexes. Tarsi similar. 

Male—Elongate and rather slender. Elytra rather gradually 
narrowed and arcuately declivous posteriorly. Abdomen rather 
moderately convex, somewhat flattened on the first two segments and 
very feebly impressed at middle. 

Female.—Elongate and slightly robust. Elytra rather suddenly 
and arcuately declivous posteriorly. Abdomen evenly and rather 
strongly convex. 

Measurements —Males: Length, 21-26 mm.; width, 6.5-9 mm. 
Females: Length, 20-27.5 mm.; width, 8-10.8 mm. 

Genital characters, male—KEdeagophore oblong-ovate, somewhat 
flattened and not arched. 

Basale about twice as long as wide, evenly convex, sides evenly and 
feebly arcuate. 

Apicale long triangular, evenly convex above, with a moderate 
median groove in apical third; sides evenly and slightly arcuate from 
base to apex, the latter subacute; base rather broadly lobed at middle 
and feebly sinuate laterally. 


REVISION OF ELEODITNI—BLAISDELL. 239 


Sternite transverse. Each lobe with the external border quite 
evenly arcuate, or rather straight in basal half, and quite broadly 
rounded at apical border, angle introrse and narrowly rounded; in- 
ternal border more or less sinuate; surface feebly convex, rather 
densely punctate in apical two-thirds and setose, sete quite long in 
apical third. Sinus rather broad with membrane not setose across 
the bottom. The lobes are separated cephalad to the sinus by a rather 
broad oval membranous area, which is not as sharply defined @s in 
the Obscura group. 

Female.—Genital segment trapezoidal in outline and finely setose. 

Valvula (Plate 4, fig. 19).—Dorsal plate oblong, the internal third 
in apical half strongly reflexed and glabrous; surface longitudinally 
concave, shining, finely and not densely punctate, setose, sete fine; 
outwardly the surface is obliquely reflexed; external margin rather 
feebly and broadly sinuate; internal border arcuate in apical half, 
thence to base sinuate; apical margin not reflexed and more or less 
evenly arcuate and not easily differentiated from the base of the 
appendage, angle rounded. Apex triangular and moderate, clothed 
with very fine inconspicuous sete, which are not longer at tip. 

Appendage flattened, as broad at base as the apical margin of the 
dorsal plate, rather irregularly semioval, shorter than the apex, 
finely setose, sete short and soft; margins of the fossa thin and 
closely applied to base of the appendage. 

Superior pudendal membrane attaining the middle of the dorsal 
plate, finely and longitudinally rugulose. 

Basal prominences rather small. 

Ventrolateral surfaces—Body of the genital segment with the sur- 
face evenly convex but not noticeably inflated in basal half, quite 
strongly and transversely concave before the apices: surface shining, 
finely punctate and setose, sete fine. Internal margins of the valves 
contiguous in basal fourth. Genital fissure long and evenly fusiform, 
inferior pudendal membrane visible in basal half. 

Habitat—New Mexico (LeConte and Horn; Warren Knaus; 
H. C. Fall; collection of Hubbard and Schwarz); Arizona (H. K. 
Morrison; Walnut, July, M. L. Linell; Santa Rita Mountains, June: 
Sulphur Springs, May; Fort Grant, July; San Simon, July: Chi- 
ricahua Mountains, May, Hubbard and Schwarz; collections Chas. 
Fuchs and C. V. Riley) ; Texas (Alpine, June, elevation 4,400-4,600 
feet, H. F. Wickham; Fort Bliss, collection H. C. Fall); Mexico 
(State of Sonora, Morrison). 

Number of specimens studied, 40. 

Type in the LeConte collection. 

Type-locality—Arizona (or Sonora, type bears a silver dise indic- 
ative as given) ; collectors, Webb and Schott, 


240 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Salient type-characters—HElongate. Thorax rotundato-quadrate, 
moderately convex; disc finely and_ sparsely punctulate ; sides 
rounded, posteriorly subsinuate; posterior angles obtuse, anterior 
angles acute and more or less acuminate. Elytra with dorsum 
slightly convex, more strongly so laterally, scarcely sulcate, punctato- 
striate, intervals with very few punctures. (LeConte.) 

Diag gnostic characters—Known by its elongate form and smooth 
surface. Elytra not at all muricate; pronotum subquadrate, with 
sides not strongly rounded. The elytra are not at all caudate. 

The prothorax is much narrower than in dentipes. It is with more 
difficulty separated from sponsa, and especially forma convexa (see 
p.215). Here its general form, glabrous integuments, and finer punc- 
tuation must be depended upon. The character of the sculpturing 
is the most important. Occasionally rather strongly punctate ex- 
amples are met with, but the sculpturing is simple; if muricate it 
is to be referred to sponsa. Sponsa forma convexa is smooth, and 
here the interstitial punctures are usually large and somewhat in- 
dented. 

Var. distans is usually more robust and has thicker integuments, 
and is found in a different geographical region; the elytral inter- 
vals are wider and the punctures are very distinctly defined. 

For other specific differentiations consult the diagnostic characters 
of Group A of the present subgenus. 

I can not see that gracilis eats any close affinity or resemblance to 
luce as suggested by Doctor Horn. 

The specimens collected in Lower California and referred to this 
species by Dr. George Horn® and in the collection of the California 
Academy of Sciences, do not belong to this species at all, but to a 
distinct species described in this paper as ¢ibzalis. 

I doubt that true gracilis occurs in Lower California. All data 
thus far collected are surely negative. It does occur in the State of 
Sonora, Mexico, and might pecsibly extend westward upon the 
peninsula. 

The mentum is moderate. The middle lobe is rather short and 
somewhat equilaterally triangular, with apex rather narrowly 
rounded; at times reflexed at tip so as to appear truncate; surface 
coarsely punctate, rather broadly and feebly convex centrally and 
more or less distinctly foveate laterally, not noticeably setose. 

The prosternum is variable; convex between and behind the coxe, 
sometimes feebly mucronate. 

The mesosternum is more or less arcuately precipitous or oblique, 
and more or less concave. 


“See Proc. Calif. Acad. Sci., 2d ser., IV, p. 349, 


REVISION OF ELEODITNI—BLAISDELL. 241 


The abdominal process is subquadrate, just noticeably transverse, 
and in width about equal to the length of the post-coxal portion of 
the same segment, and also to that of the third; the second segment 
is about a fourth of its length longer than the third, the latter is 
slightly less than twice as long as the fourth. 

The abdominal salient is about a fifth wider than the metasternal 
process. ; 

The metasternum laterally between the coxe is as long as the width 
of a mesofemur at base. 

The tibial grooves of the profemora are rather broad apically; the 
sides, which are distinct, converge regularly and rather rapidly to 
become contiguous a short distance from the femoral base; the floors 
are nearly flat and quite glabrous, each anterior margin is more or 
less acutely and dentately laminate at the outer fourth in both sexes. 

The grooves of the meso- and metafemora are rather narrow; sides 
rather well defined and asperate, becoming obsoletely contiguous at 
about basal third; the floors are more or less opaque, glabrous or 
rugulose and subasperate. 

The protibiw are more or less distinctly carinate externally, and 
slightly compressed; the tarsal grooves are not very well defined and 
usually scabrous, with the posterior margins more or less serrulate; 
the articular cavities are more or less slightly open. 

The meso- and metatibiw are not at all carinate externally in the 
series before me, and the articular cavities are quite closed; grooves 
of the mesotibizw are more or less defined, and at times moderately 
deep, extending to about the middle of the tibiz, and their floors are 
more or less glabrous or scabrous. 

The external surfaces of the metatibize are flattened, more or less 
scabrous, and the grooves may be distinct. 

The tarsi are moderate and variable in length and stoutness. 

The protarsi are about one-sixth of their length shorter than a 
mesotarsus. Joints two, three, and four subequal, quite short and 
wider than long, and together equal to the fifth; the first is distinctly 
longer than wide. 

The mesotarsi are about a seventh of their length shorter than a 
metatarsus. Joints two, three, and four distinctly longer than wide 
and slightly shortening in length in the order named, and together 
a little longer than the fifth; the second and third combined about 
equal in length to the first. 

The metatarsi are about half as long as a metatibia. Joints two 
and three are about one and a half times longer than wide and not 
equal, together equal in length to the fourth, the first being very 
slightly shorter. 

59780—Bull, 683—09——16 


242 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 
ELEODES GRACILIS var. DISTANS, new variety. 


Elongate ovate or oval, slightly robust, integuments firm, glabrous 
and shining; elytra with distant strie of very sharply defined punc- 
tures. 

Head about twice as wide as long, feebly and quite evenly convex, 
broadly and slightly impressed laterally, frontal suture not generally 
evident, finely, rather sparsely and somewhat evenly punctate, slightly 
more densely so laterally. Antenne rather long, attaining a short 
distance beyond the prothoracic base, feebly compressed and a little 
incrassate in outer four joints, third joint about equal to the next two 
taken together, the fourth a little longer than the fifth, the latter to 
the eighth inclusive subequal in length, eighth about as wide as long 
and triangular, ninth and tenth feebly transversely oval, eleventh 
short and subtrapezoidal or ovate. 

Pronotum widest at about the middle and about one-fifth wider 
than long; disc moderately and quite evenly convex, more or less 
declivous at the apical angles, polished and shining, rarely subalu- 
taceous, very finely-and sparsely punctulate, laterally in anterior 
four-fifths narrowly and feebly impressed, impressed area subopaque 
and more or less obsoletely granulato-rugulose; apex feebly and 
broadly emarginate, rather coarsely beaded laterally, obsoletely so at 
the middle; sides broadly, evenly, and not strongly arcuate in ante- 
rior three-fourths, thence to base less arcuate, straight or scarcely 
subsinuate, rather coarsely beaded; base broadly and slightly areu- 
ate, coarsely beaded, about one-fourth wider than the apex and equal 
to the length; apical angles acute, not large, subdentiform and fre- 
quently slightly everted; basal angles obtuse, distinct, and not 
rounded. 

Propleure smooth and opaque, very sparsely and obsoletely punc- 
tulate, more or less rugulose. 

Elytra oval, widest at the middle, polished and shining; base 
broadly, feebly, and distinctly emarginate, very slightly wider than 
the contiguous prothoracic base; Awmeri obtuse, distinct, and not 
prominent; sédes evenly arcuate, somewhat gradually narrowed pos- 
teriorly, apex obtusely rounded; disc more or less moderately convex, 
rarely slightly flattened on the dorsum, evenly and broadly rounded 
laterally, arcuately but not suddenly declivous behind; surface 
striato-punctate, strie distant and not impressed, regular and usually 
very distinct, punctures distinctly defined, those of the striz close 
and separated by a distance equal to their diameters, moderately small, 
intervals flat with a series of very fine and distantly spaced minute 
punctules; punctures simple, but somewhat feebly submuricate on 
apex, scarcely confused laterally. 


REVISION OF ELEODIINI—BLAISDELL. 243 


_Epipleure moderate in width beneath the humeri, thence grad- 
ually narrowing to apex; surface smooth, subopaque, and usually 
impunctate. . 

Sterna not strongly punctate and more or less rugulose. 

Parapleure sparsely and obsoletely punctate. 

Abdomen horizontal, obsoletely punctate on first three segments, 
more strongly so on the fourth and fifth segments. 

Legs moderate in length, slender to somewhat robust; anterior 
femora armed with an acute tooth in both sexes; tibial spurs similar 
in the sexes, the anterior subequal and acute. The tarsi are simple 
and similar in the sexes. 

Male—Elongate and scarcely robust. Elytra somewhat gradually 
narrowing posteriorly, arcuately and slightly obliquely declivous. 
Abdomen moderately convex, slightly and broadly impressed on 
first two segments. 

Female.—Elongate and rather robust. Elytra not noticeably nar- 
rowed behind and arcuately declivous, abdomen somewhat strongly 
and evenly convex. 

Measurements.—Males: Length, 19-26 mm.; width, 8-9 mm. /e- 
males: Length, 20-28.5 mm.; width, 8-10.8 mm. 

Genital characters, male—As in gracilis; except that the median 
groove of the apicale extends to the basal fourth and becomes wider 
and more membranous. 

Female (Plate 5, fig. 19)—Genital segment trapezoidal, moder- 
ately robust and sparsely setose. 

Valvula—aApparently oblique at the apex. Dorsal plate oblong, 
concave, inner fourth reflexed and glabrous, external three-fourths 
obliquely and gradually reflexed; surface glabrous and_ shining, 
sparsely punctate and setose, setee moderately short and fine; external 
border nearly straight; internal margin arcuate in apical three- 
fourths thence sinuate to base; apical margin arcuate and at the 
internal fourth continuous with the surface of the apex, angle not 
evident, each set with a few rather long sete. Apex triangular, with 
its external border giving the valve an obliquely truncate appearance 
as it passes into the inferior margin of the fossa, chitino-membranous, 
setose, setee mostly fine, a few longer on tip. 

Appendage semi-oval, slightly longer than wide, quite evenly 
rounded, flattened and transverse, finely setose, with a very loose 
tuft of longer setw at apex. Fossa transverse between the apical 
margin of the dorsal plate and external border of the apex. 

Basal prominences not large but distinct. 

Superior pudendal membrane reaching to about the middle of the 
dorsal plate and somewhat distantly longitudinally rugulose. 


244 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Ventrolateral surfaces—Body not very stout, moderately convex 
and very slightly swollen, broadly and feebly concave before the 
apices; surface smooth and sparsely punctate in apical two-thirds, 
setose, setee fine and small; submarginal groove small and distinct 
between external margin of the dorsal plate and the thin external 
border of the lateral plate, terminating at the external angle of the 
fossa. Internal margins of the valves contiguous in basal fourth. 

Genital fissure in middle two-fourths and fusiform, closed in basal 
half by the inferior pudendal membrane. 

Habitat—California (Fort Tejon, April 19-21, Fuchs and Hop- 
ping; Los Angeles County, Dr. E. C. Van Dyke; Fairmount, April 
23, Fuchs and Hopping; Norwalk, December 25, Antelope Valley, 
August 5, H. C. Fall; Oak Creek, Kern County; Tehachapi Valley, 
June 15). 

Number of specimens studied, 60. 

Types in my own collection (males and females). 

Type-locality —F airmount, Los Angeles County, California. Col- 
lectors, Fuchs and Hopping. 

Salient type-characters.——Somewhat robust and elongate, integu- 
ments firm, smooth and polished. Pronotum scarcely subquadrate, 
sides broadly arcuate in» anterior two-thirds, thence quite straight 
and converging to base; apical angles acuminate and dentiform, 
slightly everted; basal angles obtuse and distinct; dise very sparsely 
and minutely punctulate. Elytra with distant and distinct unim- 
pressed strive of rather small punctures, intervals flat with a series 
of very minute and distantly spaced punctules. 

Diagnostic characters.—In collections distans is usually associated 
with gracilis and is to be differentiated like that species. It is more 
robust than gracilis; the intense black and firm body wall, highly 
polished and shining surface are quite characteristic. Elytral striz 
are more distinct than in gracilis and the punctuation is more dis- 
tinctly defined. 

A very interesting fact is that the most heterotypical form—a 
large robust example with pronotal dise subalutaceous, is amphi- 
typical of hispilabris forma levis, but the punctuation is perfectly 
simple and typical of the present race; the other heterotypes of the 
series resemble gracilis in form, but the sculpturing still remains char- 
acteristic of distans. 

The interesting series at hand leads me to the following specula- 
tion: It appears that distans is the smooth modification of hispilabris 
forma levis, the latter being more northern and the former more 
southern in habitat, distans passing into the typical form of gracilis, 
from which it can not be sharply defined; in fact it may be the trans- 
itional form uniting gracilis and hispilabris—if there is a break in 


REVISION OF ELEODITNI—BLAISDELL. 245 


the chain it is between forma levis and distans, for the latter is truly 
more closely united to graci/is than to the former. 

General observations —The mentum is nearly as in gracilis. It 
is larger and the middle lobe is less foveate. 

The prosternum is prominent ventrally with the coxe, convex be- 
tween the same and vertically truncate behind, the angle a mucro; 
sometimes simply with a small mucro and not truncate. 

Mesosternum usually rather vertically declivous and broadly con- 
cave. 

The abdominal process is subquadrate, slightly transverse, about 
equal in width to the length of the post-coxal portion of the same 
segment; the second segment is slightly more than twice as long as 
the fourth; the third about a third of its length shorter than the 
second, 

The abdominal salient is about a fourth of its width wider than 
the metasternal process. 

The metasternum laterally between the coxe is about equal in 
length to the width of a mesofemur at base. 

The tibial grooves of the profemora are rather broad apically, the 
sides are rather strong and finely scabrous, becoming contiguous a 
short distance from the base; their floors are more or less concave and 
smooth. On the mesofemora the grooves are moderate in width; the 
sides converge but become obsolete before becoming contiguous, one- 
fourth from the base. The grooves of the metafemora are narrower, 
margins rather strong and obsolete at basal third, before becoming 
contiguous. 

The protibiz are quite distinctly carinate externally and slightly 
arcuate. In the males the base is somewhat constricted. The tarsal 
grooves behind the carine are narrow and scabrous; the articular 
cavities are more or less open. 

The articular cavities of the meso- and metatibizw are narrowly 
open or closed. The grooves of the mesotibiw are usually strongly 
developed, concave with the floors, smooth and opaque, margins muri- 
cately scabfous, and they extend to the basal third. The metatibie 
are flattened externally and also grooved in apical three-fourths, and 
the grooves are more or less smooth or asperate at bottom, 

The tarsi are rather stout. The protarsi are about two-fifths of 
their length shorter than a mesotarsus. Joints two, three, and four 
short, distinctly wider than long and subequal, together slightly 
shorter than the fifth; the first about as long as wide. 

The mesotarsi are about a seventh of their length shorter than a 
metatarsus. Joints two, three and four subequal and about as long 
as wide, together about as long as the fifth; the first is a little shorter 
than the next two taken together. 


246 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


The metatarsi are distinctly less than a half as long as a metatibia. 
Joints two and three subequal, a little longer than wide and together 
about equal to the first and not quite as long as the fourth. 


ELEODES SUBCYLINDRICA Casey. 
- Bleodes subcylindrica Casry, Annals N. Y. Acad. Sci., V, Nov. 1890, p. 400. 


Form cylindrically convex, rather slender, finely, very strongly 
alutaceous and smooth throughout, black; elytra castaneous. 

Head moderate, sparsely and rather finely punctate. Antenne 
rather long and slender, about as long as the head and prothorax, 
third joint nearly four times as long as wide. 

Prothorax subcylindrical, about one-fifth wider than long; disc 
transversely, moderately but very evenly convex throughout, longi- 
tudinally very feebly convex, very sparsely and. extremely minutely 
punctate throughout; apex very nearly as wide as the base, subtrun- 
cate between the apical angles; sides evenly and feebly arcuate 
throughout; base broadly and very feebly arcuate; apical angles 
acute, very strongly advanced but not at all everted; basal angles 
obtuse, not in the least rounded, and not at all prominent. 

Elytra more than three times as long as the prothorax and rather 
less than one-third wider; base very feebly emarginate, exactly equal 
in width to the contiguous base of the pronotum; humeri very ob- 
tuse, not rounded, not at all prominent; sides feebly convergent and 
arcuate near the humeri, parallel and almost perfectly straight thence 
to the posterior third; apea narrowed rather gradually in apical 
third, acutely rounded at tip; disc with unimpressed distant rows of 
very small, nearly simple punctures, the intervals with single rows of 
extremely widely distant but almost similar punctures, the punctua- 
tion not denser towards the sides. 

Legs long and very slender; anterior femora with a small but acute 
tooth slightly beyond apical third, the remaining femora simple, but 
with the groove of the lower edge fine, very deep and conspicuous, 
extending almost to the base, with the cariniform edges finely, un- 
evenly serrate; tibie strongly arcuate, spurs of the anterior short, 
nearly equal and slender. 

Measurements.—Length, 21-23 mm.; width, 7.5-7.8 mm. 

Habitat—Arizona (collected by Mr. G. W. Dunn). 

Species unknown to me. Number of specimens studied by Colonel 
Casey, 2. 

Type in Colonel Casey’s collection. 

T ype-locality —Arizona (exact place not given). 

Colonel Casey considers this species totally distinct in general 
habitus from any other which is known to inhabit the United States, 


a a 


ea oe 


REVISION OF ELEODITINI—BLAISDELL, 247 


and not described in the Biologia Centrali-Americana. He advises 
that it be placed near gracilis. I have been unable to recognize it in 
the large amount of material which has passed through my hands. 


ELEODES SUBPINGUIS, new species. 


Elongate oval to ovate, rather robust, subopaque to feebly shining, 
somewhat ventricose and not muricately punctured; elytra striato- 
punctate, intervals feebly convex, more strongly so on apex. 

Head less than twice as wide as long, feebly convex, broadly 
impressed laterally and along the frontal suture, sometimes trans- 
versely so before the vertex, somewhat sparsely and irregularly 
punctate at middle and rather densely so laterally, punctures 
moderate. Antenne long, distinctly reaching beyond the prothoracic 
base in both sexes, slightly compressed and incrassate in outer four 
joints, third joint about equal in length to the next two combined, 
fourth just slightly longer than the fifth, the latter with sixth and 
seventh subequal, eighth triangular and about as long as wide, ninth 
and tenth slightly transversely oval, eleventh trapezoidal. 

Pronotum noticeably transverse, less than one-half wider than long, 
widest at the middle; disc very moderately convex, somewhat im- 
pressed laterally, very finely and sparsely punctulate; apex sub- 
truncate, scarcely emarginate, very distinctly and rather coarsely 
margined laterally, less strongly so at middle; sides evenly and 
moderately arcuate to basal fourth, thence feebly sinuate to base, 
margin distinct, moderate and slightly reflexed; base distinctly but 
not strongly arcuate, one-third wider than the apex, and coarsely 
margined; apical angles simply acute or minutely dentiform and not 
everted; basal angles obtuse, distinct, not rounded nor prominent. 

Propleure opaque, impunctate, and more or less obsoletely rugulose. 

Elytra oval, about twice as long as wide and somewhat ventricose, 
widest at the middle; base very feebly emarginate, scarcely wider 
than the contiguous prothoracic base; humeri obtuse, not prominent 
nor rounded; s‘des evenly arcuate from base to apex, somewhat 
gradually converging posteriorly, apex obtusely rounded ; disc moder- 
ately and quite evenly convex on the dorsum, more strongly and 
broadly so laterally, areuately declivous posteriorly; surface striato- 
punctate, some of the strize apparently duplicated, punctures quite 
coarse, usually separated by less than their own diameters and some- 
what impressed on the dorsum and more strongly so laterally; 
strie near the suture not duplicated, intervals more or less feebly 
convex, quite strongly so on the apical declivity, widest near the 
suture and narrower laterally, with a single series of minute and not 
very distantly spaced punctures. 


248 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


FE’ pipleure rather narrow, gradually narrowing from base to apex; 
surface obsoletely punctulate and smooth. “ 

Sterna subglabrous, more or less obsoletely punctate and rugose. 

Parapleure more or less obsoletely punctate. 

Abdomen horizontal, smooth and more or less shining, very sparsely 
and obsoletely punctulate, most strongly punctate on the last two 
segments, more or less rugulose. 

Legs moderate in length and quite slender; anterior femora armed 
in both sexes; tibial spurs similar, the anterior subequal and acute; 
anterior tarsi simple. 

Male.—FElongate, oval, scarcely robust or ventricose, somewhat 
attenuated behind; elytra arcuately and obliquely declivous pos- 
teriorly, apex rather narrowly rounded; abdomen moderately convex 
and rather broadly impressed on the first three segments; anterior 
femora with an acute and moderate tooth. 

Female.—Ovate and somewhat ventricose, not noticeably attenu- 
ated behind; elytra arcuately declivous posteriorly. Abdomen rather 
strongly and evenly convex. Anterior femora obtusely and not 
strongly dentate. 

Measurements—Male: Length, 25 mm.; width, 10.2 mm. Fe- 
male: Length, 29 mm.; width, 11.5 mm. 

Genital characters, male——Edeagophore elongately fusiform, 
scarcely arched and somewhat depressed. 

Basale oblong oval, narrowing cephalad, moderately convex, sides 
slightly arcuate. 

Apicale triangular, rather broad, subdepressed; surface very mod- 
erately convex, with a median and slightly widened linear membra- 
nous groove at middle two-fourths; sides feebly arcuate from base to 
apex, the latter subacute and not in the least produced; base rather 
broadly lobed at middle and broadly sinuate laterally. 

Sternite transverse. Each lobe rather short and broad, and sub- 
triangulo-parabolic in outline; external border arcuate to apex, the 
latter broadly rounded and quite continuously so with the internal 
margin; surface slightly convex externally, glabrous and shining in 
basal two-thirds and coarsely punctate, setose in the apico-internal 
third; seta quite long and rather dense. Sinus broad and apparently 
closed by membrane, the latter not setose at bottom. 

Female.—Genital segment trapezoidal, comparatively small, sides 
reflexed and more or less concave, not conspicuously setose. 

Valvula (Plate 5, fig. 18).—Dorsal plate oblong-oval, with a mod- 
erate antero-posterior convexity, concave, internal fourth reflexed, 
external three-fourths gradually and obliquely so; surface smooth 
and glabrous, not noticeably punctate nor setose; external and inter- 
nal borders more or less feebly arcuate; apical margin evenly arcuate, 


; 
) 
q 
; 
| 
| 
| 


REVISION OF ELEODIINI—BLAISDELL. 249 


not setose. Apex short, triangular, chitino-membranous and very 
finely setose. 

Appendage short, rather broad and obtusely subtriangular, flat- 
tened and partly concealed by the swollen superior margin of the 
fossa, finely setose, with two or three-longer sete at tip. Fossa trans- 
verse, margins more or less swollen and very finely setose and without 
conspicuous sete. 

Superior pudendal membrane reaching to about the middle third 
of the dorsal plate and rather distantly longitudinally rugulose. 

Basal prominences moderately prominent laterally; submarginal 
groove not visible from above. 

Ventrolateral surfaces—Body moderately stout, rather strongly 
convex and swollen in basal half and transversely concave before the 
apices; surface glabrous, very finely punctulate and sparsely setose 
in apical half, setee minute and fine. Submarginal groove small 
beneath the feebly laterally prominent external margin of the dorsal 
plate, and terminating at the external angle of the fossa. Inferior 
margin of the apex continuous with the inferior margin of the fossa. 
‘Internal margins of the valves contiguous in basal fourth. Genital 
fissure long and rather broadly fusiform, with basal half closed by 
the inferior pudendal membrane. 

Habitat—Texas (Cameron County, August, H. F. Wickham). 

Number of specimens studied, 2 (male and female). 

Sexitypes in Prof. Wickham’s and my own collections. 

Type-locality—Cameron County, Texas. 

Salient type-characters.—Somewhat opaque to feebly shining, ely- 
tra slightly ventricose. Thorax very moderately convex and evidently 
transverse; sides moderately arcuate and very slightly sinuate before 
the base; apical angles small, simply acute to feebly dentiform; basal 
angles obtuse. Elytra with striew of rather coarse, closely placed 
punctures, intervals feebly convex, distinctly convex on the apical 
declivity. Some of the strie duplicated. Intervals with a single 
series of minute punctules. 

Diagnostic characters.—The two specimens before me appear to 
be quite distinct. The slightly ventricose elytra, transverse and 
rather short pronotum, with the sides rather strongly rounded, small 
apical angles; elytra with striae of rather coarse punctures, which are 
coarser laterally, the intervals more or less convex and distinctly so 
on the apical extremity, general surface smooth and not at all muricate, 
with the integuments dull, give quite a characteristic facies. 

The female in form somewhat resembles the larger form of nupta 
(see hispilabris). 

Subpinguis is apparently most closely related to gracilis and 
dentipes. From the former it is separated by the much broader 


250 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


pronotum; from the latter by the subopaque integuments and shorter 
and broader pronotum, and it inhabits a different goological region. 

General observations.—The mentum is triangular, rounded at apex, 
sometimes reflexed so as to appear truncate. The inflexed lobes are 
moderate in size. Surface coarsely punctate and feebly convex cen- 
trally, foveate laterally within the margins. The prosternum is 
strongly arcuate antero-posteriorly between the cox and strongly 
protuberant ventrally with the coxe; triangular behind the equator 
of the coxe and vertically truncate, also compressed behind, with the 
angle rounded or quite strongly mucronate; surface longitudinally 
grooved between the coxe. 

Mesosternum quite vertical and more or less arcuate, broadly con- 
cave, 

The abdominal process is subquadrate and noticeably transverse, 
in width about equal to the length of the second segment; the latter 
is slightly longer than the post-coxal part of the first; the third is a 
little more than twice as long as the fourth, while the second is 
about a third of its length longer than the third. 

The abdominal salient is about a fifth wider than the metasternal 
salient. Metasternum laterally between the coxe is scarcely as long 
as the width of a mesofemur at base. 

The tibial grooves of the profemora are moderate in width and 
slightly concave, floors glabrous; sides carinate, quite straight and 
contiguous before the base; the anterior margin of each groove is 
more or less dentately laminate at about the outer fourth. 

On the mesofemora the grooves are glabrous, with the floors quite 
flat, the sides moderately cariniform, becoming evanescent at basal 
fourth without becoming contiguous. Those of the metafemora are 
similar but evanescent at basal third. 

The protibiz are slightly compressed and carinate externally. The 
tarsal grooves are more or less feeble, with their floors glabrous in 
apical moiety; the articular cavities are more or less feebly open. 

The meso- and metatibie are not carinate externally and the artic- 
ular cavities are quite closed; the external surfaces are flattened and 
scarcely grooved, glabrous on the mesotibiw, subglabrous and obso- 
letely sculptured on the metatibie. 

The tarsi are moderately stout and somewhat elongate. 

The protarsi are about a third of their length shorter than a 
mesotarsus. Joints two, three, and four subequal, apparently just 
a little wider than long and together about equal to the fifth; the 
first is slightly longer than wide. 

The mesotarsi are about one-eighth of their length shorter than 
a metatarsus. Joints two, three, and four subequal, just a little longer 
than wide and together just a little longer than the fifth; the first is 
scarcely as long as the third and fourth combined. 


REVISION OF ELEODIINI—BLAISDELL, 251 


The metatarsi are equal in length to half the length of a metatibia. 
' Joints two and three are subequal and distinctly longer than wide, 
and together equal to the fourth; the first is slightly shorter. 


ELEODES DENTIPES Eschscholtz. 


Bleodes dentipes Escuscuortz, Zool, Atlas, ITI, 1833, p. 10, pl. xrv, fig. 4.— 
MANNERHEIM, Bull. Soc. Nat. Moscow, XVI, 1843, p. 267; Mag. Zool., 
1834, p. 31, pl. cxvi.—LeContTrE, Proc. Acad. Nat. Sci. Phila., TS85s, 
p. 181.—Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 314. 

Eleodes prominens Casry, Annals N. Y. Acad. Sci. V, Nov., 1890, p. 401, 

Bleodes elegans Casry, Annals N. Y. Acad. Sci., V, Nov., 1890, p. 401. 

Bleodes conjinis BLAISDELL, Ent. News, VI, no. 7, Sept., 1895, p. 237. 


Elongate oval or ovate, smooth, and convex. 

Head twice as wide as long, feebly convex, very feebly and vaguely 
impressed laterally, rather coarsely, irregularly and densely punctate. 
Antenne usually moderate, but variable in length, outer four joints 
compressed, slightly and gradually dilated, third joint quite equal 
to the next two combined, fourth slightly longer than the fifth, the 
latter, sixth, and seventh subequal, eighth subtriangular in outline 
and very slightly shorter than the seventh, ninth, and tenth trans- 
versely oval or suborbicular, eleventh trapezoido-ovate. 

Pronotum widest at middle, about one-third wider than long, less 
than twice as wide as the head; disc more or less bright and shining, 
moderately convex, rather finely, sparsely, and irregularly punctate, 
usually obsoletely impressed, narrowly opaque, and granulo-punctate 
laterally along the margin in middle four-sixths, rather declivous at 
apical and basal angles; apex moderately and rather arcuately emar- 
ginate, obsoletely margined; sides broadly and evenly arcuate, dis- 
tinctly sinuate in basal fifth, and briefly subparallel before the angles, 
more or less finely margined, margin reflexed; base feebly arcuate 
and very finely margined, slightly wider than the apex; apical angles 
moderately acuminately produced and more or less everted; basal 
angles subrectangular or slightly prominent. 

Propleure more or less finely and irregularly granulato-muricate 
with a large smooth area posteriorly. 

Elytra oval, smooth and shining, slightly more than twice as long 
as wide, widest at the middle; base feebly emarginate, scarcely wider 
than the contiguous prothoracic base; humeri obtuse, not prominent 
nor rounded; sides evenly arcuate, apex obtuse and not rounded ; disc 
evenly and rather strongly convex from side to side, more or less 
gradually arcuately declivous posteriorly ; surface rather finely punc- 
tate, punctures arranged in moderately distant unimpressed series 
and quite closely spaced, intervals with a series of finer punctules, 
which are more distantly spaced, and become more irregular and 
denser along the suture; the punctures become more confused and 


252 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


very feebly muricate laterally and on apex; sometimes the punctures 
are subequal and diffuse. 

EB pipleure moderately wide at the humeri and ieee gradually 
narrowing to apex; surface very finely and obsoletely punctate. 

Sterna more or less densely punctate and rugulose. 

Parapleure rather coarsely punctate. 

Abdomen horizontal, rather finely and evenly punctate, rugulose, 
the punctuation denser on the fifth segment. 

Legs variable in length, form and stoutness. Anterior femora 
armed in both sexes; tibial spurs similar in the sexes, the anterior 
equal and acute; tarsi similar in the sexes. 

Male—FElongate, subfusiform oval, and somewhat narrow. An- 
tenne usually scarcely reaching to or slightly beyond the protho- 
racic base. Elytra gradually narrowed, and evenly, not rapidly 
arcuately declivous posteriorly; apex narrowly rounded and obtuse. 
Abdomen moderately convex, first two segments scarcely im- 
pressed. Anterior femora with a moderate and acute tooth at about 
outer third; posterior tibiz straight or arcuate. 

Female—Rather robust. Antenne usually attaining the posterior 
third or about to the base of the prothorax. Elytra more or less 
broadly oval, rather suddenly arcuately declivous posteriorly, with 
apex rather broadly rounded. Abdomen quite strongly convex and 
not at all impressed. Anterior femora armed with a more or less 
obtuse tooth; posterior tibiz not arcuate. 

Genital characters, male—Edeagophore of the usual flattened 
oblong-ovate form and rather small. 

Basale quite short and oblong, scarcely arched, but gradually gib- 
bous dorsally toward base. 

Apicale triangular; surface evenly convex, rather slightly de- 
pressed and with a feeble membranous groove in middle two-fourths; 
sides slightly arcuate and apex simply acute; base more or less ar- 
cuately lobed at middle and broadly sinuate laterally. 

Sternite slightly transverse. Each lobe with the external border 
evenly and broadly arcuate in-apical half and quite parallel toward 
base; internal border straight; apex narrowly rounded; surface 
densely and strongly punctate in apical moiety. Sete rather long 
and dense. Membrane not setose across the bottom of the sinus, the 
latter moderately narrow. 

Female.—Genital segment subtrapezoidal, rather robust, apex 
more or less chitinous and setose. 

Valvula (Plate 3, figs. 9 and 11).—Dorsal plate oblong, glabrous; 
surface moderately concave, very sparsely punctate, sete rather 
short; sides feebly reflexed; margins quite parallel, the external 
straight to feebly sinuate; internal border slightly arcuate; apical 


REVISION OF ELEODIINI—BLAISDELL. 253 


margin truncate to somewhat arcuate, not always well defined from 
apex internally; angle subrectangular. Apex short, rather coarsely 
punctate, sete rather short; fossa fringed with quite long hairs at 
outer margin, the hairs extending inward along the apical margin 
of the dorsal plate. 

Appendage short, scarcely as long as the apex, rather stout and 
conical, rounded at tip and set with rather short setee which form a 
loose tuft. 

Basal prominences rather strongly developed. 

Superior pudendal membrane not quite reaching to the middle of 
the dorsal plate and distantly longitudinally rugulose. 

Ventrolateral surfaces (Plate 3, fig. 10) moderately swollen at base 
and rather broadly concave laterally before the apices, glabrous, 
sparsely punctate and setose in apical moiety, sete moderately long, 
especially along the ventral margins of the fosse. Submarginal 
groove more or less developed. Internal margins of the valves con- 
tiguous in basal fourth; genital fissure fusiform and rather wide, 
with the inferior pudendal membrane visible in basal half. 

The above descriptions are applicable to the greater number of 
examples in the series before me. The present species is indeed the 
most variable and polymorphic of any in the genus. 

The large series at my disposal presents a complex of incipient 
races that are most bewildering. It is not wise nor practicable to give 
names to these forms and thus burden our faunal lists with names of 
races which can not be distinctively defined. These forms appear to 
diverge along certain lines of development and at the same time to 
pass directly into one another by the gradual diminution or augmen- 
tation of characters, as the case may be. It seems to me that it is best 
to select a few of the heterotypical forms as centers, around which the 
more incipient races may be clustered. 

Undoubtedly dentipes is unusually reactive to its environment, for 
each geographical region produces some peculiar modification of the 
typical form, and yet more or less typical examples are taken in the 
same regions. 

In accordance with the ideas just expressed I shall indicate the 
following forms for cabinet arrangement: 

Forma pertenuis.—Comparatively small and slender. Pronotum 
widest at or in advance of the middle; sides evenly arcuate and con- 
verging posteriorly, scarcely sinuate to distinctly so before the angles. 
Legs long and very slender. In the males the hind tibiwe are feebly 
arcuate in basal half, thence scarcely straight or gradually thickened 
to apex. 

In one specimen the elytral surface is very minutely reticulately 
rugulose, punctuation scarcely distinguishable. 


254 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Measurements —Males: Length, 18.5-19.5 mm.; width, 7-7.8 mm, 
Females: Length, 17.8 mm.; width, 7.2 mm. e 

Forma elongata—Comparatively large and distinctly elongate. 
Two subforms can be recognized. 

Forma elongata levis.—Alutaceous; antenne reaching to or beyond 
the prothoracic base; elytra very finely and obsoletely punctate, legs 
long, posterior tibiz very slender, and more or less arcuate in basal 
half in the males. 

Measurements.—Males: Length, 19.5-29 mm.; width, 6.5-9 -mm. 
Females; Length, 24-29.5 mm.; width, 8.5-10.5 mm. 

Forma elongata punctata.—This is more or less shining; antenne 
reaching to or beyond the prothoracic base; elytra with the serial 
punctures rather coarse and more or less impressed, the interstitial 
punctures are small but distinct. Legs moderately long and rather 
stout, posterior tibiz in the males stout and straight. 

Measurements.—M ales: Length, 24-28.5 mm.; width, 7.5-9.5 mm. 
Females: Length, 27.5 mm.; width, 10.5 mm. 

Forma typica.—Glabrous, very bright and shining; thorax with the 
sides strongly sinuate in front of the basal angles; legs moderate (see 
salient type-characters). 

Variations Examples of the typical form taken about San Fran- 
cisco Bay and in the series before me frequently present the follow- 
ing variations: Antenne rather stout or slender, varying in length 
irrespective of sex, rarely two of different lengths in the same in- 
dividual; the relative length of joints differs in individuals of the 
same sex; outer four joints may be noticeably dilated, subclavate, 
again not in the least dilated; anterior pronotal angles simply acute 
and not prominent to acuminately produced and very strongly 
everted, the pronotum itself may be subquadrate; elytra sometimes 
widest behind the middle. 

Measurements—Males: Length, 22-28.5 mm.; width, 6.5-8.5 mm. 
Females: Length, 23-28 mm.; width, 8.5-9.5 mm. 

Forma prominens Casey.—Iorm somewhat as in dentipes, convex, 
smooth, and alutaceous throughout. 

Head moderate, finely, sparsely punctate, more densely so toward 
the sides of the epistoma. Antenne short and very robust, much 
shorter than the head and prothorax, third joint about twice as long 
as wide and equal in length to the next two. 

Prothorax about one-sixth wider than long; disc rather strongly 
convex throughout, finely and sparsely punctate, strongly aluta- 
ceous, the side margins very minutely beaded; qpea broadly arcuate, 
sinuate laterally, and subequal to the base; sides broadly arcuate an- 
teriorly, rather strongly convergent in basal half, and strongly sinu- 
ate near the basal angles; base broadly, evenly, very feebly arcuate; 


See 


REVISION OF ELEODIINI—BLAISDELL, 255 


apical angles acute, very prominent anteriorly, and distinctly everted ; 
basal angles acute and prominent. 

_Scutellum strongly transverse, broadly rounded behind, highly 
polished, and impunctate. 

Elytra at base broadly, feebly emarginate and about as wide as the 
contiguous prothoracic base; Awmeri obtuse, but not rounded nor 
prominent; s¢des evenly arcuate, the elytra gradually narrowed and 
acute at apex; disc widest at the middle, where it is rather more than 
one-third wider than the prothorax, feebly alutaceous, finely, sparsely 
punctate, the punctures nearly simple, arranged in widely distant, 
rather feebly defined, unimpressed rows, the intervals with a few 
widely distant and nearly similar punctures. 

Legs rather short and slender, the anterior femora with a short 
obtuse tooth near the outer third; middle and hind femora simple: 
spurs of anterior tibiz moderate in length, slender, similar, and very 
nearly equal. 

Measurements.—Length, 19 mm.; width, 7.8 mm. 

Forma robusta.—Very robust, elytra less narrowed toward base, 
prothoracic base wide, integuments very smooth and subalutaceous; 
punctuation very fine or obsolete. Prothorax comparatively large, 
apex arcuate at middle and sinuate laterally ; sides more or less sinuate 
before the base. Antenne and legs quite stout to somewhat slender. 

Measurements.—M ales: Length, 24-28 mm.; width, 9-10 mm. 
Females; Length, 22-27 mm.; width, 9-10.5 mm. 

Forma confinis Blaisdell.— (See Plate 3, fig. 13.) 

Rather robust and alutaceous, very convex. Head large; antenne 
robust, reaching to the posterior third of the prothorax. Pronotum 
widest just in front of the middle, sometimes at the middle, poste- 
riorly straight, convergent and not in the least constricted or sinuate 
in front of the basal angles, the latter obtuse. Legs moderately 
slender. 

Measurements.—M ales: Length, 24 mm.; width, 7.5 mm. /emales: 
Length, 24 mm.; width, 8.5 mm. 

The following unique, described by Colonel Casey, may be simply 
a form of dentipes or an aberration: 

Eleodes elegans Casey.—Moderately robust, strongly convex, 
strongly alutaceous, the elytra moderately shining; integuments 
nearly smooth. 

Head moderate, somewhat finely, sparsely punctate. Anfenna 
rather short but slender, not quite as long as the head and prothorax, 
third joint about two and one-half times as long as wide, scarcely as 
long as the next two, fourth much longer than the fifth. 

Prothorax nearly one-third wider than long; disc rather strongly 
convex throughout, widest just before the middle, somewhat sparsely 


256- BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


but deeply and rather coarsely punctate: apex transversely truncate, 
a very little narrower than the base; s¢des strongly, almost evenly 
arcuate throughout, and very minutely beaded; base very feebly, 
evenly arcuate; apical angles small but acute, anteriorly prominent, 
dentiform, and feebly everted; basal angles very obtuse, not rounded, 
but not in the least prominent. 

Elytra about three times as long as the prothorax, gradually acute 
at apex, inflated, widest at the middle, where they are about three- 
fourths wider than the prothorax; base just visibly wider than the 
base of the prothorax; humeri obtuse but not rounded, not at all 
prominent; sides evenly arcuate; disc confusedly, finely creased, 
rather coarsely and distinctly granulato-reticulate, very finely and 
sparsely punctate, the punctures nearly simple throughout and not 
denser laterally, arranged in very feebly defined unimpressed rows, 
approximate toward the suture, then widely distant, with an uneven 
and sparser row along the middle of the intervals. 

Legs moderate in length and very slender; anterior femora not 
dentate but with a very broad and obtuse rounded salient near the 
apex: spurs of the anterior tibie very slender and almost exactly 
equal. 

Measurements.—Length, 13 mm.; width, 6 mm. 

Colonel Casey writes that there is no described species with which 
it can be compared, and says that it should be placed near dentipes. 
If the sexes have the anterior femora armed it can be placed nowhere 
else. The type is evidently a female and the femoral tooth should 
be more acute in the male. It does not differ any more than many 
other aberrant forms of dentipes which I have seen. 

The following Aberrations represented by uniques may be men- 
tioned to make known extreme forms that are probably fortuitous: 

Aberration No. 1.—Antenne slender; sides of the pronotum evenly 
arcuate from apex to base; basal angles obtuse. Elytra widest be- 
hind the middle. Legs slender and moderate in length; anterior 
femora with a small subacute tooth at about outer third; posterior 
tibiz quite slender, the anterior moderately arcuate. Integuments 
glabrous and shining. Female——Length, 20.5 mm.; width, 8.5 mm. 
Taken with typical specimens at Berkeley, California. 

Aberration No. 2—Moderately robust, subopaque, and alutaceous, 
antenne reaching slightly beyond base of the pronotum, the latter 
subquadrate; base and apex truncate and equal to each other; sides 
evenly but not strongly arcuate, feebly sinuate just before the basal 
angles, the latter rectangular. 

Legs rather long and slender. 

This specimen might at first glance be taken for a female guad- 
ricollis. The apical angles of ‘the pronotum are very acute and 
everted at right angles. 


uw 


el, 


REVISION OF ELEODIINI 


BLAISDELL. 257 


Female —Length, 21.5 mm.; width, 9 mm. 

Taken at Mokelumne Hill, Calaveras County, California. 

In the latter aberration the anterior femora have a small obtuse 
salient. In other respects both possess other characters as in dentipes. 

Habitat.—California. 

Forma pertenuis.—Kaweah, Tulare County, Ralph Hopping; Wat- 
son Springs, May; Martinez, Contra Costa County, January, collec- 
tion U. S. National Museum; Colony Mills, Tulare County, elevation 
5,415 feet, Fuchs and Hopping. 

Forma elongata—/wvis.—Mokelumne Hill, Calaveras County, ele- 
vation 2,400 feet, March to August; Fresno County. 

Punctata—Placerville, Eldorado County; Corning, Tehama 
County; Martinez, Contra Costa County; San Jose, San Mateo 
County. Y 

Forma typica.—Counties bordering the Bay of San Francisco; Port 
Harford, San Luis Obispo County; Ocean Beach, Los Angeles 
County, J. J. Rivers; Napa County, F. C. Clark; Trinity County, 
BE. C. Van Dyke; San Joaquin County; Argus Mountains; Yosemite 
Valley; San Bernardino Mountains. 

Forma prominens.—Port Harford, San Luis Obispo County, Col. 
Thomas Casey. 

Forma robusta.—Fairmount, Los Angeles County; Fort Tejon, 
Fuchs and Hopping; Kern County, Fordyce Grinnell; Tulare 
County; Calaveras County; Napa County. . 

Forma confinis—Mokelumne Hill, Calaveras County, elevation 
2.300 feet; Napa County; Santa Clara’ County; Tulare County; 
Sonoma County. 

Forma elegans.—Hoopa Valley, Humboldt County, Col. Thomas 
Casey. 

Number of specimens studied, 960. 

Disposition of the type unknown to me. 

Type-locality—San Francisco; “on the hills beneath rocks.” 

Salient type-characters—Elongate and shining. Head and thorax 
finely granulato-punctate; thorax with sides rounded and strongly 
sinuate at base; apical angles acuminate. Elytra finely striato-punc- 
tate, the punctures granuliferous (Eschscholtz). 

Diagnostic characters.—<A very variable species both as to sculptur- 
ing and general form. 

From acuticauda it is recognized by the Jess strongly rounded sides 


-of the pronotum and in being less strongly and rather less suddenly 


constricted at base, not broadly granular at the sides of the dise and 
the elytra are never decidedly produced. 
In forma elongata the sides of the pronotum are most distinctly 
granulate of any of the variations, but less so than in aeuticauda var. 
59780—Bull. 63 —09——17 


258 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


laticollis, from which it is at times difficult to separate, but here the 
form of the pronotum must be relied upon. : 

Armata and militaris have all of the femora armed, while in the 
present species only the anterior bear a tooth. 

Dentipes is undoubtedly genetically related to armata (see p. 262). 

The chief incipient races have already been sufficiently indicated 
and must only serve as centers for cabinet arrangement, for the meso- 
types are multitudinous and defy limitations. 

The middle lobe of the mentum is moderate and subtriangular, at 
times sub-trapezoidal or more or less parabolic, and varies independ- 
ent of race or form; the apex is usually more or less distinctly areu- 
ate; surface coarsely punctate and very faintly convex and quite 
narrowly but not strongly foveate laterally within the margin. 

The prosternum variable; usually rounded between and behind the 
coxe; frequently with a small, and less frequently with a well- 
developed mucro. In forma confinis some specimens have it sub- 
truncate and feebly compressed behind, with the angle mucroid. 

Mesosternum quite obliquely precipitous and more or less concave. 

The abdominal process is subquadrate, slightly transverse and in 
width about equal to the length of the third abdominal segment; the 
post-coxal part of the first is about equal to the second, and the latter 
is about twice as long as the fourth. 

The abdominal salient is about a fourth of its width wider than the 
metasternal process. 

The metasternum laterally between the coxe is about as long as the 
width of a mesotibia at apex. 

The tibial grooves of the femora are well developed, with edges 
more or less cariniform, and their floors vary in the degree of con- 
cavity. Those of the profemora do not quite attain the femoral base; 
on the mesofemora the grooves extend inwards to about the basal 
fourth and there the margins become evanescent before becoming con- 
tiguous; on the metafemora they only attain the internal third and 
there become evanescent before uniting. 

The tibie vary greatly in sculpturing, at times very densely and 
coarsely muricate and at others rather feebly so. The articular cavi- 
ties are nearly always closed and the grooves are always more or less 
roughly sculptured with their floors more or less shining, and 

variable in degree of development. The external borders of the pro- 
tibiz are more or less distinctly carinate, the mesotibiz scarcely at 
all so. 

The tarsi are very variable in length and stoutness, even in the 
same race, 

In an average typical specimen the protarsi are about a sixth of 
their length shorter than a mesotarsus. Joints two, three, and four 


itil 


— —— <<. | 


SC 


REVISION OF ELEODITNI—BLAISDELL, 259 


are subequal and slightly wider than long, and together about equal 
to the fifth; the first about a half longer than wide. 

The mesotarsi are about a seventh of their length shorter than a 
metatarsus. Joints two, three, and four about as long as wide and 
subequal, and together about equal to the fifth; the first is about one 
and a half times longer than wide. 

The metatarsi are about half as long as a metatibia. Joints two 
and three subequal and distinctly longer than wide, together scarcely 
as long as the fourth, and slightly longer than the first. 


ELEODES ARMATA LeConte. 


Bleodes armata LrContr, Ann, Lye. Nat. Hist. N. Y., V, 1851, p. 134; also 
in Thomson’s Arcana Nat., 1859, p. 125, pl. x1, fig. 2; Proc. Acad. 
Nat. Sci. Phila., 1858, p. 181.—Horn, Trans. Amer. Phil. Soc., XIV, 
1870, p. 310. 

var. impotens BLAISDELL, Ent. News, VI, no. 7, Sept., 1895, p. 2386. 


Elongate, somewhat oblong-ovate, shining, sometimes alutaceous. 

ITead twice as wide as long, moderately convex, more or less im- 
pressed laterally, frequently the frons is broadly and transversely 
impressed with vertex more strongly convex; surface rather coarsely, 
moderately, densely, and irregularly punctate. Antenne not reach-. 
ing to the base of the prothorax, outer four joints slightly compressed 
and searcely dilated, third joint scarcely longer than the next two 
combined, fourth longer than the fifth, the latter to the seventh, inclu- 
sive, subequal, eighth obconico-triangular, ninth and tenth scarcely 
transversely oval, eleventh ovate. 

Pronotum widest at or slightly in advance of the middle, about a 
third wider than long; disc smooth, moderately and evenly convex, 


‘rather declivous at the apical angles, rather sparsely and evenly 


punctulate; apex rather deeply and broadly emarginate, usually 
finely margined laterally; sdes broadly arcuate anteriorly, more or 
less straight, or feebly arcuate to base, sometimes sinuate before 
the base, finely beaded ; ase not strongly arcuate and rather coarsely 
margined, about a sixth wider than the apex: apical angles acute, 
more or less dentiform, frequently everted; basal angles obtuse. 

Propleure smooth, with small setigerous punctures anteriorly, 
along posterior margin and on the acetabular convexity. 

Elytra oval, smooth, about twice as long as wide, widest at the 
middle, rarely behind the same; base scarcely emarginate, and hardly 
wider than the contiguous prothoracic base; Awmeri obtuse and not 
prominent; sides evenly arcuate, apex obtuse and more or less nar- 
rowly rounded; dise smooth, evenly convex from side to side, more 
or less arcuately declivous posteriorly; swrface more or less finely 
punctate, punctures arranged in moderately distant, unimpressed 


260 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


series, the punctures sometimes rather coarse, at other times fine and 
quite closely placed in the series, intervals flat, with a single more 
or less regular series of widely spaced and very fine punctures; the 
punctuation becomes more or less irregular on the sides and apex. 

E'pipleure moderate in width at the humeri, thence gradually nar- 
rowing to the apex; surface smooth, finely and very sparsely punc- 
tulate, or impunctate, frequently finely rugulose and feebly concave. 

Sterna more or less punctate. 

Parapleure rather coarsely and evenly punctate. 

Abdomen horizontal, sparsely, finely, and quite evenly punctured, 
punctures slightly denser on the fifth segment. 

Legs moderately long; all of the femora coarsely and quite densely 
punctate, and each armed with a long, acute, and strongly devel- 
oped tooth; all of the tibiz generally more or less arcuate in basal 
half; anterior tibial spurs equal and acute; tarsi simple and similar 
in the sexes. 

Male-—Elongate, slightly narrower than the female. Antenne 
reaching about to the posterior fifth of the prothorax. Elytra some- 
times distinctly narrowed posteriorly and more or less obliquely de- 
clivous; apex obtuse, not produced, and rather narrowly rounded. 
Abdomen moderately convex, and broadly, very feebly impressed on 
the first two or three segments. 

Female.—More or less robust. Antenne scarcely attaiming the 
posterior fifth of the prothorax. Elytra more or less evenly arcu- 
ately declivous posteriorly ; apex rather broadly rounded. Abdomen 
strongly convex and not impressed. 

Measurements.—Males: Wength, 25-28.5 mm.; width, 8-9 mm. 
Females: Length, 24-32 mm.; width, 8-11.5 mm. 

Genital characters, male—Edeagophore of the usual flattened 
oblong-ovate form and but slightly arched. 

Basale about a third of its length longer than the apicale, evenly 
convex above, with the sides evenly arcuate. 

Apicale very slightly elongate, triangular, evenly convex above, 
with a narrow, membranous median groove in apical half; sides 
shghtly arcuate or straight, apex subacute, not produced and nar- 
rowly rounded; base broadly and evenly rounded at middle, feebly 
sinuate laterally. 

Sternite with sides converging apicad. Each lobe rather small; 
external border straight in basal half and arcuate apically, with the 
apex broadly and evenly rounded; internal margin slightly arcuate; 
surface rather densely punctate and set with quite long and densely 
placed seta, especially at apex and apical border. Membrane not 
setose at bottom of the sinus, the latter rather broad. 

Female.—Genital segment trapezoidal, moderate in size, usually 
not strongly setose, 


REVISION OF ELEODITNI—BLAISDELL. 261 


Valvula (Plate 1, fig. 18).—Dorsal plate oblong, with the internal 
border strongly reflexed in apical half, external three-fourths grad- 
ually reflexed and outwardly oblique; surface glabrous, very slightly 
concave, strongly so along the reflexed internal border, very sparsely 
and finely punctate, sete short and not conspicuous; external border 
nearly straight or slightly arcuate; internal margin more or less 
sinuous; apical margin subtruncate, angle broadly and_ eyenly 
rounded, with a few rather short and not very conspicuous seter, 
Apex small and about as long as the appendage, finely setose and 
_ without tuft. 

Appendage somewhat flattened, subconical, and more or less con- 
cealed by the apical margin of the dorsal plate; finely setose and 
without a conspicuous tuft. 

Superior pudendal membrane attaining the middle of the dorsal 
plate. ; 

Basal prominences usually not weil developed. 

Ventrolateral surfaces—Body not inflated nor strongly convex, 
slightly concave before the apex and laterally; surface smooth, with 
few fine punctures and setw about the fossa; apices finely setose. 
Submarginal groove fine and normal under the slightly prominent 
external margin of the dorsal plate. Internal margins of the valves 
contiguous in basal third. Genital fissure narrowly fusiform, with 
the inferior pudendal membrane closing its basal half. 

Habitat—California (eastern San Diego County; Los Angeles 
County; Kern County; Coso Valley; Panamint Valley, and Death 
Valley, April; Needles; Fresno; Tulare County; Palm Springs; 
Merced County; Mono County; Amedee, Lassen County, July, ele- 
vation, 4,200 feet, H. F. Wickham; Fort Tejon; Argus Mountains) ; 
Arizona (Tacna, April, Hubbard and Schwarz; Fort Yuma; Pres- 
cott; Martinez (Congress Junction); Bill Williams Fork, F. H. 
Snow). Lower California (coll. Calif. Acad. Sciences). 

Number of specimens studied, 112. 

Type in the LeConte collection. 

Type-locality—California (gold dise label) in the desert near the 
Colorado ‘River. 

Salient type-characters.—Thorax finely and scarcely densely punc- 
tate; sides rounded, and narrowed posteriorly; basal angles obtuse. 
Elytra convex, apex slightly attenuate, punctures more or less dis- 
tinetly seriately placed. Femora always with a long and acute tooth 
beneath (LeConte). 

Diagnostic characters —Quickly recognized from all other species 
north of Mexico—except-militaris and femorata—by having all of the 
femora armed. 

Armata differs from the above two species mainly in its larger 
size and in having the elytra less attenuate, also, rather more sud- 


262 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


denly declivous behind. The femoral teeth are also usually much 
stouter, longer and more acute, but these last charfcters are variable 
and in a large series lose their differential value. 

In the typical race the apical angles of the pronotum are acute, 
usually prominent anteriorly and dentiform. The angles present 
considerable variation in a large series of specimens, and if the 
examples be arranged according to the form of the angles, they will 
be found to pass from that observed in the typical, to small, acute, 
dentiform and everted in the other extreme of the series. 

These remarks apply also to the femoral teeth; in the extreme 
forms as opposed to the normal, they become long, slender, and acic- 
ular, or even shorter and triangular, even obtuse or entirely aborted 
on the meso- and metafemora (forma subedentata), even where there 
is no change in the bodily form. 

In the collection of the California Academy of Sciences there is a 
specimen from Lower California, and referred to /ucw by Dr. George 
Horn, that has only the anterior femora armed and without any evi- 
dence of the other femoral teeth; this specimen has the facies of 
armata—the elytra being unusually and very gradually attenuated 
posteriorly, without any signs of the formation of a definite cauda. I 
believe this specimen to be a subedentate armata. 

A most interesting series was received from Fresno County, Cali- 
fornia, by Mr. Fuchs for the California Academy of Sciences. To 
my mind this series demonstrated the fact that armata passes into 
dentipes by the failure of development in the teeth upon the meso- 
and metafemora. In the series were specimens with the teeth in dif- 
ferent degrees of atrophy, and some with only the anterior femora 
armed; in general facies they could not be separated from the un- 
doubted typical forms in the same series (see impotens). Many 
specimens of fully armed armata have the pronotum nearly as 
strongly sinuate posteriorly as in dentipes. 

General observations—The mentum is also variable. The middle 
lobe may be moderate or smaller in size, subtriangular to trapezoidal, 
the apex evenly rounded or subtruncate. The surface is coarsely 
punctate and not noticeably setose, feebly convex centrally:and more 
or less strongly foveate laterally, or quite plane and not foveate. 

The prosternum is strongly sculptured and strongly mucronate, 
usually horizontally so, and flat between the cox; frequently convex 
between the same and horizontally mucronate, or rarely with a small 
mucro; sometimes vertically truncate behind, with the angle rectan- 
gular or strongly mucroid and prominent. 

Mesosternum short, scarcely at all horizontal, nearly entirely and 
obliquely vertical, more or less strongly concave. 

The abdominal process is subquadrate, a little transverse and just 
a little wider than the third segment is long. The second segment is 


———— 


REVISION OF ELEODIINI—BLAISDELL. 263 


about equal in length to the post-coxal part of the first; the fourth is 
half as long as the third. 

The abdominal salient and that of the metasternum are subequal 
in width. 

The metasternum laterally between the coxe is about equal in 
length to the width of a mesofemur at base. 

The tibial grooves of the femora are strongly developed and, de- 
fined; the margins are more or less cariniform; the floors are gla- 
brous and more or less concave. The profemoral grooves nearly 
attain the femoral base, while those of the meso- and metafemora 
become more or less obsolete at the basal fourth or third, respectively. 

The anterior margin of each groove is more or less laminate, and 
dentately so at about the outer fourth. The femora are distinctly 
sinuate external to the teeth, and the anterior surface is longitudinally 
and inwardly impressed from the bases of the teeth. It must also be 
noted that the metatibiaw, while arcuate in basal half, may also be 
straight and slightly widened in the apical half (see wickhami). 

The tibiw are distinctly arcuate in basal half. The articular 
cavities are more or less open and the tarsal grooves more or less well 
developed, although rather narrow and usually defined by asperate 
margins. The floors are quite smooth. 

The protibiz are distinctly carinate externally, and the meso- and 
metatibiw are scarcely at all carinate externally. 

The tarsi are usually quite stout and moderate in length. 

The protarsi are about a sixth of their length shorter than a meso- 
tarsus. Joints two, three, and four are short, subequal, and distinctly 
wider than long, and together distinctly shorter than the fifth; the 
first is slightly longer than the second. 

Each mesotarsus is about a seventh of its length shorter than a 
metatarsus. Joints two, three, and four are subequal and evidently 
just a little wider than long, and together about equal to the fifth; 
the first is about as long as wide. 

The metatarsi are distinctly less than half as long as a metatibia. 
Joints two and three are subequal and longer than wide, and together 
decidedly shorter than the fourth; the first is aboutne-half longer 
than wide. 


ELEODES ARMATA var. IMPOTENS Blaisdell. 


Elongate, subovate, convex, and moderately shining. 

Head about twice as wide as long, moderately convex, irregularly 
and rather sparsely punctate, punc.ares moderate in size and denser 
at the periphery. Antenne short, moderately slender, subclavate, 
outer four joints compressed and rather moderately dilated, third 
joint equal to the next two combined, fourth slightly longer than the 


264 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


fifth, the latter, sixth, and seventh equal and cylindrical, eighth tri- 
angular, ninth and tenth transversely oval, eleventh ovate. 

Pronotum widest a little in front of the middle and slightly wider 
than long; désc evenly and a little more than moderately convex, 
more or less declivous at the apical angles, finely and sparsely punc- 
tate; apex truncate to feebly emarginate, finely margined laterally; 
sides evenly arcuate in the anterior two-thirds, thence less so or some- 
what straight and converging to base, sometimes feebly sinuate, very 
finely margined; base feebly arcuate, not coarsely margined and 
slightly wider than the apex; apical angles small and acutely denti- 
form, at times strongly everte&; basal angles subrectangular and not 
prominent. 

Propleure opaque, more or less finely and very sparsely punctate, 
frequently the acetabular convexities are strongly rugulose. 

Elytra oval and more cr less elongate, smooth, widest at the mid- 
dle; base truncate or feebly emarginate, and about equal to the con- 
tiguous prothoracic base; humeri obtuse and not prominent; sides 
evenly arcuate, apex obtuse; disc evenly and quite strongly convex 
or somewhat depressed, more or less arcuately declivous posteriorly ; 
surface finely punctate, punctures closely and quite regularly placed 
in the rather close series, intervals with a single series of more dis- 
tantly and regularly placed punctures, that often become larger in 
the outer intervals; both more irregular and denser at the periphery. 

E pipleure moderately wide and gradually narrowing from base to 
apex; surface obsoletely and sparsely punctulate. 

Sterna usually rather finely punctate and frequently quite strongly 
rugose. 

Parapleure rather opaque, more or less sparsely, irregularly, and 
rather coarsely punctate. 

Abdomen finely and sparsely punctate, punctures denser on the 
fifth segment, the first frequently rugulose. Horizontal. 

Legs slender to moderately stout, and also moderate in length; 
femora quite densely punctate, and all armed; .the anterior with an 
acute tooth, the middle with a smaller and obtuse salient, the pos- 
terior with a sll smaller and very obtuse salient; tibial spurs rather 
slender and moderately long and acute, the anterior subequal; tarsi 
similar in the sexes. 

Male.—Elongate and rather narrow. Antenne reaching to about 
the posterior fourth of the prothorax. Elytra rather gradually 
arcuato-obliquely declivous posteriorly. Abdomen moderately con- 
vex and broadly impressed and with a stronger longitudinal median 
impression. Posterior tibizv more or less arcuate in basal half. 

Female.—Elongate and rather robust, antenne usually not attain- 
ing the posterior fourth of the prothorax. Elytra more or less 
broadly oval and somewhat suddenly arcuato-obliquely declivous 


REVISION OF ELEODIINI—BLAISDELL, 265 


posteriorly. Abdomen strongly convex, not impressed. Posterior 


tibizw scarcely arcuate. 

Measurements.—M ales: Length, 21.5-28 mm.; width, 7-9.5 mm. 
Females; Length, 21-28 mm.; width, 7.5-10.5 mm. 

Genital characters, male-—Edeagophore of the usual depressed 
oblong-ovate form, and slender. 

Basale not arched, sides nearly parallel; surface evenly convex, 
scarcely gibbous. 

Apicale triangular, slightly elongate, feebly decurved; surface 
evenly convex, feebly depressed, with a linear groove in apical half 
and not extending upon the apex; sides evenly and very moderately 
arcuate; apex very narrowly rounded and subacute; base broadly 
lobed at the middle, scarcely sinuate laterally. 

Sternite moderately, transverse. Each lobe subparabolic with the 
external border evenly arcuate and moderately converging to apex; 
apical margin rather broadly rounded, angle scarcely evident; in- 
ternal border subarcuate; surface feebly convex and rather densely 
punctate and setose; sete long and quite dense at apical margin. 
Membrane not setose across thé bottom of the sinus. 

Female—Genital segment subtrapezoidal, moderate in size, and 
setose. 

Valvula—Dorsal plate oblong, concave internal border rather 
strongly refiexed in apical two-thirds and impunctate, longitudinally 
subcanaliculate external to the same; surface glabrous, finely, evenly, 
and sparsely punctate and setose in apical two-thirds; sete reclinate 
and rather short; external border feebly arcuate; internal margin 
more or less arcuate, sinuate near base; apical margin evenly arcuate, 
angle rounded. Apex rather long, subacute at tip, not densely 
punctate, setw small and slightly denser at tip; superior margin of 
the fossa set with rather long sete, not denser at angle. 

Appendage flattened, moderately wide, subconico-semiellipsoidal, 
and covered with rather long set in apical half. 

Superior pudendal membrane reaching to the middle of the dorsal 
plate and longitudinally rugulose. 

Basal protuberances moderate. 

Ventrolateral surfaces.—Body slightly inflated, not strongly con- 
eave laterally, but feebly transversely so before the apices; surface 
finely and sparsely punctate, setose, sete short, longer on the inferior 
margin of the fossa. Submarginal groove small; external margin of 
the dorsal plate slightly prominent laterally. Internal margins of 
the valves contiguous in basal third. Genital fissure narrowly fusi- 
form, inferior pudendal membrane visible in the basal half. 

Variations.—Apex small; appendage as long as the apex and sub- 
conical; apical margin of the dorsal plate less arcuate, with angle | 


266 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


narrowly rounded. Dorsal plate with a slight antero-posterior con- 
vexity. 

Habitat—California (Livingston, Merced County, July, F. E. 
Blaisdell, Charles Fuchs; Fresno, collection of the California Acad- 
emy of Sciences; and the collection of E. A. Schwarz; San Joaquin 
County, D. W. Coquillett). 

Number of specimens studied, 22. 

Type in my own collection, female. 

Type-locality Livingston, Merced County, California. 

Salient type-characters—Pronotum moderately convex; dise 
smooth, very finely and sparsely punctate; apex truncate; apical 
angles dentiform, small, acute, and strongly divergent; sides moder- 
ately arcuate in the anterior two-thirds, thence feebly convergent to 
base, scarcely sinuate; basal angles almost. rectangular and not 
prominent. Elytra with punctures moderately fine, arranged in 
rather closely placed, feebly impressed series, intervals with similar, 
sparsely, and irregularly placed punctures. The anterior femora 
with an acute tooth, the middle with an obtuse process, the posterior 
with a very small and obtuse salient. ° 

Diagnostic characters.—Form variable. It differs in a general way 
from armata by being more slender and usually considerably smaller, 
and especially in having the teeth of the hind femora rudimentary 
or obsolete, while those of the mesofemora are distinct or rudimen- 
tary, and no doubt obsolete at times, and such individuals (forma 
subedentata) are scarcely separable from dentipes. The thorax may 
be as in typical armata or more or less constricted at base as in 
dentipes. Individuals of armata with normal femoral teeth occasion- 
ally have the prothorax constricted nearly as in dentipes and can be 
spoken of as forma sinuata. 

No doubt but that zmpotens is a transitional form between armata 
and dentipes, as it occurs chiefly at the periphery of the area of 
distribution of armata. Dentipes appears to be found mainly be- 
yond the area inhabited by armata. Some examples of impotens 
have the general form of gigantea. 

General observations.—The middle lobe of the mentum is rather 
large, trapezoidal and slightly broader than usual, apex more or less 
arcuate, and the surface evenly and feebly convex. 

The sterna and abdomen as in armata. 

The articular cavities of the protibie are more or less open. The 
other tibiz and ferhora are as in armata. 

The tarsi may be slender or stout and moderate in length. 

The protarsi are scarcely a fourth of their length shorter than a 
mesotarsus. First four joints subequal, the first apparently as long 
as wide, the others a little wider than long, and together about equal 
to the fifth. 


“Te. ea 
 s* 


* 


REVISION OF ELEODIINI—BLAISDELL. 267 


The mesotarsi are about equal in length to a metatarsus. First 
four joints apparently subequal and just a little longer than wide; 
joints two to four, inclusive, together a little longer than the fifth. 

The metatarsi are a little less than a half as long as their metatibiz. 
Joints two and three subequal and distinctly longer than wide, and 
together shorter than the fourth; first joint the least bit longer than 
the second. 


ELEODES MILITARIS Horn. 
Bleodes militaris Torn, Trans. Amer. Phil. Soc., XTV, 1870, p. 310. 


Elongate oval, moderately shining and convex. 

Head twice as wide as long, usually moderately convex, and more 
or less impressed about the eyes, not coarsely and rather sparsely 
punctate at middle, punctures becoming denser at the periphery. 
Antenne moderate, outer four joints feebly or not at all dilated, 
slightly compressed, third joint nearly twice as long as the fourth, 
the latter subequal to the fifth, sixth, and seventh individually, the 
latter and eighth about as wide as long, ninth and tenth suborbicular, 
eleventh truncate ovate. 

Pronotum subquadrate, widest at the middle, a fifth to a third 
wider than long; disc evenly and rather strongly convex, more or 
less declivous laterally at apex, finely and evenly punctate, punctures 
but slightly denser at the sides; apex rather deeply emarginate, finely 
margined; sides evenly and moderately arcuate, scarcely oblique 
posteriorly or at times converging to base, finely margined; base 
truncate or feebly arcuate and not very coarsely margined; apical 
angles acute, not dentiform and moderately advanced, or rarely sub- 
dentiform and feebly everted; basal angles obtuse and not prominent. 

Propleure smooth, with a few scattered punctures posteriorly and 


on the acetabular convexities, more or less rugulose. 


Elytra elongate oval, smooth, moderately attenuated behind, widest 
at the middle; dase subtruncate or feebly emarginate, and equal to 
the contiguous base of the prothorax; humeri subobtuse, not rounded 
nor prominent; sides evenly arcuate, apex more or less produced and 
rather narrowly rounded; disc strongly convex, gradually arcuately 
declivous behind; surface finely punctate, punctures arranged in dis- 
tant unimpressed series, those of each series rather closely placed, 
intervals with a single series of distantly placed punctures, punctua- 
tion becoming denser and irregular at the sides and apex. 

Epiplura rather wide and gradually narrowing from base to 
apex; surface smooth and obsoletely punctate. 

Sterna not densely and more or less coarsely punctate; prosternum 
anteriorly finely and sparsely punctate and more or less transversely 
rugulose. 

Parapleure moderately sparsely and not coarsely punctate. 


268 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Abdomen horizontal, rather finely and sparsely punctate, denser on 
the fifth segment. ad 

Legs moderate, sometimes quite robust; all of the femora armed 
with an acute triangular tooth in both sexes, and coarsely, densely 
punctate; all of the tibize more or less arcuate in basal half; tibial 
spurs acute and subequal on their respective tibie; tarsi similar in 
the sexes. 

Male.—Antenne reaching to the posterior fifth of the prothorax, 
the latter with base one-half wider than the apex and a fourth 
greater than the length, a third wider than long. Abdomen moder- 
ately convex, broadly and not strongly impressed. Femora robust, 
very coarsely and rather strongly punctate, punctures often deeply 
impressed. Sometimes the femoral teeth are very strong. 

Female.—Antenne scarcely reaching to the posterior fifth of the 
prothorax, the latter with the base about equal to the length, about 
a fifth wider than long, and a fourth wider than the apex. Abdo- 
men evenly and strongly convex. 

Measurements— Male: Length, 22.5-28 mm.; width, 7.5-9.5 mm. 
Female: Length, 22-25 mm.; width, 7.5-8.5 mm. 

Genital characters, male. agophore of the usual flattened 
oblong-ovate form, and somewhat oblong. 

Basale with the sides scarcely parallel, about two and a half times 
longer than wide, slightly or not arched, very strongly and quite ab- 
ruptly convex from side to side, with rather a strong antero- “posterior 
convexity, the disc appearing gibbous at middle. y 

Apicale triangular, slightly elongate and decurved apically; sur- 
face evenly and moderately convex, and with a linear groove in apical 
half, that does not extend upon the apex; sides arcuate at base, 
feebly sinuate towards apex, the latter slightly produced and nar- 
rowly rounded at tip; base broadly lobed at middle, scarcely sinuate 
laterally. 

Sternite moderately transverse. Each lobe subtriangular; exter- 
nal border nearly evenly arcuate; internal margin nearly straight, 
with apex rounded; surface strongly punctate in apical half, setose, 
setee not dense, and moderate in length. Sinus rather broad, mem- 
brane not setose across the bottom of the same. 

Female.—Genital segment trapezo-triangular, moderate in size 
and rather strongly setose. 

Valvula (Plate 1, fig. 20).—Dorsal plate oblong, concave, sides 
evenly reflexed; surface finely, evenly, and not very densely punctate 
throughout, sete short and reclinate; external border quite straight; 
internal border more or less sinuous; apical margin more or less arecu- 
ate, angle obtuse. Dorsal margin of the fossa conspicuously fringed 
with rather long sete, those at the angle a little longer and denser. 
Apex moderate and triangular, tip acute with few short hairs. 


REVISION OF ELEODIINI—BLAISDELL. 269 


Appendage flattened, semi-ellipsoidal, comparatively large, pune- 
tate; and strongly setose at apical moiety, sete long. 

Superior pudendal membrane attaining the middle of the dorsal 
plate and longitudinally rugulose. 

Basal protuberances rather well developed. 

Ventrolateral surfaces—Body not inflated and very feebly con- 
cave laterally and before the apéx; finely and sparsely punctate, 
setose throughout, sete small and reclinate; inferior margin of the 
fossa with a few rather long sete. Submarginal groove not well de- 
veloped, dorsal plate scarcely expanded laterally. Internal margins 
of the valves contiguous in basal third. Genital fissure narrowly 
fusiform, closed in basal half by the inferior pudendal membrane. 

Habitat—Lower California (collection of Hubbard and Schwarz; 
Comondu, San José del Cabo, San Quintin, collection of California 
Academy of Sciences; Santa Margarita Island, March, collected by 
Charles D. Haines, collection of-Charles Fuchs). Arizona (Charles 
Fuchs). 

Number of specimens studied, 12. 

Type in the Horn collection ; collected by William M. Gabb. 

Type-locality Lower California. 

Salient type-characters.—Male, thorax subquadrate, sides rounded, 
feebly converging at base; apex rather deeply emarginate, angles 
acute, not dentiform, basal angles not prominent. Female, thorax a 
third broader than long, moderately rounded at the sides, not con- 
verging at base, apex as in the male, base notably broader than apex. 
Elytra strongly attenuated behind, with distant rows of rather fine 
and distinct punctures (Horn). 

Diagnostic characters —Militaris is separated from armata by the 
femoral teeth being less produced, broader and more equilaterally tri- 
angular. The anterior angles of the prothorax are less prominent 
and less strongly dentiform, everted or not. The elytra are more 
attenuated behind. 

The insect appears more obtuse in front on account of the stronger 
discal convexity of the pronotum. <A large series would no doubt 
show variations in the femoral teeth; the male specimen from Santa 
Margarita Island has the teeth more produced than those from the 
mainland. Ina single specimen from Calamajuet, Lower California, 
all of the elytral punctures are rather coarse and very distinct, not 
impressed and the femoral teeth approach those of armata in form. 

There is in this species a greater antero-posterior dorsal convexity 
than is observed in armata. 

In another specimen before me the general form is that of a male 
acuticauda—the elytra being gradually produced at apex with the: 
dorsal outline continuously arcuate; the punctures are small; length, 
28 mm.; width, 9 mm. 


270 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


The series at hand does not support Doctor Horn’s statement in 
regard to the form of the prothorax in the two se&es. A male from 
Santa Margarita Island has the base of the prothorax notably wider 
than the apex and the sides do not converge at base; a series of six 
females from Lower California (mainland) have the prothorax sub- 
quadrate, base slightly wider than the apex, and the sides converge. 

The series is not large enough to decide this question; evidently 
the prothorax is variable; it is to be noted that the specimens are 
from two separate and distinct regions. The male is often more 
robust than the female. 

In several males before me the femora are very stout, compressed, 
and quite broad; the femoral teeth are stout and subequilaterally 
triangular. 

The elytral strize may be quite strongly impressed at times, with 
the intervals feebly or moderately convex. 

This species is probably a southwestern modification of armata. 
Forms occur in Arizona which can not be separated from this species. 

A specimen collected at Palm Springs, California, and recently re- 
ceived for identification, has the integuments more opaque, form 
more robust and oblong, elytra with the sides feebly arcuate and more 
rapidly narrowing in apical fourth; elytral punctures fine and equal 
in both series; antenne slender and elongate, the femora compara- 
tively slender and the anterior femoral teeth small. 

Two specimens under examination have the general form of mili- 
taris, although more elongate; the anterior femora are alone armed, 
the middle and posterior being edentate. One of these specimens was 
collected at San Quentin, Lower California (collection California 
Academy of Sciences), and referred to Zucw by Doctor Horn. I do 
not believe that it should be referred to that species, for the facies is 
not that of Zwcw, and I have found that the prothorax may vary sufli- 
ciently in almost any species to simulate that of another. T have 
demonstrated to my own mind that armata may become subedentate 
by loss of the teeth of the middle and posterior femora, and thus pass 
into dentipes; furthermore, I see no reason why militaris and femo- 
rata should not vary in a similar way. 

In both specimens the epipleure at the elytral apices are dilated a 
little more than usual for species of the present section, but I consider 
this character more of a fortuitous exaggeration of the indifferent 
stage of the caudal development, or possibly it is atavistic reversion. 
The relationship of this interesting form can not be determined until 
better collecting has been done on the peninsula. I therefore for the 
present define this form as follows: 

Forma subedentata.—Form as in mlitaris, with the anterior femora 
alone armed, 


REVISION OF ELEODIINI-——BLAISDELL, 271 


Tt will be observed that the sides of the front at the junction with 
the epistoma are frequently slightly emarginate at the suture. 

General observations.—The mentum is moderate and the middle 
lobe is nearly parabolic, being arcuately rounded in the examples that 
I have seen; the surface is very feebly convex and sometimes very 
feebly foveate laterally. 

The prosternum is horizontal or more or less convex between the 
coxee, surface longitudinally grooved or concave posteriorly ; mucro- 
nate behind, truncate or subtruncate. The ante-coxal portion is quite 
short at times and with the coxie prominent ventrally. 

Mesosternum as in armata. 

The abdominal process is subquadrate, slightly transverse, and 
about as wide as the post-coxal portion is long; the second segment is 
about twice as long as the fourth; the third about a third of its length 
shorter than the second. 

The abdominal salient is slightly wider than the metasternal salient. 

The metasternum laterally between the coxe is about as long as the 
width of,a mesotibia at apex. 

The tibial grooves of the femora are well developed and strongly 
defined, broader in the peninsular examples and somewhat contracted 
in the insular specimens. The anterior margins of all are quite carin- 
iform, most strongly so in the robust examples, and strongly den- 
tately laminate, internally rather gradually passing into the teeth. 
The floors are opaque, sometimes subasperate on the profemora, 
generally so on the meso- and metafemora. The anterior grooves 
have the sides more or less arcuate and quite attaining the femoral 
base; on the other femora the sides of the grooves are more or less 
evanescent internally; on the mesofemora the grooves attain the in- 
ner fifth and on the metafemora to about the inner fourth. All of the 
femora are longitudinally impressed inwardly from the tooth along 
the lower part of the anterior surface.’ 

The protibie are more or less arcuate externally and distinctly 
carinate; the tarsal grooves are more or less developed and _ sca- 
brously sculptured. In the peninsular specimens the articular cavities 
are closed and in the insular more or less open. 

The mesotibiz are more or less carinate in basal third externally. 
The metatibizw are not carinate, and both have the articular cavities 
closed. The tarsal grooves of the mesotibiw are more or less evident 
and scabrous. The metatibie are flattened or feebly grooved and 
muricate externally; quite cylindrical in transverse section. 

The tarsi are variable, moderately slender to quite stout, quite 
moderate in length. 

The protarsi are about a fifth of their length shorter than a meso- 
tarsus. Joints quite closely articulated, second, third, and fourth 


272 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


subequal, wider than long, and together about equal to the micas the 
first but little longer than the second. : 

The mesotarsi are about a sixth of their length shorter than a meta- 
tarsus. Joints two, three, and four are sabeael and about as long 
as wide, together a little longer than the fifth; the first is about as 
long as wide. 

The metatarsi are just a little more than a third as long as a meta- 
tibia. Joints two and three subequal, just a little longer than wide, 
together subequal to the fourth and a little longer than the first. 


ELEODES MILITARIS var. FEMORATA LeConte. 


-Eleodes femorata LECONTE, Ann. Lyc. Nat. Hist. N. Y., V, 1851, p. 184; 
Proc. Acad. Nat. Sci. Phila., 1858; p. 181.—Horn, Trans. Amer. Phil. 
Soc., XIV, 1870, p. 311. 

Elongate oval, more or less shining, smooth and convex. 

ITead rather ell twice as wide as long, moderately convex, obso- 
letely impressed laterally and along the frontal suture; rather finely 
but not densely punctate, punctures slightly denser at the sides. 
Antennw moderate, feebly compressed in outer five joints, very grad- 
ually and very focbly dilated, third joint about equal in length to the 
next two taken together, fourth just noticeably longer than the fifth, 
the latter to the seventh, inclusive, subequal in length, the seventh 
nearly quadrate, eighth as wide as long, ninth and tenth very feebly 
transversely oval, eleventh subtrapezoidal. 

Pronotum widest at middle, slightly wider than long and nearly 
twice as wide as the head; disc quite strongly and evenly convex, 
finely and evenly punctulate; apex broadly and feebly emarginate. 
very finely beaded; sides evenly, broadly and moderately arcuate 
with a feeble tendency to become sinuate posteriorly, very finely 
headed ; base very feebly arcuate or truncate and finely beaded, about 
a fourth wider than the apex-and about equal to the length; apical — 
angles small, subacute, not prominent anteriorly, with a slight ten- 
dency to eversion; basal angles obtuse, not rounded nor prominent. 

Propleure smooth, obsoletely punctulate and more or less rugulose. 

‘Elytra oval, more or less gradually attenuated posteriorly, feebly 
emarginate or truncate at base and equal in width to the base of the 
prothorax; Awmeri obtuse, not rounded nor prominent; stdes evenly 
arcuate, apex subacute: d¢/sc quite evenly convex from side to side, 
gradually and arcuately declivous. posteriorly; surface finely and 
irregularly but not densely punctate, punctures never impressed 
and scarcely denser at the sides, frequently showing a serial arrange- 
ment centrally, with the interstitial punctures distanthy and irregu- 
lar placed: suture on the apical declivity usually more or less im- 
pressed. 


-. - 


; REVISION OF ELEODIINI—BLAISDELL. 273 


Epipleure moderately wide at the humeri and gradually narrow- 
ing to apex; surface subopaque and obsoletely punctulate. 

Sterna more or less sparsely punctate and irregularly rugose. 

Parapleure more or less sparsely punctate. 

Abdomen horizontal, finely, quite evenly, and rather sparsely punc- 
tate, more or less rugulose. 

Legs rather long and quite densely sculptured. All of the femora 
armed in both sexes with small and more or less acute teeth, the 
acuity diminishing from before backward, so that the teeth of the 
metafemora are very small and obtuse; tibiz more or less arcuate 
in basal third; anterior tibial spurs about equal in size and length. 
Tarsi similar in the sexes. 

Male—Moderately narrow. Antenne attaining the posterior fifth 
of the prothorax. Elytra strongly attenuate behind, very evenly and 
very gradually arcuately declivous posteriorly. Abdomen rather 
strongly convex and feebly or obsoletely impressed at middle of first 
two segments. Posterior tibiz rather strongly arcuate in basal third. 

Female.—Rather robust. Antenne attaining the posterior fourth 
of the prothorax. Elytra not strongly attenuated behind, moderately 
and gradually arcuately declivous posteriorly. Abdomen evenly con- 
vex from side to side. Posterior tibiz not more strongly arcuate than 
the mesotibie. > 

Measurements—Males: Length, 22-23 mm.; width, 8-8.5 mm. 
Females: Length, 25 mm.; width, 9.6 mm. 

Genital characters, male-—Edeagophore flattened oblong-ovate, very 
feebly arched in apical half, moderate in size and elongate. 

Basale oblong, evenly convex from side to side, sides evenly and 
feebly arcuate. 

Apicale rather elongately triangular; surface strongly convex, 
without or with a very faint median membranous groove; sides feebly 
sinuate in apical two-thirds or straight; apex more or less produced 
and rather slender, quite acute; base with a median subacute lobe, 
quite broadly and distinctly sinuate laterally. 

Sternite transverse. Each lobe subquadrate, with the external bor- 
der quite straight, meeting the apical margin at somewhat of an 
angle; apical margin feebly and evenly arcuate and nearly trans- 
verse, angle introrse and subacute; internal border more or less sinu- 
ous; surface scarcely convex, subglabrous, finely and evenly but not 
densely punctate, setose, setee quite long and subequal and not dense. 
Membrane not setose across the bottom of the sinus, the latter short 
and oblong. 

Female.—Genital segment subtrapezoidal in outline, quite strongly 
chitinized, setose and rather shining. 


597T80—Bull, 68—09——18 


274 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Valvula (Plate 4, fig. 18).—Dorsal plate suboblong, and widened" 
somewhat in basal half; surface deeply concave, Very sparsely and 
finely punctate, setulose; margins more or less reflexed; external 
border subangulate at junction of the middle and basal thirds, thence 
straight or feebly sinuate to apical margin, the latter more or less — 
feebly arcuate with the angle evenly and rather broadly rounded; 
internal margin more or less feebly arcuate. Apex narrow and more 
or less chitinous, rather acute and finely setose at tip, setee quite short 
and not noticeable over the general surface. 

Appendage strongly flattened, quite large and semioval, about as 
long as the apex, quite densely punctate in apical half and rather 
densely set with long hairs. Fossa narrow, margin fringed with 
sparsely placed and moderately long sete that become rather dense 
at the angle. Submarginal groove very fine and more or less visible 
from above. 

Superior pudendal membrane reaching slightly beyond the middle 
of the dorsal plate, and with three or four longitudinal rugule. 
Valvular membranes visible beyond the pudendal membrane. 

Ventrolateral surfaces—Body evenly convex in basal half and 
more or less transversely and rather broadly impressed before the 
apices; surface smooth and shining, rather evenly punctate and 
setose, setae short and reclinate. Internal margins of the valves 
contiguous in basal half. Genital fissure very narrowly fusiform and 
nearly closed. . 

Habitat—Three specimens before me were collected by myself at 
San Diego on the Coronado peninsula. They were in company with 
omissa, beneath some roots of Yucca baccata. 

Number of specimens studied, 3. More recently 9 others. 

Type in the LeConte collection. 

Type-locality—San Diego, California. 

Salient type-characters.—Thorax finely and not densely punctate, 
sides rounded, somewhat narrowed posteriorly, basal angles obtuse. 
Elytra with apex attenuated, punctures more or less distinctly 
serially arranged, intervals sparsely punctulate. Femora always 
somewhat briefly dentate (LeConte). 

Diagnostic characters——This race is to be separated from militaris 
by the femoral teeth being more obtuse. The thorax is less broad, 
especially in the females (7), and the elytra are slightly more atten- 
uated at apex; the antero-posterior convexity of the dorsal surface 
of the insect is slightly more evident. 

In my opinion femorata is undoubtedly a more northern modifi- 
cation of mélitaris, and in a large series this view would, no doubt, 
be verified. 

It is interesting to note that LeConte’s descriptions of armata 
and femorata are the same, word for word, except: “ elytris parum 


REVISION OF ELEODIINI—BLAISDELL. 275 


attenuatis, femoribus * * longe acuteque dentatis” (armata), 


and “elytris * * * apice attenuatis, *. * *  femoribus 
* * *  breviter dentatis” (femorata). 

Femorata is quite distinct from armata, the chief differences 
being the form of the elytra and the femoral teeth—both very varia- 
ble characters. 

General observations.—The mentum is subparabolic. In one speci- 
men it is subtrapezoidal; surface scarcely convex and not foveate 
laterally. 

The prosternum in the specimens at hand is arcuately rounded 
antero-posteriorly, with a small and abruptly developed mucro be- 
hind, which may be horizontal or deflexed. Mesosternum as ir 
armata. 

The abdominal process is subquadrate, feebly transverse; the post- 
coxal portion of the first segment is equal to the second in length; 
the third is as long as the width of the metasternal salient; the fourth 
is about one-half as long as the second. ? 

The intercoxal process of the abdomen is about a fourth wider than 
the metasternal salient. 

The metasternum laterally between the coxe is as long as the width 
of a mesotibia at apex. 

The tibial grooves of the femora are well developed and distinctly 
defined; their floors are opaque and quite plane. Those of the pro- 
femora are subentire, those of the meso- and metafemora nearly 
attain the femoral base, and the carinate margins are somewhat eva- 
nescent internally. The anterior margin of each groove is more or 
less dentately laminate externally. 

The protibiz are somewhat compressed and distinctly carinate 
externally; their tarsal grooves are more or less developed, opaque, 
and asperately sculptured. The articular cavities are more or less 
open. 

The mesotibiz are carinate externally in basal third; the grooves 
are more or less indicated, opaque, and muricate. The articular cavi- 
ties closed. . 

The metatibiew are more or less finely subcarinate externally, and 
the grooves are scarcely evident, muricately sculptured; articular 
cavities closed. Both the meso- and metatibiw are slightly com- 
pressed. 5 

Tarsi moderate in length and rather stout. The protarsi are about 
a sixth of their length shorter than a mesotarsus. Joints one to four, 
inclusive, subequal and a little wider than long; joints two to four 
combined equal in length to the fifth. 

The mesotarsi are about a seventh of their length shorter than a 
metatarsus. Joints two to four inclusive subequal and very little 


276 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


longer than wide; together scarcely longer than the fifth; the first 
joint is about one and a half times longer than wide. 

The metatarsi are a little less than half as long as a metatibia. 
Joint two is evidently just the least longer than the third, both are 
longer than wide, and together distinctly shorter than the fourth, 
also a little longer than the first. 


ELEODES ACUTICAUDA LeConte. 


Eleodes acuticauda LEContTE, Ann. Lyc. Nat. Hist. N. Y., V, 1851, p. 135; 
Proc. Acad. Nat. Sci. Phila., 1858, p. 181.—Horn, Trans. Amer. Phil. 
Soc., XIV, 1870, p. 314. 

Var. laticollis LeContTE, Ann. Lyc. Nat. Hist. N. Y., V, 1851, p. 135. 


Elongate, more or less shining, strongly convex, elytra more or 
less strongly produced and not suddenly caudate. 

Head slightly more than twice as wide as long, quite evenly convex, 
moderately finely and evenly punctate. Antenne rather stout, mod- 
erately short, outer four joints slightly compressed, more or less 
feebly and gradually dilated, third joint equal in length to the next 
two combined, fifth, sixth, and seventh subequal in length and each 
slightly shorter than the fourth, eighth trapezoidal in outline, ninth 
and tenth wider than long, eleventh subtrapezoidal. 

Pronotum widest at about the middle, about four-sevenths wider 
than long and more than twice as wide as the head; disc more or less 
bright and shining, moderately convex, noticeably declivous at base 
and apex, rather narrowly and more or less impressed and opaque 
laterally, finely, irregularly and somewhat densely punctate, some- 
times granulate laterally; apex moderately emarginate, obsoletely 
or distinctly margined; sides very strongly arcuate in anterior two- 
thirds, thence to base less so or somewhat straight and converging, 
and strongly sinuate in front of the basal angles, more or less finely 
margined; base moderately arcuate and rather coarsely margined, a 
little narower than the apex; apical angles small, acuminately pro- 
duced and dentiform, very acute and more or less everted; basal 
angles obtuse, sometimes quite prominent. 

Propleure smooth, obsoletely punctulate and rugulose. 

Elytra elongate oval, smooth and more or less shining, widest at 
the middle; base feebly emarginate and not wider than the contiguous 
prothoracic base, Awmeri obtuse and not at all prominent; sides 
evenly arcuate, frequently broadly and feebly sinuate at sides of 
apex, the latter gradually produced; disc quite evenly and strongly 
convex from side to side, more or less gradually and arcuately de- 
clivous posteriorly, or sometimes more or less broadly sinuate before 
the apex; surface smooth, punctate, the punctures closely placed in 
longitudinal series, the latter moderately closely placed, intervals 


Se eS 


REVISION OF ELEODIINI—BLAISDELL, 277 


irregularly and more or less sparsely punctulate, punctures usually 
not noticeably denser laterally. Elytral suture on apex more or less 
deeply impressed. Apex or cauda concave beneath, the surface of 
which is formed by the internal elytral surface. 

Epipleure slightly widened at base, thence gradually narrowing 
to apex, where it narrowly margins the inferior surface of the pro- 
duced apex or cauda; surface smooth and usually impunctate. 

Sterna more or less punctate and rugose. 

Parapleure more or less punctate. 

Abdomen horizontal, obsoletely or rather strongly punctate, rugu- 
lose. 

Legs more or less moderate, anterior femora armed in the sexes; 
anterior tibial spurs equal and acute. Tarsi similar in the sexes, 

Male.—Elongate. Antenne attaining the posterior fourth of the 
prothorax. Elytra usually more than twice as long as wide and more 
or less strongly and gradually produced at apex.. Abdomen rather 
strongly convex, scarcely at all flattened or impressed. Legs some- 
what stout, especially the anterior femora, which are armed with 
a very acute tooth; posterior tibiz more or less arcuate. 

Female.—Robust. Antenne attaining the posterior third of the 
prothorax. Elytra suboval, and usually less than twice as long as 
wide, rather moderately obliquely declivous posteriorly; apex more 
or less produced. Abdomen strongly convex. Legs moderate; ante- 
rior femora with a more or less acute tooth; posterior tibie straight. 

Genital characters, male.—Edeagophore of the usual oblong-ovate 
form and rather elongate. 

Basale slightly or not arched, quite evenly convex. 

Apicale triangular, slightly elongate, not decurved at tip; surface 
rather evenly convex more strongly so apically where the sides appear 
compressed, groove at middle two-fourths, fine toward apex and 
slightly dilated toward base; sides feebly sinuate in apical two- 
thirds; apex rather acute and gradually narrowed; base broadly 
lobed at middle, lobe rounded, laterally sinuate. 

Sternite transverse, sometimes quite oblong, lobes longer than wide. 
Each lobe with the external border evenly and broadly arcuate apic- 
ally, quite straight toward base; internal margin nearly straight; 
apex narrowly rounded; surface punctate and setose. Membrane not 
setose across the bottom of the sinus, the latter rather broad. 

Female.—Genital segment trapezoidal, moderate in size, glabrous 
and setose. 

Valvula (Plate 3, fig. 7) —Dorsal plate oblong, slightly concave, 
sparsely punctate and setose in apical two-thirds, sete moderately 
short, semirecumbent ; external border straight or feebly and broadly 
sinuate; internal margin quite strongly reflexed in apical half, more 


278 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


or less sinuate toward base; apical margin squarely truncate and 
transverse; angle rectangular or narrowly rounded, with a few rather 
long set that are also sparsely distributed along the apical margin. 
Apex small, subacute, with a tuft of a few rather short sete. Fossa 
rather large. 

Appendage quite large, flattened and triangular or semi-elliptical, 
broad as long, setose toward tip, with a tuft of quite long sete. 

Superior pudendal membrane attaining the middle of the dorsal 
plate and quite distantly longitudinally rugulose. 

Ventrolateral surfaces (Plate 3, fig. 8)—Body slightly inflated at 
base, rather strongly and transversely concave before the apex, rather 
strongly and sparsely punctate; sete along the ventral margin of the 
fossa rather long and flying. Apex quite densely setose beneath. 
Submarginal groove well developed beneath the projecting external 
border of the dorsal plate. Internal margins of the valves contiguous 
in basal third, genital fissure rather broadly fusiform and closed in 
basal half by the inferior pudendal membrane. 

Two forms may be recognized as follows: 

Forma typica—Glabrous. Elytra with striz of small unimpressed 
punctules or punctures which are frequently not very evident, inter- 
vals somewhat or very irregularly punctulate. Males dimorphic. 
The typical males are very strongly but not suddenly caudate; the 
atypical males with the females have the elytral apices moderately or 
scarcely produced, and not caudate. 

Measurements.—Males: Length, 16.5-30 mm.; width, 5-10.5 mm. 
Females: Length, 24.5-28.5 mm.; width, 8-9.5 mm. 

Forma punctata—Glabrous. Antenne stout, attaining the pos- 
terior third of the prothorax. Elytra with serial punctures moder- 
ately large, often eroded and feebly impressed, the series rather closely 
placed and often irregularly duplicated, intervals irregularly and 
finely punctulate, rather more densely so laterally, males apparently 
homomorphic and with the females have the elytral apex moderately 
produced and not strongly caudate. Legs rather stout. 

Measurements.—Males: Length, 29-98 mm.: width, 8-8.5 mm, 
Females: Length, 22.5-26.5 mm.; width, 8.5-10.5 mm. 

Habitat.—Forma typica——California (San Diego County; San 
Diego, elevation 50-300 feet. F. E. Blaisdell, E. C. Van Dyke, 
Charles Fuchs, and collection U. S. National Museum; Poway, eleva- 
tion 700 feet, F. E. Blaisdell; Los Angeles County, E. C. Van Dyke; 
Santa Barbara; Fort Tejon, April 19-21, Fuchs and Hopping; Tulare 
County; San Bernardino County); Lower California (San Pedro 
Martir, collection California Academy of Sciences). 

Forma punctata.—California (Los Angeles County, E. C. Van 
Dyke). 


Number of specimens studied, 120. 


REVISION OF ELEODIINI—BLAISDELL. 279 


Type in the LeConte collection. 

Type-locality.—San Diego, California. 

Salient type-characters.—Thorax finely punctate, broad and greatly 
rounded at the sides, which are subdepressed, narrowed behind ; basal 
angles obtuse, somewhat more prominent than usual. Elytra strongly 
convex, finely and seriately punctate, posteriorly declivous, strongly 
attenuated, and produced (LeConte). 

Diagnostic characters—The prothorax is usually at least one-half 
wider than long, sides very strongly rounded and strongly con- 
stricted at base; anterior angles acute, prominent and dentiform, but 
generally smaller than in Jaficollis, and frequently everted. The ely- 
tra in the typical form are elongate and strongly produced in the 
males, not suddenly caudate, and directed obliquely downward. 

In examples from Fort Tejon the males have the ‘elytra distinctly 
caudate, the cauda horizontal and formed nearly as in duce. In the 
other forms the elytra are less elongate and moderately produced. 
The elytra are never sulcate, but simply with striz of small punce- 
tures that are at times rather coarse and rarely impressed. 

The elytral surface is never muricate, and is in this character sepa- 
rated from /aticollis, the latter having the pronotum more strongly 
punctate, duller, and rather broadly impressed and granulate later- 
ally; in acuticauda the pronotum is narrowly or not at all impressed 
nor granulate laterally. 

From dentipes it must be separated by the form of the pronotum. 
In acuticauda the sides of the pronotum are more strongly and less 
broadly arcuate and the constriction is more rapidly formed ‘and 
larger; in dentipes the sides of the pronotum are less strongly and 
more broadly arcuate, the sinuation is shorter and more gradually 
formed. In some examples of aeuticauda the pronotum is not wider 
than in dentipes, and these are the difficult specimens to place, but 
the form of the arcuation still remains the same, and a quick eye can 
readily catch the difference. 

In forma typica the males may have the hind tibiz arcuate in basal 
half and more or less gradually thickened apically. 

General observations —The middle lobe of the mentum is moder- 
ate and rather wide, subparabolic, sometimes subtrapezoidal in out- 
line, the apex is seldom truncate; surface slightly convex centrally 
and more or less feebly foveate laterally within the margin. 

The prosternum is variable, usually convex or horizontal between 
the cox and with a moderate mucro behind; rarely rounded or sub- 
truncate behind. The mucro may be horizontal or deflexed. 

Mesosternum is more or less oblique and concave. 

The abdominal process is quite quadrate and in width equal to the 
length of the third segment; the post-coxal part of the first segment 


280 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


is twice as long as the fourth; the second is a little longer than the 
post-coxal part of the first. ~ 

The abdominal salient is about a fourth of its width broader than 
that of the metasternal process. 

The metasternum laterally between the cox in length is about 
equal to a mesotibia at middle. 

The tibial grooves of the femora are well developed and scarcely 
entire, the margins are cariniform and become evanescent, usually at 
the internal fourth; their floors are smooth and quite flat. 

The anterior margin of the profemoral grooves become quite sud- 
denly dentately laminate at about the external fourth. 

The protibiz are more or less distinctly carinate externally and 
feebly compressed, their tarsal grooves are more or less evident and 
scabrous and the articular cavities are closed. 

The mesotibiz have the grooves more or less developed and muri- 
cate and the articular cavities are closed; externally obsoletely cari- 
nate in basal fourth. The metatibiz are more or less flattened exter- 
nally and roughly sculptured, not at all carinate and the articular 
cavities closed. 

The tarsi are moderate in length and quite stout. 

The protarsi are about a third of their length shorter than a meso- 
tarsus. Joints two, three, and four are quite closely articulated, sub- 
equal, wider than long and together about equal to the fifth; the first 
is about as long as wide. 

The mesotarsi are very little shorter—about an eighth of their 
length—than a metatarsus. Joints two, three, and four are subequal, 
as long as wide, and together quite equal to the fifth; the first is 
slightly shorter than the next two taken together. 

The metatarsi are a little less than half as long as a metatibia. 
Joints two and three are subequal and distinctly longer than wide, 
together just the least shorter than the fourth, and a little longer 
than the first. 


ELEODES ACUTICAUDA var. LATICOLLIS LeConte. 


Elongate oval, more or less shining and smooth, strongly convex 
and not caudate, interstitial punctures of the elytra more or less 
muricate. 

Head rather large, twice as wide as long, rather moderately con- 
vex, very feebly and broadly impressed laterally, finely, irregularly 
and rather sparsely punctate, quite densely so laterally, punctures 
finely muricate, frontal suture ‘obsolete. Antenne moderately stout, 
reaching to the base of the prothorax, outer four joints feebly com- 
pressed and very slightly dilated, third joint very slightly longer than 
the next two combined, fourth slightly longer than the fifth, the 


REVISION OF ELEODIINI—BLAISDELL. 281 


latter, sixth, and seventh subequal, eighth triangular to triangulo- 
conical, ninth and tenth suborbicular in outline, eleventh subtrape- 
zoidal. 

Pronotum widest at about the middle, almost a half wider than 
long; disc moderately convex, more strongly so at the angles, rather 
finely, distinctly, irregularly and not densely punctate, rather 
broadly impressed and granulo-punctate laterally; apea broadly 
and quite evenly emarginate, distinctly margined laterally, ‘and 
obsoletely so at the middle; s¢des greatly rounded in anterior three- 
fourths, thence strongly sinuate and converging to base, rather 
coarsely margined; base feebly arcuate, equal to the apex and 
coarsely margined; apical angles strong, acuminately produced and 
dentiform, more or less everted, and somewhat reflexed at times; 
basal angles subrectangular and more or less prominent. 

Propleure rather smooth, irregularly and sparsely granulo-muri- 
cate, obsoletely rugulose. 

Elytra oval to elongate oval, widest at the middle; Jase subtrun- 
eate, more or less slightly wider than the contiguous prothoracic 
base; humeri obtuse, not rounded nor prominent; sides evenly 
arcuate, apex gradually narrowed, rarely attenuated and not pro- 
duced; disc moderately, convex on the dorsum and more strongly 
so laterally, arcuately declivous posteriorly; surface punctate, punc- 
tures arranged in rather distant series, moderately small and rather 
closely placed, intervals with a series of similar punctures, slightly 
more distantly spaced, on the dorsum very faintly muricate, laterally 
distinctly so and frequently more distantly spaced, both series be- 
coming more irregular and finely muricate on the apex, punctures 
minutely setigerous. 

Epipleure moderate at the humeri and gradually narrowing to 
apex; surface sparsely and finely submuricately and_ obsoletely 
punctate. 

Sterna more or less densely, submuricately punctate, and rugose. 

Parapleure rather coarsely and irregularly punctate, punctures 

rather shallow. 
' Abdomen horizontal, rather densely, somewhat finely, submuri- 
cately punctate and rugulose. 

Legs rather long and somewhat stout; anterior femora armed in 
both sexes; anterior tibial spurs subequal and acute. Tarsi similar 
in the sexes. 

Male.—Somewhat narrow. Elytra arcuately and rather obliquely 
declivous behind. Abdomen moderately convex and slightly flat- 
tened at middle on the first three segments. Anterior femora armed 
with an acute tooth. Metatibiw more or less arcuate. 


“O82 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Female—Somewhat robust. Elytra arcuately declivous posteri- 
orly. Abdomen evenly convex. Anterior femora with an obtuse or 
subacute tooth. Metatibie straight. 

Genital characters, male-—Edeagophore oblong-oval, depressed and 
not arched. 

Basale oblong; surface moderately convex and the sides quite 
parallel. 

Apicale subequilaterally triangular; surface evenly convex, 
coarsely and sparsely punctate, median groove linear and fine in 
middle three-fifths; sides feebly arcuate from base to apex, the latter 
subacute and not produced; base broadly and arcuately lobed at 
middle, feebly sinuate laterally. 

Sternite transversely subcrescentiform. Each lobe wider than long; 
external border broadly and evenly arcuate from base to apical 
angle, which is introrse at the internal margin, the latter short; angle 
rounded; surface moderately convex, quite densely punctate in apical 
three-fourths; setose, sete quite long and quite dense. Membrane 
not setose across the bottom of the sinus, the latter short. 

Female.—Genital segment rather small, setose, and triangularly 
trapezoidal in outline, moderately short. 

Valvula (Plate 3, fig. 12).—Dorsal plate gblong with sides slightly 
reflexed and moderately concave; surface glabrous, very sparsely 
punctate and setose, sete moderate in length and somewhat erect; 
external border straight or feebly sinuate; internal margin feebly 
arcuate; apical margin subtruncate, angle narrowly rounded, both 
fringed with rather long hairs, which are somewhat dense at the 
angle; apex small and short, triangular, finely setose, with a tuft of 
shghtly longer hairs at tip. 

Appendage rather small ‘and conical, equal in length to the apex, 
finely setose, with a tuft of longer hairs at tip. 

Basal prominence small and not very evident. 

Superior pudendal membrane attaining the middle of the dorsal 
plate; finely but not strongly longitudinally rugulose. 

Ventrolateral surfaces—Body scarcely swollen in basal half. 
broadly concave laterally before the apices; setose, sete rather long, 
especially upon the inferior margin of the fossa and at angle; surface 
glabrous. Internal margins of the valves contiguous in basal fourth. 
Genital fissure fusiform, widest in apical half, closed by the inferior 
pudendal membrane in basal moiety. 

It is proper to recognize two forms, as follows: 

Forma typica.—Elongate, usually large and more or less shining. 
Interstitial punctures finely muricate and moderately distant. the 
punctures laterally and on the apex more decidedly prickly, each with 
a very small stiff seta. Elytral apex gradually narrowed and slightly 


REVISION OF ELEODIINI—BLAISDELL. 283 


produced in the male; scarcely produced and obtuse in the female. 
Legs long and rather stout. 

Measurements.—M ales: Length, 24.5-34 mm.; width, 9-10 mm. 
Females: Length, 26-32 mm.; width, 10.5-10.5 mi. 

Forma minor.—Somewhat robust and more or less shining. Inter- 
stitial punctures quite distant and very distinetly prickly laterally 
and on the apex. Sexes quite homomorphic. Elytral apex scarcely 
produced and more or less abtuse. Legs moderate. 

Measurements —M ales: Length, 22.5-27 mm.; width. 8-9.5 mm. 
Females: Length, 21.5-26.5 mm.; width, 9.2-8.5 mm. 

Habitat—Forma typica.— California (San Diego County, Los 
Angeles County; collections Charles Fuchs, E. C. Van Dyke, F. E. 
Blaisdell, and U. S. National Museum; Fort Tejon, April 19-21, 
Fuchs and Hopping) ; St. Nicholas Island (collection U. S. National 
Museum). 

Forma minor.—California (Poway, elevation 700 feet, F. I. Blais- 
dell; Fort Tejon, Fuchs and Hopping: collection U. S. National 
Museum). 

Number of specimens studied, 125. 

Type in the LeConte collection. 

Type-locality.—San Diego, California. 

Salient type-characters.—Thorax broad, sides greatly rounded and 
subdepressed ; narrowed behind; basal angles rectangular, somewhat 
unusually prominent; anterior angles acuminate. Elytra finely and 
seriately punctate; apex declivously attenuated (LeConte). 

Diagnostic characters.—This race is separated from acuticauda by 
the form of the pronotum, elytral apices, and sculpturing. 

Acuticauda has the pronotum very narrowly granulate and very 
feebly impressed laterally along the margin; the disc is nearly evenly 
convex from side to side; in /aficollis the sides are quite broadly 
impressed and granulate; consequently the surface is less evenly and 
less strongly convex. 

In acuticauda the punctuation is simple, while Jaticollis is remark- 
able for having the interstitial punctures very finely muricate; on 
the elytra laterally the interstitial punctures are distinctly prickly, 


on the apex irregular and likewise muricate. 


In /aticollis the elytra are never caudately produced, although at 
times somewhat gradually narrowed, but never as strongly as in 
acuticauda., 

Dentipes forma elongata may have the pronotum somewhat im- 
pressed laterally and more or less granulate, but here the form of the 
prothorax must be the diagnostic criterion. (See aeuticauda and 
synopsis. ) 


284 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


A series of laticollis from St. Nicholas Island is almost distinctive 
of another race (forma insularis) or insular fornf. The pronotum is 
notably bright and shining, and the general form is rather more 
elongately robust. 

General observations.—The middle lobe of the mentum is moderate 
and subparabolic or subtrapezoidal in outline; the surface is very 
feebly convex and coarsely punctate, rarely feebly foveate iaterally. 

The prosternum is but moderately prominent ventrally with the 
cox, evenly arcuate antero-posteriorly and mucronate behind, or 
subhorizontal with the tip mucronate; frequently deeply and longi- 
tudinally grooved between the coxe. 

The mesosternum is more or less arcuately declivous and more or 
less broadly concave. 

The abdominal intercoxal salient is quite quadrate and in width is 
a little shorter than the third segment; the post-coxal part of the 
first is a little longer than the third; the second twice as long as the 
fourth, and the third is a third of its length shorter than the second. 

The abdominal process is about a fourth of its width wider than 
the metasternal salient. 

The metasternum laterally between the coxz is quite short and in 
length is about equal to the width of a mesotibia at middle. 

The tibial grooves of the femora are well developed, and the mar- 
gins are quite cariniform; the floors are quite flat, opaque, and fre- 
quently slightly granular, especially on the meso- and metafemora. 

The profemoral grooves are subentire, the margins slightly evanes- 
‘cent before the base, and the anterior margins are dentately laminate 
near the outer third. The mesofemoral grooves are evanescent at 
basal fourth, those of the metafemora at basal third. 

The external surface of the protibiz is more or less distinetly cari- 
nate and the tarsal grooves are more or less evident, sometimes feebly 
glabrous toward apex, but generally scabrous; the articular cavities 
are not strongly closed. 

The meso- and metatibize have the articular cavities strongly closed 
and margined; the tibie are not carinate. Externally both have the 
grooves evident in apical two-thirds, with rest of the surface very 
densely, strongly, and muricately sculptured. 

The tarsi are quite stout. The protarsi are about two-sevenths of 
their length shorter than a mesotarsus. Joints two, three, and four 
are subequal and shghtly wider than long, together about equal to the 
fifth ; first joint about as long as wide. 

The mesotarsi are about a ninth of their length shorter than a meta- 
tarsus. Joints two, three, and four are subequal and about as long as 
wide, together longer than the fifth; the first is about one and a half 
times longer than wide. 


REVISION OF ELEODIINI-——-BLAISDELL, 285 


The metatarsi are not quite equal to half of the length of a meta- 
tibia. Joints two and three are subequal, together shorter than the 
fourth and longer than the first. 


ELEODES ESCHSCHOLTZII Solier. 


Bleodes eschscholtzii Sovirre, Studi Entomol., 11, 1848, p. 254.—CHamPion, 
Biol. Centr.-Amer., IV, Pt. 1, 1884, p. 77, pl. 11, fig. 24, male. P 
Var. luca LeContr, Smithsonian Miscell. Coll., 167, Pt. 1, 1866, p. 114.— 

Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 312. 


Elongate, fusiform oval, smooth and more or less shining, strongly 
and more or less caudate. 

Head about twice as wide as long, more or less convex, feebly im- 
pressed within the sides of the frons, rather finely, more or less irregu- 
larly, and not very densely punctate. .tnfennew long and moder- 
ately slender, outer four joints feebly compressed, gradually and fee- 
bly dilated, third joint scarcely as long as the next two combined, 
fourth to the seventh inclusive subequal, eighth subtriangular, ninth 
and tenth more or less orbicular to feebly oval, eleventh ovate and 
obliquely truncate at tip. 

Pronotum quadrate, widest at the middle, an eighth to a fourth 
wider than long; dsc moderately and evenly convex, declivous at the 
apical angles, finely to obsoletely, evenly and rather sparsely punctu- 
late ; apex truncate to more or less evenly emarginate, finely margined, 
obsoletely so at the middle; sides evenly and feebly to moderately 
arcuate, finely margined; base feebly arcuate and finely beaded, and 
about a fourth wider than the apex; apical angles subrectangular to 
obtuse and not rounded, or acute-and more or less advanced; basal 
angles obtuse and not rounded. 

Propleure obsoletely punctulate and more or less rugulose. 

Elytra oval, widest at the middle; base truncate to feebly emargin- 
ate, very slightly wider than the contiguous prothoracic base; humeri 
obtuse, not rounded nor prominent; sides evenly arcuate, apex more 
or less produced; disc evenly and strongly convex from side to side, 
obliquely to arcuately and more or less gradually declivous posteri- 
orly; surface striato-punctate, punctures moderate in size, closely 
and regularly placed, in moderately distant and more or less im- 
pressed series, the intervals with a series of irregularly and more or 
less distantly spaced, very fine punctures; punctures not becoming 
confused at the sides or apex, the intervals scarcely convex, but 
becoming slightly so on the apex, where the suture is more or less 
impressed. 

Epipleure rather narrow and gradually narrowing from base to 
apex, where they extend narrowly along the sides of the inferior sur- 


286 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


face of the cauda or produced apex; surface finely and obsoletely — 
punctulate. ~ 

Sterna more or less finely, densely, obsoletely punctate and rugulose. 

Parapleure more or less finely and rather densely punctate. 

Abdomen horizontal, finely, obsoletely punctulate and rugulose, 
punctures denser on the fifth segment. 

Legs quite slender and moderately long, usually not strongly sculp- 
tured; anterior femora armed in both sexes; anterior tibial spurs 
subequal and acute; tarsi similar in the sexes. ; 

Male.—Elongate and somewhat slender. Antenne attaining a 
slight distance beyond the prothoracic base, the latter a little greater 
than the width of the prothorax. Elytra very obliquely declivous, 
then subhorizontally produced at apex into a long cauda, the latter 
frequently with the sides more or less converging to tip, which is 
slightly deflexed. Abdomen quite strongly convex, feebly, longi- 
tudinally, and narrowly impressed at middle on first three segments. 
The anterior femora armed with a moderate and acute tooth; pos- 
terior tibiz quite strongly arcuate to slightly beyond the middle, then 
rather suddenly dilated and straight to apex. 

Female.—Quite robust. Antenne not quite attaining the protho- 
racic base, the latter equal to the length. Elytra moderately broadly 
oval, arcuately declivous posteriorly, apex slightly produced and sub- 
acute. Abdomen strongly convex. Anterior femora with a more or 
less obtuse tooth, and the metatibiw straight without being widened 
apically. 

Measurements. 


Males: Length, 27-29 mm.; width, 9-9 mm. Fe- 
males: Length, 24-25.5 mm.; width, 9-9.5 mm. 

Genital characters, male-—Edeagophore elongately subfusiform 
and feebly arched. 


Basale more or less oblong, evenly convex above, and the sides 
evenly and moderately arcuate. 

Apicale elongately triangular and very feebly decurved; surface 
moderately and evenly convex above, with a narrow median longi- 
tudinal groove; sides feebly arcuate or straight; apex not produced 
and narrowly rounded; base acutely lobed at middle, broadly and 
feebly sinuate laterally. 

Sternite subparabolic in outline. Each lobe triangular; external 
border more or less evenly arcuate; apex rather broadly rounded; in- 
ternal margin more or less sinuate; surface glabrous, rather evenly, 
moderately densely punctate and setose in apical three-fourths, sete 
moderate in length, becoming long on the apex. Membrane not 
setose across the bottom of the sinus, the latter rather triangular. 

Female.—Genital segment robust, subquadrate and strongly chiti- 
nized, 


REVISION OF ELEODIINI-——BLAISDELL., 287 


Valvula.-Dorsal plate more or less oblong, glabrous and shining, 
sides very strongly reflexed; surface deeply concave, very finely, 
sparsely punctate and finely setose; external border rather strongly 
arcuate; internal margin more or less sinuous; apical margin promi- 
nent, evenly and moderately arcuate, angle rounded. Apex short 
and triangular; valvular membranes strongly exposed between the 
apices caudad to the pudendal membrane. 

Appendage subsemielliptical, rather equal in length and size to 
the apex. Appendage, apex, and margin of the fossa finely punctate 
and setose, setze short and inconspicuous. 

Superior pudendal membrane attaining the middle of the dorsal 
plate and longitudinally rugulose. 

Ventrolateral surfaces —Body moderately inflated and glabrous, 
transversely and broadly concave before the apices, the latter finely 
punctate and setose. Submarginal groove more or less obsolete. In- 
ternal margins of the valves contiguous in basal third; genital fissure 
narrowly fusiform and closed in basal half by the visible inferior 
pudendal membrane. 

Habitat—Mexico (Sinaloa, coll. Chas. Fuchs; Alamos, Buchau- 
Hepburn; Presidio, Forrer). United States (Texas). 

Number of species studied, 5 (87). 

Type is probably with the Solier collection. 

Type-locality.—Mexico (Alamos?). 

Salient type-characters.—Thorax quadrate; dise very finely punc- 
tulate; apex feebly emarginate; apical angles subacute and slightly 
prominent anteriorly; sides evenly but not strongly arcuate; basal 
angles obtuse. Elytra striato-punctate, punctures moderate and 
closely placed in the series, intervals scarcely convex and with a 
series of rather distantly spaced punctules; cauda horizontal. 

Diagnostic characters—The figure given in the Biologia I take 
to represent what is probably a typical form of this species. I can 
not see that it differs very much from some of the less strongly de- 
veloped forms of /ucw. The prothorax is quadrate, with the sides 
feebly and evenly rounded from base to apex; apical margin feebly 
emarginate, angles small, acute, and just feebly prominent anteriorly. 

In the specimens before me the sides of the prothorax are rather 
less rounded, apical margin very feebly emarginate or truncate and 
the apical angles scarcely at all prominent. The general form of the 
body is rather more slender, with outline more evenly fusiform. The 
cauda narrows somewhat to tip. 

I do not believe that a large series could be separated from a simi- 
Jarly large series of luce. 

In a series of five examples (4 males and 1 female) mentioned 
in the Biologia, the largest male measures 35 mm., including the 


288 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


caudal prolongation of 6 mm.; similar large males are common in 
the series of Zucw taken in Lower California. The hind tibie are 
curved in basal half, straight and thickened toward apex as in luce. 

General observations—The middle lobe of the mentum in the ex- 
amples at hand is triangulo-trapezoidal, more or less arcuately 
rounded at apex; surface is coarsely punctate, just noticeably convex 
and scarcely at all foveate iaterally. 

The prosternum in the series under examination is rounded antero- 
posteriorly, feebly mucronate or not. In the larger examples it 
would no doubt be strongly mucronate as in ducw. In a large male 
without locality and in the U. S. National Museum collection, the 
prosternum is horizontally and triangularly mucronate, exactly as in 
the large specimens of /ucw,; the specimen measures 33 mm. Two 
accompanying females have similar prosterna. These specimens are 
without doubt from Mexico. . 

The mesosternum is the same as in ducw, except that it is less ver- 
tically oblique. 

The abdominal process is subquadrate and feebly transverse, in 
width equal to the length of the third segment; the post-coxal por- 
tion of the first segment is equal in length to that of the second; 
the third is scarcely twice the length of the fourth and about a third 
of its length shorter than the second. 

The abdominal process is about a fourth of its width greater than 
that of the metasternal salient. 

The metasternum laterally between the cox is about equal in 
length to the width of a mesotibia at apex. 

The tibial grooves of the femora are well defined, margins carini- 
form, floors plane, opaque and smooth. Those of the profemora 
nearly attain the femoral base and the margins become contiguous. 
The margins of the grooves on the mesofemora become evanescent 
at the internal fourth before becoming contiguous. On the meta- 
femora the margins are evanescent at internal third. The meso- and 
metafemora have the grooves somewhat narrow. On the profemora 
the anterior margin of each groove is dentately laminate at external 
fourth. . 

The protibie are more or less carinate externally and not com- 
pressed; the articular cavities are more or less open, and the tarsal 
grooves are more or less developed and scabrous. 

The meso- and metatibiz are not carinate externally and the artic- 
ular cavities are closed. The tarsal grooves of the mesotibize are 
quite well defined in apical two-thirds; the sides of the grooves are 
muricate and the floors smooth and opaque. The metatibiz are sim- 
ply flattened externally or feebly grooved and muricate. 

The tarsi are moderate in length and stoutness. 


REVISION OF ELEODITNI—BLAISDELL, 289 


The protarsi are about a third of their length shorter than a meso- 
tarsus. Joints two, three, and four subequal, apparently very slightly 
wider than long, and together about equal to the fifth; the first about 
as long as wide. 

The mesotarsi are very slightly shorter than a metatarsus. Joints 
two, three, and four subequal, scarcely longer than wide, and together 
just a little longer than the fifth; the first about one and a half times 
longer than wide. 

The metatarsi are a little shorter than half of the length of a meta- 
tibia. Joints two and three subequal, distinctly longer than wide 
and together scarcely as long as the fifth, a little longer than the first. 


ELEODES ESCHSCHOLZII var. LUCA LeConte. 


Elongate, more or less shining, convex and more or less caudate. 

Head twice as wide as long, moderately and evenly convex, rather 
finely, quite evenly and not densely punctate. Antenne long and 
slender, subequal in length in the sexes, outer four joints feebly com- 
pressed and slightly dilated, third joint subequal in length to the 
next two combined, fourth very slightly longer than the fifth, the 
latter to the seventh inclusive subequal, eighth a little shorter and 
obeonical, ninth and tenth scarcely wider than long, eleventh more 
or less ovate. 

Pronotum more or less quadrate, widest at the middle, usually 
about a sixth to a third wider than long, and less than twice as wide 
as the head; disc smooth, moderately and evenly convex, slightly 
declivous at the apical angles, very finely, evenly and sparsely, or 
obsoletely punctulate; apex deeply emarginate, finely or obsoletely 
margined; sides evenly and not strongly arcuate, very finely mar- 
gined; base feebly arcuate, finely beaded, and about a sixth wider 
than the apex; apical angles porrect and acute, at times dentiform 
and more or less everted; basal angles obtuse. 

Propleure smooth, more or less rugulose and finely punctulate 
anteriorly. ‘ 

Elytra more or less oval, smooth, usually widest at the middle; 
base subtruncate to feebly emarginate and a little wider than the 
contiguous prothoracice base; humeri obtuse, not rounded nor promi- 
nent; sides evenly arcuate, apex more or less caudate; dise evenly 
convex and punctato-striate, strie slightly impressed with the punc- 
tures rather coarse and closely placed, intervals scarcely convex with 
a series of extremely fine punctules that are more or less irregularly 
placed, not becoming asperate at the sides and apex. Cauda slightly 
deflexed at tip, above with the elytral suture impressed, beneath con- 
cave, the concavity being formed by the general internal surface of 
the elytra. 

59780—Bull. 68—09——19 


290 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Epipleure quite wide at the humeri, thence gradually and rather 
rapidly narrowing for a short distance, then less rapidly to apex, 
where they narrowly margin the sides of the inferior surface of the 
cauda; surface smooth, generally impunctate. 

Sterna more or less punctate and rugulose. 

Parapleure not strongly punctate. 

Abdomen horizontal, finely, sparsely, quite evenly punctate and 
more or less rugulose. 

Legs rather long and quite slender; anterior femora armed with a 
straight, acute tooth in both sexes; tibial spurs similar in the sexes 
and rather small, the anterior being equal and acute. Tarsi similar 
in the sexes. 

Male.—Elongate and dimorphic. Pronotum nearly quadrate, about 
a sixth to less than a third wider than long. Elytra gradually de- 
clivous posteriorly; apex briefly produced or strongly and horizon- 
tally caudate, and accordingly two forms may be recognized: 

Forma typica with the cauda equal to about a fourth of the entire 
length of the elytra. Abdominal segments one to three usually 
broadly and longitudinally impressed; posterior tibize quite strongly 
arcuate in basal half, thence to apex straight and distinctly widened. 

Forma ecaudata with the elytral apex simply obliquely and briefly 
produced, acute at tip. Abdomen evenly convex from side to side; 
posterior tibie not arcuate and not more suddenly widened toward 
apex. 

Female.—Robust. Pronotum slightly transverse and about a 
fourth to a third wider than long. Elytra more or less broadly oval 
and slightly ventricose, quite strongly, suddenly and obliquely decliy- 
ous posteriorly; apex simply acute. Abdomen evenly convex. Pos- 
terior tibie straight. 

Genital characters, male—Edeagophore oblong-ovate, rather small 
and slightly arched. 

Basale evenly convex, oblong, with the sides subparallel, and about 
twice as long 4s the apicale. 

Apicale longer than wide at base, triangular ; surface evenly con- 
vex, with a very fine median groove extending from near the base to 
the apex, where it becomes wider; sides nearly straight or feebly 
arcuate; apex acute and not produced; base subacutely lobed at the 
middle and sinuate laterally. 

Sternite truncately subtriangular. Each lobe with the external 
border straight in basal two-thirds converging toward apex, thence 
evenly arcuate to angle; internal margin rather arcuate, apex nar- 
rowly rounded; surface rather strongly and densely” punctate 
throughout, setose, sete quite long and dense at tip. Membrane not 
setose across the bottom of the sinus, the latter rather broad. 


REVISION OF ELEODIINI—BLAISDELL. 291 


Female.—Genital segment subquadrate, not strongly developed at 
base, moderate in size, and setose. 

Valvula (Plate 1, fig. 21).—Dorsal plate, oblong, with sides more or 
less reflexed; surface moderately coneave, finely and sparsely punc- 
tate, and with rather short sete ; external border moderately arcuate ; 
internal margin quite straight and parallel; apical margin subtrun- 
cate and fringed with a few very short hairs; angle subrectangular 
and without conspicuous sete. Apex short, stout, and triangular, 
with few very short hairs at tip. 

Appendage moderately large, as long as the apex, flattened, semi- 
elliptical, and broadly rounded at tip, with two or three long sete, 
others short and scattered over the surface. Fossa moderate in size 
and not conspicuously fringed with sete, situated beneath the ex- 
ternal half of the apical margin of the dorsal plate. 

Superior pudendal membrane attaining about the middle of the 
dorsal plate and longitudinally rugulose. Valvular membranes visi- 
ble between the apices caudad. 

Ventrolateral surfaces—Body slightly swollen at base, moder- 
ately and transversely concave before the apex, the latter covered 
with short sete. Submarginal groove normal and linear, more or 
less visible from above, especially in dried specimens; lateral plate 
not visible externally when viewed from above. Internal margins of 
the valves contiguous for a short distance at base; genital fissure long 
and fusiform, closed in basal half by the inferior pudendal mem- 
brane. 

The three following forms are to be recognized: 

Forma typica.—Moderate in size. Males more or less caudate. 
Elytra striato-punctate and not noticeably inflated. 

Measurements—Males: Caudate form: Length, 20-28 mm; with 
cauda, 23-31.5 mm.; width, 8-10.5 mm. Ecaudate form: Length, 
24-28 mm.; width, 9-9.5 mm. Females: Length, 22-27 mm.; width 
9.2-11.5 mm. 

Forma inflata—Quite robust. Elytra striato-punctate and more 
or less distinctly inflated (Plate 1, fig. 22). 

Measurements.—Males: Not at hand. Females: Length, 26.5-32 
mm.; width, 11.5-13.2 mm. 

Forma grandis.—Very robust and elongate, smooth, body very 
evenly and gradually narrowed behind. Elytral striz more or less 
obsolete. 

Measurements —Male: Length, 37 mm.; width, 13.5 mm. 

Habitat—Lower California. 

Forma typica.—Cape ‘St. Lucas; Santa Rosa, Gustav Beyer; San 
Jose del Cabo, collections of Charles Fuchs and California Academy 
of Sciences 


292 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Forma inflata.—San Jose del Cabo, collection Charles Fuchs. 

Forma grandis.—Calmalli mines, collection California Academy 
of Sciences. 

Number of specimens studied, 140. 

Type in the LeConte collection. 

Type-locality—Cape St. Lucas, Lower California. 

Salient type-characters——Thorax quadrate, apex deeply emargi- 
nate, apical angles porrect and acute, but not acuminate; sides mod- 
erately rounded, basal angles obtuse; disc moderately convex and 
sparsely punctulate. Elytra oval and convex, acutely attenuate be- 
hind, strongly striato-punctate, strize slightly impressed, intervals 
punctulate (LeConte). 

Diagnostic characters.—The characters that separate this race from 
eschscholtzii are very feeble. In duce the apical angles of the pro- 
thorax are more strongly prominent anteriorly and the apical mar- 
gin more strongly emarginate. The prothorax is less typically quad- 
rate and the cauda is less attenuated toward tip. From the differ- 
ences presented by the series before me, I have no doubt but that a 
larger series from the different geographical regions would prove 
these differential characters of no value. 

In the strongly developed males with long caude the hind tibie 
are more or less arcuate in the basal half and rather abruptly dilated 
in the apical moiety, these characters are not evident in the ecaudate 
form. 

From the forms of acuticauda it is separated by the striato-punc- 
tate elytra, by the sides of the pronotum being quite feebly rounded, 
and by the elytral apices being more abruptly attenuate, the cauda 
being horizontal. 

From dentipes it is readily known by the quadrate prothorax with 
feebly rounded sides. 

Forma inflata has to be separated from ventricosa by the sides of 
the pronotum being less convergent anteriorly and by the greater 
width of the apex as compared with the base; the elytra are not so 
decidedly inflated and the apices are not produced in the females. 
This form appears to lead toward ventricosa—in fact may be inter- 
mediate, but undoubtedly has great affinity with duce. A large 
series of both species would no doubt demonstrate this relationship. 
The males are probably ecaudate. 

Forma grandis is a wonderful specimen of gigantism—such phe- 
nomenal development makes the limiting of species exceedingly dif_i- 
cult in the absence of an ample series. The epipleurw are wide at 
the humeri and rather rapidly narrowing to opposite the meta- 
sternal epimera, thence narrower, with margins gradually converging 
to apex. Fourteen series of very fine punctules are clearly indicated 
upon each elytron. 


REVISION OF ELEODITNI—BLAISDELL. 293 


General observations.—The middle lobe of the mentum is variable, 
but nearly always triangular, in the large examples quite equilaterally 
so; the apex is scarcely truncate, generally distinctly rounded; sur- 
face more or less irregular from the broad, shallow, lateral foveee. 

The prosterna in the series before me are exceedingly variable. In 
the more strongly developed examples the prosternum is horizontal 
and triangularly, strongly mucronate; in the smaller specimens it is 
rounded antero-posteriorly and without a mucro, at other times it is 
more or less mucronate behind and more or less convex between the 
coxe. The prosternum is not strongly prominent ventrally with the 
COX. 

The prosternum in forma grandis has an extraordinary develop- 
ment. It is horizontal between the coxe and uncinately produced 
and slightly but evenly decurved, arcuately sinuate above the mucro. 

The mesosternum is more or less oblique and coneave. 

The abdominal process is subquadrate, slightly transverse, with its 
width about equal to the length of the post-coxal part of the first 
segment, and also to the length of the third; the second is about twice 
the length of the fourth. 

The abdominal salient is about a fourth of its width broader than 
the metasternal process. 

The metasternum laterally between the coxe is about as long as the 
width of a mesotibia at middle. 

The tibial grooves of the femora are well developed, with the 
margins more or less distinctly cariniform. The profemoral grooves 
are moderately wide, with the sides feebly arcuate or quite straight. 
the latter becoming contiguous at the femoral base; the anterior 
margins are quite acutely dentately laminate at the outer third; the 
floors are nearly flat, opaque, and smooth. 

The mesofemoral grooves have the sides evanescent at about basal 
fifth, while those of the metafemora become evanescent at basal 
fourth. The floors of the grooves on both are flat and glabrous. 

The protibix are feebly compressed and the mesotibiz just notice- 
ably so. The protibix are distinctly carinate externally; the tarsal 
grooves are more or less well marked and scabrous; the articular 
cavities are nearly always closed. 

The metatibiew are variable in form and carinate to a slight extent 
the grooves are more or less well marked and at times rather deep, 
subglabrous, and opaque; the articular cavities are closed. 

The metatibiw are variable in form and carinate to a slight extent 
externally in basal fourth; the grooves are more or less obsolete or 
feeble and the articular cavities are closed. 

The tarsi are more or less variable in length and thickness; usually 
moderate. The protarsi are about a third of their length shorter 
than a mesotarsus. Joints two, three, and four are subequal, the 


994 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


fourth is at times very slightly longer than the second, all are wider 
than long and together a little shorter than the fifth} the first is about 
as long as wide. 

The mesotarsi are about an eighth of their length shorter than a 
metatarsus. Joints two, three, and four are subequal, scarcely longer 
than wide and together slightly longer than the fifth; the first is 
about one and a half times longer than wide. 

The metatarsi are slightly shorter than half the length of a meta- 
tibia. Joints two and three are subequal and a little longer than 
wide, together scarcely as long as the fourth, and noticeably longer 
than the first. 

ELEODES TENUIPES Casey. 


Eleodes tenuipes Casky, Ann, N. Y. Acad. Sci., V. Nov., 1890, p. 399. 


Elongate-ovate, rather smooth and shining. Caudate. 

Head twice as wide as long, finely and sparsely punctate, very 
slightly convex. Antenne moderate, scarcely compressed and not 
dilated in outer four joints, third joint a little longer than the next 
two taken together, fourth just a trifle longer than the fifth, the lat- 
ter to the eighth, inclusive, subequal, eighth subtriangular, ninth and 
tenth suborbicular, eleventh ovate. 

Pronotum quite quadrate, about a seventh wider than long, widest 
just at or a little in advance of the middle; disc evenly and moder- 
ately convex, minutely and very sparsely punctulate; apex quite 
truncate, more or less finely and obsoletely margined; sides quite 
evenly and not strongly arcuate from base to apex, finely beaded ; base 
broadly and very feebly arcuate, about equal to the apex and about 
a sixth shorter than the length; apical angles very acute, small, denti- 
form, and everted ; basal angles obtuse, feebly or scarcely rounded. 

Propleure quite smooth externally, more or less obsoletely punctate 
and rugulose. 

Elytra elongate oval, widest at the middle; base feebly emarginate, 
and equal to the contiguous prothoracic base; humeri obtuse, not in 
the least prominent; s/des evenly arcuate, apex more or less pro- 
duced ; disc moderately convex on the dorsum, quite evenly rounded 
laterally, arcuately and obliquely declivous behind, punctate, pune- 
tures arranged in distant unimpressed rows of fine, simple, moder- 
ately approximate punctures, the intervals each with a single series 
of still finer and widely spaced punctures, which are generally simple, 
but which laterally toward apex become very coarse sparse asperities. 

E'pipleure moderate in width beneath the humeri, thence gradually 
narrowing to apex, where it narrowly margins externally the infe- 
rior surface of the elytral prolongation; surface smooth, sparsely and 
very finely punctate toward apex. 


OC —————— ——— 


REVISION OF ELEODITINI—BLAISDELL. 295 


Sterna more or less obsoletely punctate and very feebly rugulose. 

Parapleure coarsely and somewhat obsoletely punctate. 

Abdomen horizontal, quite smooth, very finely and sparsely punc- 
tulate, finely rugulose. 

Legs rather long and slender. Anterior femora more or less 
armed; anterior tibial spurs subequal and rather stout. Tarsi similar 
in the sexes, with the spinules beneath quite slender. 

Male-—Body moderately slender, antennx attaining the base of the 
prothorax. Elytra produced at tip forming a cauda, which is nearly 
horizontal and 4 mm. more or less in length, with its inferior surface 
formed by the general internal surface of the elytra. Abdomen 
moderately convex, more or less feebly impressed along the median 
line on the first three segments. Anterior femora armed with a 
slender and acute tooth at the outer third. 

Measurements.—M ales: Length, with cauda, 30.5-31 mm.; width, 
10-10.8 mm. Females: Unknown to me. 

Genital characters, male—Edeagophore elongately fusiform, not 
arched. 

Basale elongate oblong, very moderately convex, with sides slightly 
arcuate. 

Apicale elongate, triangular, nearly twice as long as wide, mod- 
erately convex above, groove very fine and linear; sides feebly 
arcuate, just noticeably sinuate in apical half, tip subacute; base with 
a broad rounded lobe at central two-fourths, laterally feebly sinuate. 

Sternite transversely parabolic. Each lobe rather short, with the 
external border evenly arcuate and continuously so with the apical 
border, which is rather broadly arcuate, angle not evident; internal 
border short, feebly arcuate or straight; surface feebly convex, 
sparsely punctate and setose, setee quite long, longer and denser 
on the apical margin, impunctate in basal fourth. Membrane not 
setose across the bottom of the sinus, the latter rather broad. 

Habitat—Texas (El Paso). 

Number of specimens studied, 1 male; type not seen. 

Type a male in the collection of Colonel Casey. Collector, G. W. 
Dunn. 

Type-locality.—E1 Paso, Texas. 

Salient type-characters.—Prothorax with the apex very nearly as 
wide as the base, broadly, feebly emarginate in circular are, the 
apical angles very acute and in the form of small everted teeth; base 
very feebly arcuate; basal angles extremely obtuse; sides almost 
evenly and distinctly arcuate; disc widest just visibly before the 
middle, rather strongly and evenly convex, minutely and very 
sparsely punctate. Elytra with humeri not prominent; dise with 
distant unimpressed rows of fine simple moderately approximate 
punctures, the intervals each with a single line of still finer and ex- 


296 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


tremely widely spaced punctures, which are generally simple, but 
which laterally toward apex become very coars@ sparse asperities 
(Casey). 

Diagnostic characters —In tenuipes the apical angles of the pro- 
thorax are strongly everted, the sides are quite evenly arcuate; in 
wickhami the apical angles are not everted and the sides are oblique 
posteriorly; in fenuipes and in a specimen received from Professor 
Wickham—the latter I refer to wickhhami—the elytra are similar 
and more elongately oval than in ducw. In the specimen of tenuipes 
before me the elytra are very slightly widened behind the middle and 
this character is less evident in Wickham’s specimen. In both the 
length of the prothorax is comparatively greater as compared to the 
width than in duce; the latter when well developed have the apical 
angles of the prothorax acute and prominent anteriorly, with the 
apex distinctly and evenly emarginate, and in the ecaudate form these 
characters are less strongly marked. Professor Wickham’s specimen 
is of interest, combining the characters of fenuipes and wickhami. 

Some specimens of ducw have the apical angles of the prothorax 
more or less strongly everted. 

In the specimens of eschscholtzii before me the prothoracic apex 
is quite truncate and the angles are subacute and not prominent an- 
teriorly. 

General observations.—The middle lobe of the mentum is moderate 
in size, with the apex arcuate, the surface is moderately coarsely 
punctate and feebly scabrous, not noticeably setose. 

The prosternum is arcuately convex between the coxe, and with 
them protuberant ventrally. It is not mucronate nor longitudinally 
grooved in the specimen before me. 

The mesosternum is arcuately and vertically declivous, broadly and 
moderately concave. , 

The abdominal intercoxal salient is quadrate and about a third of 
its length shorter than the post-coxal portion of the segment, the lat- 
ter being slightly shorter than the second, which is about a fourth 
longer than the third, the latter being less than twice as long as the 
fourth. 

The abdominal salient is about a sixth of its width wider than the 
metasternal process. 

The metasternum laterally between the coxve is about as long as the 
width of a mesofemur at base. 

The tibial grooves of the femora are well defined by cariniform 
margins, the floors are glabrous and shining, and nearly plane. Those 
of the profemora are moderately wide and nearly attain the femoral 
base; the margins are scarcely arcuate and gradually converge to the 
base, where they become contiguous; the anterior margins are acutely 
and dentately laminate at the apical fourth. On the mesofemora the 


REVISION OF ELEODIINI—BLAISDELL. 297 


grooves are narrower and gradually converge to become evanescent 
at basal fourth without becoming contiguous. 

The metafemora have the grooves moderately narrow and evanes- 
cent at basal third, where the margins are distant. 

The protibix are slightly compressed and distinctly carinate exter- 
nally in basal two-thirds; the tarsal grooves are not strongly devel- 
oped and limited posteriorly by a series of rather coarse and sparse 
denticulations, and the floors are subscabrous; the articular cavities 
are more or less closed. 

The mesotibiw are scarcely compressed and feebly carinate exter- 
nally in basal fourth; the tarsal grooves are rather well developed, 
rather broad apically, gradually narrowing toward base, and their 
floors are subglabrous, being limited by muricate margins; articular 
cavities closed. 

The metatibix are somewhat rounded, obsoletely carinate in basal 
third externally, thence to apex more or less flattened and concave; 
surface of the grooves more or less shining and sparsely subasperate ; 
articular cavities closed. 

The tarsi in the specimen at hand are moderately long and rather 
stout. 

The protarsi are about a third of their length shorter than a meso- 
tarsus. Joints two, three, and four are subequal, wider than long, 
and together equal to the fifth; the first is about as long as wide. 

The mesotarsi are subequal in length to a metatarsus. Joints two 
and three about as wide as long, the fourth scarcely as wide as long, 
apparently very slightly decreasing in length from the second to the 
fourth, together subequal to the fifth; the first is a little longer than 
wide. 

The metatarsi are about four-tenths as long as a metatibia. Joints 
two and three about equal in length and size, a little longer than wide 
and together scarcely as long as the fourth; the first is about one and 
a half times longer than wide. 


ELEODES WICKHAMI Horn. 
Eleodes wickhami Horn, Trans. Amer, Ent. Soc., XVIII, Feb., 1891, p. 41, pl. 
I, fig. 12. 

Elongate, more or less shining and caudate, 

Head twice as wide as long, rather opaque, feebly convex, finely 
and very sparsely punctate, frontal suture distinct. Antenne rather 
long, reaching to or slightly beyond the prothoracie base, not com- 
pressed nor dilated in outer three joints, third joint a little longer 
than the next two taken.together, fourth very little longer than the 
fifth, the latter to the seventh inclusive subequal, eighth slightly 
shorter and a little more robust, ninth and tenth orbicular, éleventh 
ovate, 


298 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Pronotum subquadrate, one-sixth wider than long, slightly nar- 
rowed behind, widest at or just in advance of the middle; dise mod- 
erately convex, sparsely, very finely and indistinctly punctulate; 
apex slightly emarginate to subtruncate, finely and more or less ob- 
soletely margined; sides slightly and evenly arcuate anteriorly, and 
more or less oblique posteriorly, sometimes slightly sinuate before the 
basal angles, finely margined; base feebly arcuate and finely beaded, 
about one-seventh wider than the apex; apical angles more or less 
distinct, acute, not everted although sometimes feebly reflexed; basal 
angles obtuse, not rounded. 

Propleurw smooth, obsoletely punctate and more or less rugulose. 

Elytra elongate oval, widest at the middle; base feebly emarginate, 
very slightly wider than the contiguous prothoracic base; humeri 
obtuse, not prominent nor rounded; sides evenly arcuate or subpar- 
allel, attenuate posteriorly and arcuately declivous with the apex 
more or less caudate; dise slightly and evenly convex on the dorsum, 
more strongly so and broadly rounded laterally, punctate, punctures 
fine and more or less indistinct and arranged in series, sometimes 
feebly impressed, intervals with a single row of fine, widely spaced 
punctures, which are generally simple, but may become small asperi- 
ties on sides about apex. 

Epipleure moderate in width beneath the humeri and gradually 
narrowing to the apex; surface smooth and more or less obsoletely 
punctulate. 

Sterna rather smooth and more or less obsoletely punctate and 
rugose. 

Parapleure more or less coarsely and obsoletely punctate. 

Abdomen horizontal, somewhat shining, indistinctly, sparsely 
punctulate, and more or less transversely rugulose. 

Legs rather long and moderate in thickness; anterior femora 
armed in one sex at least; anterior tibial spurs subequal and rather 
stout. Tarsi similar in the sexes with the spinules beneath moder- 
ately stout. 

Male.—Body slightly robust. Elytra arcuately and obliquely de- 
clivous behind, prolonged into a narrow, horizontal cauda, whose 
under surface is a part of the general internal surface of the elytra, 
the epipleure passing narrowly along the sides to tip, the latter 
slightly decurved. Abdomen feebly and broadly impressed at middle, 
especially on the second segment. Anterior femora with an acute and 
rather stout tooth at outer fourth; anterior tibiz more or less con- 
stricted at base. Middle and posterior femora simple; posterior 
tibie frequently slightly arcuate and narrow in basal half, generally 
more or less abruptly dilated in apical moiety; both the middle and 
hind tibiz generally with the surface quite asperate in apical half. 


REVISION OF ELEODITNI—BLAISDELL. 299 


Measurements.—Males: Length, 27-34 mm.; including cauda, 
33-39 mm.; width, 12 mm. Females: Unknown to me. 

Genital characters, male-—Edeagophore elongate oblong oval, not 
slender. 

Basale oblong, about twice as long as wide, moderately convex, 
sides subparallel. 

Apicale rather broadly triangular, a little longer than wide, feebly 
convex, without median groove; sides nearly straight to tip, the latter 
acute; base with a rather narrowly rounded lobe at middle third, 
laterally broadly and feebly sinuate. 

Sternite as in tenuipes. 

Habitat—Arizona (Tueson, H. F. Wickham); New Mexico (Las 
Cruces). 

Number of specimens studied, 1 male. 

Type, a male in the Horn collection. 

Type-locality—Tueson, Arizona; collector, H. F. Wickham. 

Salient type-characters—Thorax subquadrate, a little wider than 
long, slightly narrowed behind, sides slightly arcuate in front, 
oblique posteriorly, anterior angles not everted; disc moderately con- 
vex, sparsely very finely and indistinctly punctate, elytra elongate 
oval, attenuate posteriorly; dise convex with rows of extremely fine 
indistinct punctures (Horn). 

Diagnostic characters—(See tenuipes.) Without doubt, to my 
mind the two species just described are extreme forms (heterotypes) 
of the same species, but on account of the paucity of material known 
to me, I have deemed it best to retain them separate until a larger 
series shall have been collected. 

At most wickhami can only be a race of tenuipes, Casey’s name 
having priority of three months (fenuipes, November, 1890; wick- 
hami, February, 1891). Unfortunately only males are known to me. 

Both forms have been compared with /uc@ by their authors. They 
do show greater affinity to that species than to any other, except 
gracilis. From luce they can be recognized by the elytra being sculp- 
tured with rows of fine punctures, and not strongly striate nor with 
the intervals at all convex; the elytra are slightly more inflated and 
more suddenly declivous behind. 

The prothoracie characters are too variable to be relied upon. By 
direct comparison I can not say that the legs are always longer than 
in luce. 

The form of the hind tibiz in wickhami is not peculiar to that 
species, as the same character is seen in the males of a number of 
other species, notably in the caudate form of /uce. It is strange 
that such characters should have been overlooked or ignored by pre- 
vious writers. 


300 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


From gracilis the two species under consideration differ in the 
caudate elytra of the males and in having the prothorax slightly 
longer. 

The prothorax in ¢enuipes is indeed almost as in the typical gra- 
cilis. The type of tenuipes and wickhami were practically collected 
in the same geographical region, the latter extending into New Mex- 
ico (Wickham’s specimen) ; gracilis occurs in the same area. 

My specimen of tenvipes was obtained from Mr. G. W. Dunn, who 
collected it at El Paso, Texas; it was probably one of the same lot 
from which Colonel Casey obtained his type. 

General observations—The following analytical remarks are 
founded upon Professor Wickham’s New Mexico specimen: 

Mentum moderate in size and triangular, with apex rounded; sur- 
face very feebly convex and quite coarsely punctate. 

The pro- and mesoternal characters are identical, as are also the 
tarsal characters, with those of tenuipes. 

The abdominal and metasternal salients and abdominal segments 
do not differ relatively from those of tenwipes. 

The metasternum laterally between the coxe is about as long as the 
width of a mesofemur at base. 

The femoral and tibial characters are the same as in tenuipes, ex- 
cept that the protibie have the articular cavities widely open. The 
metatibiz are just noticeably arcuate in basal half and gradually 
but distinctly dilated toward apex. 


ELEODES VENTRICOSA LeConte. 


Eleodes ventricosa LECONTE, Proc. Acad. Nat. Sci. Phila., 1858, p. 186.— 
Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 311.—CHAMPION, Biol. 
Centr. Amer., IV, Pt. 1, 1884, p. 78. 

Fusiform ovate, robust, elytra usually strongly inflated, smooth, 
and more or less shining. 

Head twice as wide as long, more or less moderately convex, fre- 
quently slightly and transversely impressed between the eyes, with 
the vertex more strongly convex, frontal suture distinct, finely, 
sparsely, and irregularly punctate. Antenne moderately long, at- 
taining a slight distance beyond the prothoracic base, comparatively 
slender, very feebly compressed, and slightly clavate; third joint 
about equal to the next two combined, fourth a little longer than the 
fifth, the latter to the seventh, inclusive, subequal in length, eighth 
shorter and slightly subtriangular, ninth and tenth suborbicular, 
eleventh ovate. 

Pronotum slightly transverse, widest at the middle, one-fourth to 
one-third wider than long, narrowing moderately anteriorly, slightly 
posteriorly; disc quite evenly and moderately convex, finely and 
sparsely punctate; apew slightly emarginate, finely and more or less 


REVISION OF ELEODIINI—BLAISDELL. 301 


obsoletely margined; sides moderately arcuate from base to apex, 
finely margined, the margin frequently more or less narrowly re- 
flexed; base moderately arcuate and rather coarsely margined, about 
one-third wider than the apex and distinctly greater than the length; 
apical angles distinct and more or less prominent anteriorly, gener- 
ally acute and not dentiform; basal angles distinctly rounded, fre- 
quently subobtuse. 

Propleure sparsely punctate and more or less rugulose, frequently 
smooth externally. 

Elytra broadly oval and usually strongly inflated, always less 
than twice as long as wide, widest at the middle; base moderately 
emarginate, usually as wide as the contiguous prothoracic base; 
humeri obtuse, not prominent nor rounded; sides evenly and quite 
strongly arcuate, apex more or less produced; disc quite evenly and 
strongly convex from side to side, at times more or less feebly de- 
pressed on the dorsum, and more strongly rounded laterally, evenly 
and arcuately declivous posteriorly; surface punctate and feebly sul- 
cate, punctures more or less coarse and arranged in moderately dis- 


tant series, intervals at times feebly convex, at others flat, with a 


single series of fine punctures, which are more or less regularby or 
irregularly arranged; laterally and on the apex the punctures be- 
come much coarser and the intervals at times strongly convex. 

Epipleure moderately narrow at base and very gradually narrow- 
ing to apex, superior margin near base broadly and feebly sinuate; 
surface smooth, very finely and distinctly or obsoletely, sparsely 
punctate. 

Sterna more or less shining, moderately densely punctured and 
rugose. 

Parapleure more or less coarsely punctate. 

Abdomen horizontal, moderately convex, shining, and glabrous, 
very sparsely punctulate and more or less rugulose. 

Legs rather long and moderate in thickness. Anterior femora 
armed in one sex. Anterior tibial spurs similar in the sexes, the 
anterior slightly longer than the posterior, both acute. Tarsi nearly 
similar in the sexes; each joint, except the last, of all the tarsi with a 
slender and acute tuft of golden-yellow and somewhat modified 
spines on apical angles beneath. (See male.) 

Male—Elytra rather gradually and arcuately declivous behind 
and produced at tip into a moderately long cauda, which is not hori- 
zontal; elytral sides and dorsum feebly and broadly sinuate before 
the produced apex, the latter with its inferior surface a part of the 
general internal elytral.surface and narrowly margined at the sides 
by the epipleure. Abdomen more or less broadly impressed in the 
median line on first three segments. Anterior femora armed with a 
rather long, acute, and rather strongly curved tooth at outer fourth. 


302 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


First joint of the anterior tarsi with the tip slightly and transversely 
produced beneath, the same bearing a tuft of mo@ified spines which 
obstructs the groove. 

Female.—Elytra quite evenly arcuately declivous behind, with the 
apex feebly produced or simply subacute. Abdomen not impressed. 
Anterior femora more or less sinuate in outer fourth. 

Measurements —Males: Length, 30-33 mm. (with eauda) ; width, 
13-14 mm. Females; Length, 26-33.2 mm; width, 13.5-15 mm. 

Genital characters, male-——Edeagophore elongately oval, depressed 
and not arched. 

Basale oblong, very moderately and evenly convex, sides feebly 
arcuate. 

Apicale triangular, very slightly longer than wide and rather 
broad; surface rather feebly convex and not grooved; sides evenly 
and not strongly arcuate to tip, the latter subacute; base with a sub- 
triangular lobe at middle, which is narrowly rounded at tip, feebly 
sinuate laterally. 

Sternite transversely parabolic. Each lobe with the external mar- 
gin evenly arcuate to apex, the latter somewhat broadly rounded; in- 
ternal border straight to feebly arcuate; surface very feebly convex, 
shining, not very strongly chitinized at inner and basal third, sparsely 
punctate, sete not long, those on the apical margin longer and not 
very dense. Membrane not setose across the bottom of the sinus, the 
latter broad. 

’ Female.—Genital segment robust, trapezoidal, valves reflexed ex- 
ternally, strongly chitinized and setose. 

Valvula (Plate 5, fig. 1).—Dorsal plate oblong, strongly concave, 
margins more or less reflexed ; surface glabrous, sparsely and strongly . 
punctate, setose, setee rather short and inconspicuous; external mar- 
gin straight or feebly sinuate; apical margin inwardly obliquely 
truncate, angle acute and as prominent apically as the apex, the lat- 
ter quite short, robust and triangular, both sparsely set with short 
setee; internal margin arcuate or straight. Submarginal groové more 
or less visible from above and quite moderate. 

Appendage short and flattened, semielliptical, projecting but a 
short distance beyond the margin of the fossa and eccentrically 
placed. Fossa terminal, rather large and transverse. 

Superior pudendal membrane attaining the middle of the dorsal 
plate and feebly longitudinally rugulose. 

Basal prominences quite well developed. 

Ventrolateral surfaces strongly convex, glabrous basally and more 
or less concave before the apex laterally, very sparsely and coarsely 
punctate. Internal margins of the valves contiguous in basal and 
apical fourths. Genital fissure rather broad and fusiform. Inferior 
pudendal membrane closing the fissure in basal half. 


ee i oe) me eel 
i; > ’ a“ : 


REVISION OF ELEODIINI—BLAISDELL. 303 


Habitat—Texas (Brownsville, Charles Drury, H. F. Wickham; 
Uvalde, Wickham; Wades, Alice, and San Diego, Hubbard and 
Schwarz, collection U. S. National Museum). Mexico (coll, F. 
Bates). 

Number of specimens studied, 15. 

Type in the LeConte collection. 

- Type-locality.— Texas, near the Rio Grande River. 

Salient type-characters—Thorax finely punctate, anteriorly mod- 
erately and posteriorly slightly narrowed; sides rounded; anterior 
angles acute; base broadly rounded, humeri scarcely acute. Elytra 
striato-punctate, intervals uniseriately punctulate (LeConte). 

Diagnostic characters —Quickly recognized by its robust form, the 
elytra being strongly ventricose in both sexes, moderately caudate in 
the males, the cauda being oblique and not horizontal; in the females 
the elytral apex is slightly produced. 

The apical angles of the prothorax are distinct and not dentiform; 
the basal are rather broadly rounded, the angles may be sometimes 
feebly evident; the sides narrowing moderately anteriorly. 

As LeConte writes the punctures composing the striz of the elytra 
vary very much in size; sometimes they are very large, while at others 
they are hardly different from the more distant interstitial punctures. 

Two examples in the series before me have the elytral intervals per- 
fectly flat and the striz not in the least impressed. 

The antenne are noticeably feebly clavate on account of the joints 
one to eight, inclusive, being elongate and retaining the same width 
throughout, while joints nine to eleven, inclusive, are slightly widened 
at the middle and more or less transverse. The same structure is 
observed in eschscholtzii and luce, only that the eighth joint is also 
feebly widened. This may also be the case in ventricosa. 

This species is separated from lucw forma inflata by the basal 
angles of the prothorax in the latter being obtuse, distinct, and not at 
all rounded. 

See var. falli, p. 306. 

General characters —The mentum is large, more or less trapezoidal, 
rarely subquadrate; apex truncate or feebly arcuate, slightly reflexed, 
and frequently impressed at middle so as to appear more or less emar- 
ginate; surface coarsely punctate, sete very short, subfoveate at the 
basal angles; inflexed lobes rather stout. 

The prosternum is quite variable. Convex between the coxe, with 
which it is quite strongly protuberant ventrally and also moderately 
short before the acetabula; compressed and perpendicularly truncate 
behind, rising into a small mucro at the ventral angle; at times sub- 
truncate behind, with a moderate mucro, at others rounded and not 
at all mucronate. 


304 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


The mesosternum is quite short, obliquely declivous, and rather 
strongly concave. : 

The abdominal process is about one-fourth of its width broader 
than the metasternal salient, nearly quadrate, slightly transverse, and 
equal in length to the post-coxal portion of the same segment, the 
latter being equal to the third in length; the second is twice as long 
as the fourth. 

The metasternum laterally between the coxz is about as long as the 
width of a mesotibia at apex. 

The tibial grooves of the profemora are well developed and defined 
by smooth cariniform margins, that are scarcely at all arcuate, 
quite evenly and gradually convergent, to become contiguous a short 
distance from the base; floors glabrous and shining, feebly concave; 
the anterior margins at the outer fourth are dentately laminate in 
the males; in the female more or less sinuately. atrophic in the outer 
fourth. 

On the mesofemora the sulci are less strongly defined by cariniform 
margins, that become quite evanescent at the inner third; the floors 
are glabrous, shining, and very feebly concave. 

The metafemoral tibial grooves are rather narrow and with rather 
feeble margins, that become evanescent without becoming contigu- 
ous at about the middle. 

The protibie are scarcely compressed, not carinate, but smooth ex- 
ternally; tarsal sulci more or less feebly developed and subasperate ; 
posterior surface muricately sculptured, but not coarsely so; articular 
cavities usually closed. 

The mesotibize are more or less rounded and not coarsely muri- 
cately sculptured, smooth, and not carinate in basal third externally; 
tarsal grooves more or less obsolete and subasperate ; articular cavities 
closed. 

Metatibize quite cylindrical, not coarsely muricately sculptured, 
more or less feebly flattened and rarely grooved externally; articular 
cavities distinctly closed. 

The tarsi are moderately long and stout. 

The protarsi are about one-fourth of their length shorter than a 
mesotarsus. Joints two, three, and four are subequal in length and 
slightly wider than long, together about equal to the fifth; the first 
is slightly longer than wide. 

The mesotarsi are just slightly shorter than a metatarsus. Joints 
two, three, and four subequal in length and very slightly longer than 


wide, together equal to the length of the fifth; the first is about a 


half longer than wide. 
The metatarsi are equal to half the length of a metatibia. Joints 
two and three subequal in length, distinctly longer than wide and 


REVISION OF ELEODIINI—BLAISDELL. 305 


together scarcely as long as the fourth; first joint with a total length 
of about twice as long as wide. 
The ungues are comparatively long. 


ELEODES VENTRICOSA var. FALLI, new. 


Elongate-ovate, smooth and shining, elytra scarcely inflated, simply 
punctate and more or less caudate in the male. 

Antenne as in ventricosa, 

Prothorax quadrate, about one-eighth wider than long, not notice- 
ably more narrowed anteriorly than posteriorly; apex broadly and 
quite strongly emarginate; sides evenly and quite moderately arcuate 
from base to apex; base one-fifth to one-fourth wider than the apex 
and more or less greater than the length; apical angles strongly 
prominent anteriorly, acute, rather large, slightly reflexed and not 
everted; basal angles obtuse and not rounded. 

Propleure quite smooth and quite obsoletely sculptured. 

Elytra rather broadly oval, scarcely inflated, sides eveniy and 
moderately arcuate, with the apex more or less produced; disc quite 
evenly arcuate from side to side, sometimes slightly depressed on the 
dorsum, not suleate, punctate, punctures fine and usually of the same 
size; the serial punctures may be coarser, scarcely more strongly 
sculptured laterally. Cauda feebly impressed along the suture and a 
little longer than in ventricosa. 

Sterna and parapleure more or less obsoletely sculptured and 
glabrous. 

Legs usually rather slender; anterior tibiz slightly constricted at 
base, otherwise as in ventricosa. 

Male.—Body somewhat robust. Elytral cauda about 4 mm. long 
and less oblique than in ventricosa. The tuft of spinules on the first 
joint of the anterior tarsi is interrupted at middle and does not close 
the groove. Otherwise as in ventricosa. 

Female—Body moderately robust. Elytral apex more or less 
slightly produced and subacute. Otherwise as in ventricosa. 

Measurements—M ale: Length, 32.2 mm. with cauda; width, 12.5 
mm. Females; Length, 30-33 mm.; width, 13.2-14 mm.. 

Genital characters, male——As in ventricosa. 

Female—Compares with ventricosa as follows: The genital seg- 
ment has the apical margin of the dorsal plate of a valve less oblique 
and sometimes feebly arcuate; angle about as or less prominent. 

The ventrolateral surfaces are less strongly but more evenly convex, 
and the surface lines straight when viewed longitudinally. 

The appendage is larger, subconico-semiellipsoidal, completely fill- 
ing the fossa and projecting beyond the margins of the dorsal plate, 
not as long as wide. 

59780—Bull. 68—09——20 


306 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Fissure narrower, otherwise as in ventricosa. One of the speci- 
mens from El Paso, Texas, has the apical marginof the dorsal plate 
as in ventricosa. 

Habitat.—Texas (El Paso, March, H. Soltau; Fort Bliss, collection 
of H. C. Fall). 

Number of specimens studied, 3 (1 male and 2 females). 

Co-types distributed as follows: Male in the collection of H. C. 
Fall; female types in the collection of the U. S. National Museum. 

Type-locality.—Texas (El Paso and Fort Bliss). 

Diagnostic characters —Falli is separated from ventricosa by the 
less transverse prothorax, the very acute, anteriorly prominent, and 
more or less reflexed apical angles of the same; the basal angles are 
distinctly obtuse and not rounded; the elytra are much less inflated, 
and consequently appear more elongate, and the cauda in the male is 
longer. 

From luce forma inflata it is recognized by the more strongly de- 
veloped apical angles of the prothorax, the oblique cauda of the male, 
and the mutic anterior femora of the female. 

From tenuipes, which it approaches, it is quickly recognized by the 
well-developed anterior prothoracic angles; the cauda in tenuipes is 
slightly oblique at times, but more horizontal in wickhami. Tenuipes 
is less inflated and therefore narrower and more elongate. 

Falli is undoubtedly intermediate between fenuipes and ventricosa, 
and differs with the latter from all other members of the subgenus 
by the character of the vestiture of the anterior tarsi and the mutic 
anterior femora in the female. 

General characters.—The mentum is exactly as in ventricosa, and 
the prosternum is more or less rounded behind and feebly mucronate. 
The mesostermum is as in ventricosa. 

The relative size of the metasternal and abdominal salients are the 
same as in ventricosa. 

The post-coxal portion of the first segment is a little longer than 
the process and quite equal to the third in length; the second is about 
a fourth of its length longer than the third, the latter being about 
twice as long as the fourth. 

The metasternum laterally and the femora are as in ventricosa. 

The protibiz are more or less distinctly carinate externally, and 
the articular cavities are more or less open; the tarsal sulci are quite 
well developed, as they also are on the mesotibie, 

Otherwise the tibiz and the tarsi are relatively as in ventricosa. 


Subgenus BLAPYLIS Horn. 


The present subgenus is based chiefly upon the character of tarsal 
pubescence. The males have two or three basal joints of the anterior 


REVISION OF ELEODIINI—BLAISDELL. 307 


tarsi feebly thickened at tips beneath, and there clothed with quite 
dense, silken, yellowish or brownish pubescence. 

The first joint of the middle tarsi also bears a small tuft of similar 
pubescence at tip beneath, and at times the second exhibits a very 
small one. 

The species naturally associate themselves into groups, the relation- 
ships by which this is accomplished result from their genealogical 
development; and, accordingly, the pubescent tufts may be flat, 
slightly longer and truncate, or still longer, usually smaller and 
subacute. 

In most females the first joint of the anterior tarsi is more or less 
transversely thickened at tip beneath, and simply clothed with ordi- 
nary spinules across that point where the groove is more or less 
interrupted. 

In the males the pubescent tufts always interrupt the grooves which 
are evident on the remaining joints, and in all except the first in the 
female. 

At times there is a slight tendency to a lateral compression of the 
tarsi, but this is not always evident. The femora are always mutic; 
at times the anterior femora are slightly sinuate at apex beneath. 

The pronotum is never declivous at the sides; the disc is always 
evenly convex from side to side, but in the parvicollis section there is 
some flattening along the lateral margins corresponding to the granu- 
late area, the sides appearing somewhat dilated as a result. The 
marginal head is always distinctly visible from above, entire in 
those species which are very slightly sinuate or constricted at the 
base, and obsolete or simply continuing as a mere line to the basal 
angles in those species which are more strongly constricted. The 
base is at times so strongly constricted as to appear almost pedun- 
culated. 

The general form is oval to ovate, some species are slightly more 
elongate and oblong. The elytra are oval, with sides more or less 
strongly rounded, apex never produced; the humeral angles are 
always more or less rounded, except in ¢ébialis, where they are dis- 
tinctly angulate. 

There is in this subgenus a tendency for the anterior tibix to be- 
come constricted at base, most evident in ¢ibia/is. The tibial spurs are 
quite similar in the sexes. 

Among the species of Blapylis are to be found our smallest Eleodes. 

With the subgeneric tarsal characters are to be correlated the fol- 
lowing: 

Genital characters, male—Apicale of the edeagophore triangular 
and without a median groove, except in ¢ibialis (see p. 316) for re- 
marks upon this species). 


308 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Female.—Genital segment triangular or triangulo-parabolie in out- 
line. : 

Valvule.—Dorsal plates oblong, with a few scattered punctures 
and sete, apical margins not always well defined (see Synopsis) ; 
apices membranous or chitinous, always finely setose, generally with 
a pencil of longer hairs at tip (penicillate). 

Appendage small, mammilliform and penicillate. Superior pu- 
dendal membrane nearly attaining the apical fourth of the dorsal 
plate, or only the middle (cordata) ; genital fissure in the apical third 


of the segment. 
General characters —The crural characters are quite variable with- 


in certain limits in each species. The profemora are more or less 
clavate, the mesofemora at times feebly so in external two-thirds; 
the metafemora have their opposed surfaces quite parallel. 

The protibiz are at times subcarinate in basal half externally and 
the tarsal grooves are not developed, although sometimes indicated ; 
the mesotibie are frequently obsoletely grooved on their external sur- 
faces, and the metatibiz more or less flattened on those surfaces. 

The articular cavities are closed on all the tibie. 

The relative lengths of the tarsi and of the joints in each are prac- 
tically the same as in the other subgenera. 

The protarsi of the male are a little longer than in the female. 

The stoutness and length of the legs differ very much in the same 
species, but for purposes of comparative measurements, normal ex- 
amples have been selected. 

Distribution —Of the seventeen species given in this paper all but 
three occur in California; out of six races only two are not found in 
that State. 

Tibialis is apparently peculiar to Lower California; Jecontei to 
Colorado and New Mexico; snow? to Colorado, New Mexico west- 
ward to the Colorado River in Arizona; tenebrosa and its var. nana 
to Nevada and eastern central and northern California. 

Inculta may possibly occur on the mainland in California; it is 
authentically abundant on Santa Rosa Island off the Santa Barbara 
coast; consobrina occurs from Los Angeles County to Siskiyou 
County; scabripennis is from the Fort Tejon region; blanchardii at 
Indio and the mountains in eastern San Diego County; neotome in 
southern California, and I have taken it in numbers from the wood- 
rats nests on the hillsides of Mission Valley, San Diego County. 

Fuchsii occurs in the high mountains and forests of eastern Tulare 
County; hornii from the eastern central and northern parts of the 
State. 

Parvicollis from ledges on the hills near the coast in central parts 
of the State; var. planata in the western central, and producta in the 


es 


REVISION OF ELEODIINI—BLAISDELL. 309 


mountains and foothills from the southern to the northern parts of 
California. 

Hoppingii is at present only known from Eldorado County; c/avi- 
cornis and scabrosa maritime and lovers of sand-dunes. 

Cordata occurs from Los Angeles to northern parts of the State 
and at all extreme points of its northern distribution passes into 
pimelioides, which has the largest distribution of any species of B/la- 
pylis, occurring in Colorado, Montana, Wyoming, Idaho, Utah, Ne- 
vada, Oregon, Washington, Vancouver Island, British Columbia, its 
var. brunnipes in Idaho, Wyoming, Colorado, and Nevada. 

Most of the species are developing into a considerable number of 
incipient races. 


Subgeneric Trunk 
Fic. 6.—GENEALOGICAL DIAGRAM OF THE SUBGENUS BLAPYLIS. 


Genealogy.—It is doubtful if by any herculean effort a biologist 
could in the period of a few years, with the comparatively small 
series of specimens at hand from the different regions which they 
inhabit, work out a correct histery of descent. 

The individuals of the species making up the present subgenus are 
very protean in their divergences. 

They have evidently diverged along three main lines from the an- 
cestral subgeneric trunk (see diagram). It is quite hypothetical 
whether ¢ibialis belongs to Blapylis at all; if not, it must have con- 
geners in Mexico. If it does belong here it diverged early. 

Taking the degree of constriction of the prothoracic base as the 
criterion of divergence, two lines of descent are evident—cordata and 
pimelioides with a cordate type of prothorax and the lateral margin 


310 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


obsolete upon the strong basal constriction, and the other line con- 
tinuing reactive to a greater degree, again may ha¥e diverged dichot- 
omously, evolving the tenebrosa strain, with the sides of the pro- 
thorax evenly or nearly arcuate from apex to base; the other being 
the consobrina-parvicollis ramus. Parvicollis is not only plastic, 
but prolific, populating the regions it inhabits with puzzling varia- 
tions, many of them at the beginning of their incipient racial devel- 
opment. In these forms the sculpturing is rather fine, abdomen not 
coarsely sculptured, pronotal base quite strongly constricted, the pro- 
thorax is broader and less strongly convex and more or less impressed 
along the sides. 

The consobrina ramus apparently departed from an interesting 
strain; that is, zrculta, an insular species apparently, for if it be found 
on the mainland it is rare; in énculta the thorax is broad, very feebly 
rounded at the sides, and scarcely at all sinuate near the base, with 
the marginal bead strong and reaching the basal angles—these 
characters in contrast to the more elongate consobrina, with narrower 
prothorax which is distinctly constricted at base, the marginal bead 
becoming a fine line upon the constriction, and the elytra more tuber- 
culately sculptured. Here, again, are evidences of plasticity to en- 
vironment. 

Let the student correlate the above characteristics with the genital 
characters as presented below in the synopsis, and I believe the above 
relationships will become more evident. 


SYNOPSIS OF GENITAL CHARACTERS, 
Males. 


Apicale of the edeagophore with a median groove; sides scarcely arcuate. 
tibialis. 
Apicale without a median groove. 
lecontei. 
snowii. 
tenebrosa, 
inculta. 
consobrina. 
blanchardii. 
neotome, 
fuchsia, 
hornii. 
parvicollis, 
hoppingii. 
clavicornis. 
scabrosa, 
cordata, 
pimelioides. 


Apicale larger, sides more or Jess broadly sinuate, apex appear- 
ine. attenuate... = 9 eee eee 


Apicale rather narrow; sides feebly arcuate___--------------- 


REVISION OF ELEODIINI—BLAISDELL. $11 
Females. 


Superior pudendal membrane long, attaining the apical fourth of the dorsal 


plate. 
Apical margin of the dorsal plate not defined from the apex; appendage 
flattened mammilliform and subapical____-_-------------------- tibialis. 


Apical margin usually well defined from the apex and more or less arcuate ; 
appendage true mammilliform and more at base of the apex. 
lecontei. 
snowii. 
tenebrosa, 
inculta. 
consobrina, 
blanchardii, 
neotome, 
fuchsii. 
hornii. 


Dorsal plate rather broad, not more than three times as 
Ooo CI Jay. Be ee ae 


lavi * 
Dorsal plate elongate, four times as long as wide_---~-~ ( elec ets 


scabrosa. 
Apical margin of dorsal plate not well defined from the apex, {parvicollis. 
the latter more frequently semichitinous__.____-___---_____ Hostaon 
Superior pudendal membrane short, attaining the middle of the dorsal plate. 
Apical margin of the dorsal plate more or less truncate; anglef{cordata. 
evident and more or less rounded ; appendage at base of apex T iaiobolass 


Caseyi is aberrant. In the male the apicale is grooved and in the 
female the superior pudendal membrane is long, attaining the base 
of the apex. The material is not sufficient to indicate the relation- 
ships. 


ANALYTICAL KEY TO SPECIES OF SUBGENUS BLAPYLIS, 


Lateral marginal bead of prothorax always distinct and entire, attain- 


Ne GEES REL ESUD Ey SR ENS Nera Fe Oe ee 1 

Lateral marginal bead more or less obsolete on basal constriction_____~~~ 5 

1. Thorax transversely suboblong-oval, lateral margins feebly or not at all 
EO ee et SEE eX Ee EE ee Ol ee ie es 2 

, Thorax quadrate, not at all sinuate at base; form very elongate and de- 
, Drcssee - Suriace Smoot ana pliming o's tibialis. 
. ran ENT ICTS ENGI? CbUe s Leas RSET ee a a 3 


: Surface opaque; elytra sculptured with small granules, densely and irregu- 
larly placed. 
Size larger, length 14-18.5 mm.; granules shining and rounded_tenebrosa. 
Size smaller, length 10-12 mm.; granules coarser and submuricate 


laterally, more evidently setigerous____._______._______-_-- var. nana, 
8. Thorax moderately transverse, sides rather strongly rounded___---------_ a 

) Thorax distinctly transverse, sides very feebly arcuate, scarcely sinuate at 
3 NG Da, eee es Ae ee eee ees Se ES a inculta. 
4, Seulpturing rather finely muricato-granulate laterally; sides of the pro- 
notum evenly rounded from apex to base, sometimes briefly sinuate at 

TN re tg es Re ae et Bia SS ee eee snowii. 


Sculpturing more coarsely muricato-granulate ; pronotum briefly, feebly, and 
broadly sinuate at base; elytra flattened (or convex) on the dorsum; 
surface dull__-------, eS) eS LEE Se A i A ee ee lecontei, 


312 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


5. Thorax transversely suboblong-oval; somewhat coarsely and discretely 
punctate. 32 ee ee re ees) eee 6 
Thorax subcordate and coarsely rugoso-punctate_----------------------- 15 
6; Propleure not hairy 2-> =. te ee { 
Propleurm Dairy: 2 8s eee 14 

7. Gense not produced. +25 ae ne eee 8 
Genz produced ; sculpturing more or less coarse___-_----------- blanchardii. 

8. First three abdominal segments densely, coarsely punctate and more or less 
PULORC os a ee ees 9 
First three abdominal segments more finely and sparsely punctate, the first 
somewhat rugulose..__... => eee 10 


10. 


11 


12. 


13. 


14. 


. Pronotum and elytra moderately convex, surface dull; elytra pees 


more or less tuberculate laterally, asperate_______________ scabripennis. 
Pronotum and elytra very- strongly convex; surface shining; elytra subas- 
perately sculptured. .\— 2-4 42 2 = 8 ee neotome. 
Humeri more or less obtusely rounded, frequently prominent anteriorly_-_11 
Humeri obsolete; elytra equal to the prothoracic base, very gradually widen- 
ing to middle, causing the thorax to appear somewhat remote; surface 
smooth... shining or dull; antenne2 long... = eee hornii. 
Humeri obtusely rounded and not prominent to prominent anteriorly ; thorax 
noticeably transverse__.=— 2s. ==. sn eee 12 
Humeri obtusely to broadly and evenly rounded, not at all prominent an- 
teriorly; thorax not noticeably transverse, more as in consobrina, fre- 
quently subcordate (female) ; surface dull_______________________ fuchsii. 
Surface muricato-granulate laterally and on apex; epipleural margin at 
humeri more or less visible from above_—--=-=---.--=-==2-">_ See eee 13 
Surface finely and densely muricato-granulate; thorax somewhat finely 
punctate; form oblong-ovyal and depressed; antenne short; humeri 
TOUNQCO: =" 2 = fees ee a ee eee hoppingii. 
Surface shining; elytra short oval, convex, rather rapidly declivous behind ; 
humeri obtusely rounded, not at all prominent; pronotum rather coarsely 
punctate =... -- = 5 ee eee eee parvicollis, 
Surface more or less dull. 

Elytral base not broad; humeri rounded, feebly obtuse and not conspic- 
uous; elytra somewhat attenuate at apex, sides evenly rounded, disc 
gradually declivous behind (male). Elytral base broad, humeri 
rather broadly rounded and somewhat obtuse, conspicuous and not 
prominent anteriorly; apex less attenuate; disc more rapidly decli- 
vous Dehind: (female) =. 2 = eee eee var. planata. 

Elytral base broad; humeri conspicuous and more or less prominent 
anteriorly and margined by the epipleure; elytra oblong, dise flat- 
tened, narrowing behind the middle, sides parallel; prothorax strongly 
tPANSVCIS@n ne at eee var, producta. 

Elytral base broad; humeri as in producta; elytra oblong, dise flattened 
usually, coarsely and densely rugoso-punctate centrally, muricato- 
tuberculate laterally and on apex; thorax more sparsely and coarsely 
punctate; female often with the form of cordata____~- var. constricta. 

Pronotum not noticeably transverse; disc with rather long erect hairs lat- 
erally in the granulate area; propleural hairs rather long____clavicornis. 
Pronotum distinctly transverse; disc with very short, erect set laterally ; 
propleural hairs shorter than in clavicornis____-_-------------- scabrosa,. 


. Thorax strongly. constricted at base.-.--~----_--_=-_--_=_+_ === =e 16 


Thorax less strongly and more briefly constricted; sides broadly, strongly, 
and evenly rounded as in aoe brostia. = to var, rotundipennis, 


REVISION OF ELEODIINI—BLAISDELL. 313 


16. Sides of pronotum more strongly arcuate in front of middle, behind narrow- 
ing rather suddenly and more or less subangulate, or more evenly rounded 
and less gradually narrowing behind, 

Blytra coarsely rugoso-punctate____.____-_______- i NE TT 
Elytra sculptured with distinct tubercles, which vary in size, rounded 

or submuricate, 
Tubercles distinct, rounded, not noticeably setigerous; legs 
OE Oe RS ee ee er ee ee pimelioides. 
Tubercles more evidently setigerous; legs brown___-_var. brunnipes. 
Sculpturing finer, distinctly pubescent throughout, somewhat like 
hirsuta, hairs shorter than in that species; legs reddish or 
DROW Dian. toae coe Se us Sa ees §  otacrt, 
The reader will no doubt appreciate the difficulty of formulating a 

table that will make it possible to place all specimens, especially in a 
genus so protean as L/eodes. I desire to impress on the student the 
importance of first determining the subgenus to which the specimen 
under consideration actually belongs, then try to determine whether 
it is typical specifically or exhibiting considerable variation. The 
most important diagnostic tests are those considering the general 
form, form of thorax with its sculpturing, form and sculpturing of 
the elytra. Color plays very little part in /7eodes, although surface 
luster is often helpful. 


ELEODES TIBIALIS, new species. 


Elongate, oblong-ovate, nearly three times longer than wide, very 
smooth and shining, strongly depressed. 

Head about twice as wide as long, feebly convex, more or less obso- 
letely impressed laterally, frontal suture obsoletely indicated, very 
finely, sparsely, and irregularly punctate, punctures just noticeably 
denser laterally and on the epistoma. Antenne moderate in length 
and stoutness, outer three joints very feebly compressed, feebly and 
very gradually dilated, third joint about equal to the next two taken 
together, fourth slightly longer than the fifth, the latter, sixth, and 
seventh subequal in length, eighth more or less triangular, ninth and 
tenth orbicular, eleventh short ovate. 

Pronotum quadrate, widest at or just a little in front of the middle, 
about a third wider than long; disc evenly, very moderately convex. 
frequently somewhat depressed, more or less declivous at the apical 
angles, very minutely punctulate, obsoletely so at times, punctules 
sparse ; apex truncate to moderately emarginate, very finely and more 
or less obsoletely margined; sides quite evenly and not strongly 
arcuate, slightly less so and at times somewhat straight in the pos- 
terior third, marginal bead entire and fine; base truncate, sometimes 
feebly sinuate at middle, margin fine, a little wider than the apex; 
apical angles more or less prominent anteriorly, obtusely and nar- 
rowly rounded; basal angles obtuse and not in the least rounded. 


314 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Propleure smooth, obsoletely punctulate, and more or less feebly 
rugulose. 

Elytra elongately oval, smooth, widest at the middle, sometimes 
more posteriorly; base truncate and at times just noticeably wider 
than the contiguous prothoracic base; humeri obtusely angled, not 
rounded nor prominent; sides evenly arcuate, obliquely narrowing in 
apical third, apex not very narrowly rounded; disc depressed, feebly 
convex, sides more or less suddenly inflexed and almost evenly 
rounded, quite evenly and arcuately declivous posteriorly; surface 
usually obsoletely punctulate or finely and irregularly so, a serial 
arrangement often evident, the interstitial punctures of the same size 
as those of the series, often very feeble and eroded strie are evident; 
punctules usually denser on the sides and apex, those about the latter 
with small brownish hairs. 

E'pipleure moderately wide, gradually narrowing from base to 
apex, superior margin rather strongly defined, especially toward apex, 
beneath the humeri feebly and broadly sinuate; surface at times 
feebly concave, smooth, obsoletely punctulate, and more or less feebly 
rugulose. 

Sterna rather smooth, more or less obsoletely and sparsely punctate, 
rugose, 

Parapleure rather evenly and not densely punctate. 

Abdomen finely and sparsely punctulate, more densely so on the 
fifth segment, where it is also more or less finely setose along the 
margin. 

Legs moderate in stoutness and somewhat long; femora mutie, 
finely and sparsely punctate, the profemora very feebly sinuate in 
apical fifth; anterior and middle tibizw more or less strongly con- 
stricted at base; anterior tibial spurs similar in the sexes, the ante- 
rior a little stouter and just a little longer than the posterior, both 
acute. Anterior tarsi dissimilar in the sexes. 

Male.—Rather narrow, elongately oblong-ovate. Antenne attaining 
the base of the prothorax, the latter with base equal to the length. 
Elytra with the sides not strongly arcuate. Abdomen somewhat 
oblique apically, rather feebly convex, broadly, more or less strongly 
impressed on the first two segments, and the metasternum feebly con- 
cave, the mesosternum rather prominent. Anterior tibie suddenly 
constricted at base; first joint of the anterior tarsi feebly thickened 
at tip beneath and clothed with golden hairs. 

Female.—Rather more robust. Antenne not attaining the base of 
the prothorax, the latter with the base less than the length. Elytra 
more broadly oval, sides more strongly arcuate. Abdomen horizontal 
and moderately convex. Anterior tibiz moderately and rather grad- 
ually constricted at base; first joint of the protarsi scarcely thickened 
at tip beneath, groove entire. 


REVISION OF ELEODIINI—BLAISDELL. 315 


Two forms are recognized— 
Forma typica.—Elongate-ovate, strongly depressed and very finely 
punctulate; anterior tibiw quite strongly constricted at the base. 

Forma oblonga.—Elongate oblong-ovate, not strongly depressed, 
rather strongly punctate; anterior tibie not constricted at base. 

Measurements.—Males; Length, 20-22 mm.; width, 6.5-8 mm. 
Females: Length, 20-23 mm.; width, 9-9.2 mm. ' 

Genital characters, male—Edeagophore of the usual oblong-ovate 
form and depressed. 

Basale oblong; sides broadly and evenly arcuate; surface rather 
evenly convex, not gibbous nor noticeably arched. 

Apicale triangular; surface rather evenly convex, with a median 
shallow and rather broad groove extending to the apical fourth; 
sides scarcely arcuate; apex acute and slightly produced; base with 
a rounded lobe at middle and feebly sinuate laterally. 

Sternite feebly transverse and subparabolic in outline. Each lobe 
with the external border evenly arcuate; apex obtuse; internal mar- 
gin straight to feebly arcuate; surface glabrous and shining, not 
densely punctate, setose, setze moderately long and not dense. Mem- 
brane at bottom of the sinus, not setose. 

Female.—Genital segment elongately triangulo-parabolic in out- 
line. 

Valvula (Plate 4, fig. 16)—Dorsal plate elongate oblong, moder- 
ately narrow, sides subparallel and not reflexed; surface more or less 
feebly concave, glabrous, finely and sparsely punctate in apical third; 
external margin nearly straight in basal three-fourths and rather 
moderately arcuate in apical fourth; internal border more or less 
sinuous; apex very short, subacute, very feebly chitinized and setose ; 
sete moderately long and fine, those on apical fourth of the dorsal 
plate sparsely distributed. 

Appendage short, more or less flattened and somewhat semicircular 
in outline, sete at tip rather short, and subapical in position upon the 
apex. 

Superior pudendal membrane longitudinally rugulose, attaining 
the apical fifth of the dorsal plate. 

Ventrolateral surfaces not inflated, surface lines straight when 
viewed longitudinally, regularly convex from side to side, glabrous 
and very sparsely punctate. Apex finely and sparsely setose. Sub- 
marginal groove rather broad beneath the ,more or less explanate 
external border of the dorsal plate, the latter sometimes more or less 
translucent. Internal margins of the valves contiguous in basal 
three-fourths; fissure apical and triangular between the slightly 
diverging ventral plates. Inferior pudendal membrane attaining 
the base of the apices and visible in the apical fissure. 
Habitat—Lower California. 


316 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Forma typica.—Sierra Laguna and La Chuparosa. 

Forma oblonga.—San Francisquito. © 

Remarks.—Tibialis has been heretofore wrongly identified, and un- 
recognized as a distinct species. The following will correct the 
erroneous report as given by Doctor Horn.*| The specimens were in 
the collection of the academy, but were destroyed in the recent con- 
flagration (April 18, 1906) in San Francisco. 

Forma typica.—Female from Sierra Laguna labeled and reported 
as humeralis; two females from La Chuparosa reported as gracilis, 
and a female from the same place was also given as gracilis. 

Forma oblonga.—A male and female from San Francisquito, re- 
ported as humeralis. 

Number of specimens studied, 6. 

Type in my own collection foaleyg co-types in that of Mr. Furhet 

Ty pe-locality.—Sierra Laguna, Lower California. 

Salient type-characters—Surface smooth and more or less aluta- 
ceous. Body elongate and depressed. Prothorax widest at the mid- 
dle, sides not strongly arcuate, less so posteriorly, marginal bead en- 
tire; apical angles slightly prominent anteriorly and narrowly 
rounded; basal angles obtuse and not rounded. Elytra with the 
humeri obtusely angled; surface finely, irregularly and obsoletely 
punctulate. Anterior tibiz strongly and suddenly constricted at base. 

Diagnostic characters —Tibialis is a unique and very distinct 
species in the United States fauna. I could not recognize it among 
the Mexican species given in the Biologia. Although aberrant it will 
have to constitute a group in the present subgenus until more material 
can be studied; it is not far out of place, as the first joint of the 
protarsi is clothed with yellow pubescence beneath and the genital 
characters are more in harmony here than elsewhere. It differs from 
all other species in its elongate, depressed, and subnyctobatoid form ; 
the protibiw are constricted at base in the typical form, but not in 
the oblong form. 

General observations —The mentum is moderate in size, parabolo- 
quadrate in form; inflexed lobes small, apex truncate or more or less 
arcuate and more or less deflexed; surface moderately densely pune- 
tate, punctures not crowded nor noticeably setigerous, laterally more 
or less feebly foveate and feebly convex centrally; base of the ligula 
rather more exposed than usual. 

The prosternum is not strongly prominent ventrally, evenly convex 
antero- posteriorly between the cox and also behind, smooth and in 
the specimens before me not in the least produced posteriorly. 


4 Proc pedinet California ‘Aradeny of Sciences, 2d Ser., IV, Pt. 1, pp. 349-350. 


A 


REVISION OF ELEODIINI—BLAISDELL. 317 


The mesosternum is subvertical in its anterior two-thirds and more 
or less arcuate; surface smooth, broadly and more or less concave. 

The abdominal process is quadrate (male) or slightly transverse 
(female) and about a fourth of its width wider than the metasternal 
salient. 

The post-coxal part of the first segment is longer than the process 
and equal in length to the third (male) or a little longer in the 
female; second segment is about twice as long as the fourth. 

The metasternum laterally between the coxe is as long as the width 
of a metafemur at base. 

The profemora are moderately clavate; the tibial grooves are rather 
wide externally, gradually and quite evenly narrowing in basal two- 
thirds; floors quite concave externally, margins distinct, irregularly 
and unevenly, minutely subcrenulate, becoming contiguous at base ; 
both margins are more or less obsolete opposite the tibial constriction. 

The mesofemora are scarcely subclavate, although feebly and evenly 
tumid; grooves rather narrow, with margins as on the profemora, 
but less strongly marked and less sinuate at apex. 

Metafemora very feebly increasing in width from base to apex; 
grooves less developed than on the mesofemora, margins obsolete at 
middle and continued to base as lines of asperities. 

The tibiz may be more or less feebly arcuate, most noticeably so in 
the protibie, the latter more or less carinate externally in basal half 
or two-thirds; tarsal grooves not present and the articular cavities are 
closed. 

The tarsi are apparently slightly elongate and moderate in stout- 
ness. 

The protarsi in the male are just noticeably stouter and slightly 
longer than in the female. They are about a third (male) or a fourth 
(female) of their length shorter than a mesotarsus. 

The mesotarsi are about an eighth (male) or a seventh (female) of 
their length shorter than a metatarsus, the latter about a half of its 
own length shorter than its metatibia. 


ELEODES SNOWII, new species. 


Oblong-ovate, more or less shining, about twice as long as wide; 
sides of the pronotum evenly rounded from apex to base; elytra more 
or less muricately granulate and not distinctly flattened on the 
dorsum. 

Head twice as wide as long, more or less feebly convex, more or less 
feebly impressed laterally, frontal suture more or less evident, rather 
finely and irregularly punctate, slightly denser and coarser laterally 
and on the epistoma, sparser on the vertex. Antenne moderate, very 
feebly compressed and not noticeably incrassate, third joint about 


318 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


equal in length to the next two combined, fourth to the seventh inelu- 
sive subequal, eighth subtriangular, ninth and tenth suborbicular, 
eleventh short ovate. 

Pronotum transversely oval, about one-half wider than long, widest 
at the middle; disc moderately and evenly convex, not very finely and 
quite densely punctate, denser and granulate laterally; apea more or 
less feebly emarginate and more or less obsoletely margined; sides 
quite evenly and moderately rounded, rarely just the least feebly sin- 
uate before the basal angles, marginal bead fine and entire; base quite 
truncate, more or less finely and obsoletely margined, subequal to or 
about a seventh wider than the apex, frequently a little shorter than 
the length; apical angles obtuse, narrowly rounded at tip and not 
prominent; basal angles obtuse and not rounded. 

Propleure more or less shining, irregularly muricato-granulate and 
rugulose. : 

Elytra oval, less than twice as long as wide, widest at the middle; 
base not emarginate and evidently wider than the contiguous pro- 
thoracic base; humeri usually broadly rounded, sometimes obtusely 
and less broadly so; stdes evenly arcuate, apex not very narrowly 
rounded; disc more or less feebly or moderately convex on the dorsum, 
laterally more strongly rounded, not suddenly inflexed, although 
somewhat so occasionally, arcuately declivous posteriorly; surface 
rather finely and submuricately punctate centrally, laterally and on 
apex muricato-granulate, granules and punctures irregularly but 
closely placed; punctures usually more or less simple at centre of the 
disc. 

Epipleure at middle as wide as the mesofemur at base, gradually 
narrowing from base to apex, superior margin straight beneath the 
humeri when viewed longitudinally from the apex; surface smooth, 
and obsoletely punctate. 

Sterna more or less densely punctate. 

Parapleure quite densely punctate. 

Abdomen shining, more or less densely punctate, more or less 
rugulose, frequently reticulately so; last segment occasionally rufous. 

Legs moderate in length and stoutness; anterior femora more or 
less sinuate in outer fifth and mutic; anterior tibize more or less con- 
stricted at base, spurs quite similar in the sexes, the anterior slightly 
longer and stouter than the posterior; anterior tarsi with the first 
joint feebly thickened at tip beneath, and dissimilar in the sexes. 

Male.—Body somewhat slender, antennez attaining the base of the 
prothorax; elytra quite evenly, arcuately and feebly obliquely de- 
clivous posteriorly. Abdomen slightly oblique, not very moderately 
convex; first two segments broadly and not strongly impressed at 
middle. Anterior tarsi with the first two joints clothed with golden 


REVISION OF ELEODIINI—BLAISDELL. 319 


pubescence at tips beneath, tuft of the second joints about one-half 
the size of the first. 

Female.—Rather robust, antenne scarcely attaining the base of 
the prothorax. Elytra somewhat broadly oval, arcuately and more 
or less vertically declivous behind. Abdomen horizontal, and rather 
strongly convex. Anterior tarsi without golden pubescence on the 
first two joints beneath, the produced tips clothed with the ordinary 
ferrugineous spinules that may interrupt the groove. 

Measurements.—Madles: Length, 14-17.5 mm.; width, 6-7 mm. 
Females: Length, 15.2-18 mm.; width, 7-8.5 mm. 

Genital characters, male—Edeagophore oblong fusiform; not 
arched. 

Basale oblong, scarcely three times as long as wide, evenly convex; 
sides quite parallel. 

Apicale small, nearly equilaterally triangular; surface moderately 
convex, without groove; sides more or less sinuate, tip acute; base 
broadly lobed, feebly sinuate laterally. 

Sternite transversely quadrate. Each lobe long and narrow, tri- 
angular; external margin sinuate basally, obliquely arcuate apically ; 
apex very narrowly rounded; internal border quite straight; sur- 
face very feebly convex, rather densely punctate in apical half, 
setose, setae moderately long. Membrane not setose across the bot- 
tom of the sinus, the latter broad. 

Female.—Genital segment triangular, slightly longer than wide, 
superior surface plane. 

Valvula (Plate 4, fig. 12).—Dorsal plate very slightly narrowed 
from base to apex, sides feebly convergent or parallel; surface plane, 
glabrous, sparsely punctate, sete not evident; margins nearly straight; 
apical margin more or less evenly rounded, often rather prominent 
internally. Apex long and slender, sometimes moderate, subacute, 
external lobe obsolete, finely setose. 

Appendage short mammilliform, clothed with a few short sete, 
three or four longer ones at tip. Fossa at the base and external sur- 
face of apex. 

Basal prominences not evident. 

Superior pudendal membrane reaching to near the apical margin 
of the dorsal plate, longitudinally rugulose. 

Ventrolateral surfaces as in tenebrosa. 

Habitat.—Colorado (Ouray, elevation 7,500-8,000 feet, July, H. F. 
Wickham); New Mexico (Santa Fé Canyon, elevation 7,000 feet, 
August, F. H. Snow; Clouderoft, March and August, Warren 
Knaus); Arizona (Williams and Flagstaff, Barber and Schwarz; 
Oak Creek, August 2, Eug. Smith, from Knaus; along the Colorado 
River, Charles Fuchs). 

Number of specimens studied, 26. 


320 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Sexitypes in my own collection. 

Ty pe-locality—Santa Fé Canyon, New Mexico® Prof. F. H. Snow, 
collector. 

Salient type-characters—More or less shining. Sides of the pro- 
notum evenly arcuate from base to apex, surface not very finely punc- 
tate, punctures rather densely placed, denser laterally, where it is also 
granulate. Elytra convex not distinctly flattened, although somewhat 
so in the male; not very suddenly rounded, and inflexed laterally ; 
rather finely and submuricately punctate centrally; muricato-granu- 
late laterally and on apex. 

Diagnostic characters.—In snowii the sculpturing is not strongly 
muricato-granulate as in decontez, where the elytra are more muri- 
cato-tuberculately sculptured. In snow the tendency is toward a 
simpler form of muricate punctuation, and, besides, the form is 
more elongate and the elytra usually moderately convex. Snowii has 
not the facies of a planipennis; typical leconte? has. 

The side margins of the pronotum are at times similar in snowii, 
inculta, tenebrosa, and hornii. In ineulta the body is much more 
robust and the pronotum distinctly transverse, the elytral punctua- 
tion is rather finely muricate; tenebrosa is opaque and the elytra 
sculptured with fine shining granules; hornd has the pronotum more 
narrowed at base, antenne longer, humeri absent, and the elytra 
slowly widening from the base. 

A series of specimens collected at Williams and along the Colorado 
River in Arizona are more elongate than the types or other New 
Mexican specimens, and several of them have the pronotal sides dis- 
tinctly and briefly sinuate in front of the basal angles. Others from 
the same place, and taken at the same time, have the sides evenly 
arcuate from apex to base; in every other respect they are identically 
the same; one female is more robust and quite like a female from 
Cloudcroft, New Mexico. Specimens from Colorado and New Mexico 
also show this variation; tenebrosa and inculta present analogous 
variations. 

A female example collected at Oak Creek, Arizona, and before me, 
is more robust, somewhat like a large variety of parvicollis, but 
from some specimens of nigrina of similar facies it is almost impos- 
sible to separate it, in fact it was impossible until I examined the 
genital segment, which agreed exactly with those from Williams 
and along the Colorado River. 

I have no hesitation in referring the above specimens to the present 
species, as there is nothing to warrant me in separating them; the 
genital characters agree. It might be suggested that those specimens 
with sinuate side pronotal margins approach consobrina, but the 
latter is more robust in form and with coarser sculpturing and more 
strongly punctate abdomen. I have never seen a true consobrina 


REVISION OF ELEODIINI—BLAISDELL. 321 


east of the Colorado River. I would sooner believe that they were 
variations of tenebrosa. <A single specimen taken in New Mexico 
and in the U. S. National Museum collection is robust and resembles 
consobrina in this respect, but I would still refer it to snow7ii, even 
if I did not know the locality from which it came. 

A much larger amount of collecting must be done and larger series 
obtained before the relationship can be worked out, and besides we 
must learn something about hybrids and atavistic reversions. 

The Arizona specimens are not as strongly sculptured, and more 
shining, with the humeri more obtusely angled than the New Mexican 
examples, which have the humeri broadly and evenly rounded. 

The typical, sinuate, and robust forms must be recognized and so 
labeled in our collections. 

General observations —The mentum is somewhat small and para- 
bolic or trapezoido-parabolic in outline, the apex may be slightly 
deflexed ; the surface is feebly convex, very slightly foveate laterally 
and not strongly punctate, the punctures are not noticeably setigerous. 

The prosternum is more or less prominent ventrally with the coxe, 
sometimes very slightly so; horizontal between the cox, submu- 
cronate to vertically truncate behind, or arcuately convex antero- 
posteriorly ; surface more or less longitudinally grooved between the 
Coxe. 

The mesosternum is arcuately convex and more or less broadly 
concave. 

The metasternum laterally between the coxe is as long as the width 
of a mesotibia at apex. 

The abdominal process is quadrate (male) or very slightly trans- 
verse (female) and from a fourth to a third of its width broader 
than the metasternal salient. 

The post-coxal part of the first abdominal segment is equal in 
length to the process and also to that of the third; the second is 
about one-half (male) or not twice as long (female) as the fourth. 

The tarsi are moderate, the anterior in the male is noticeably 
slightly longer than in the female. 

The protarsi are about one-half (male) or a fourth (female) of 
their own length shorter than a mesotarsus. 

The mesotarsi are subequal to (male) or a little shorter (female) 
than a metatarsus. 

The metatarsi are about two-thirds of their length shorter than a 


metatibia. 
ELEODES LECONTEI Horn. 


Bleodes lecontei Horn, Trans. Amer. Phil. Soc., XTV, 1870, p. 316. 
Bleodes subaspera LeConte, Smithson, Miscell. Coll, 167, Pt. 1, 1866, p. 114. 
Ovate, about twice as long as wide, surface rather dull, elytra dis- 
tinctly flattened and muricately punctured. 
59780—Bull. 68—09——21 


322 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Head about twice as wide as long, scarcely gonvex between the 
eyes, very feebly impressed laterally, punctures rather fine, some- 
what densely placed and quite evenly distributed. Antenne long, 
rather slender, outer three joints very feebly compressed and not 
dilated, third joint scarcely as long as the next two combined, fourth 
just noticeably longer than the fifth, the latter, sixth, and seventh 
subequal in length, eighth just the least shorter, slightly wider and 
subtriangular, ninth and tenth suborbicular, eleventh irregularly 
short-ovate. 

Pronotum somewhat transversely oval, about a third wider than 
long; disc moderately and evenly convex, rather coarsely and densely 
punctate, the punctures crowded and granular laterally ; ape scarcely 
emarginate and more or less obsoletely beaded; sides rather strongly 
rounded, scarcely briefly sinuate before the basal angles and finely 
beaded ; base truncate to feebly arcuate, obsoletely margined, about a 
seventh wider than the apex and quite equal to the length; apical 
angles distinct and subacute, not in the least prominent anteriorly; 
basal angles obtuse. 

Propleurw rather densely punctate and rugulose. 

Elytra oval, about a half wider than the thorax, widest at the 
middle; base truncate; humeri very obtuse and rounded; sides evenly 
arcuate, apex rather narrowly rounded; disc very feebly convex, 
typically depressed, rather suddenly and somewhat narrowly rounded 
laterally and inflexed, dorsum frequently more strongly convex, 
arcuately declivous posteriorly; surface muricato-granulately punc- 
tate, punctures irregularly and quite densely placed, becoming dis- 
tinctly granular and setigerous laterally and on apex, sete short and 
inconspicuous, the granules are small and scarcely subseriately 
arranged. — 

Epipleure moderate in width, gradually narrowing from base 
to apex, superior margin quite straight beneath the humeri; surface 
more or less obsoletely punctate. 

Sterna rather densely punctate and more or less rugose. 

Parapleure rather densely but not strongly punctate. 

Abdomen more or less shining, quite evenly and not very densely 
punctate, punctures denser on the fifth segment, the first more 
strongly sculptured. 

Legs moderate in length and somewhat slender; anterior femora 
mutic; anterior tibial spurs rather short and apparently subequal 
in length and stoutness, and probably similar in the sexes; anterior 
tarsi dissimilar in the sexes, 

Male—Unknown to me. 

Female.—Body rather broadly ovate, antenne reaching a little 
beyond the base of the prothorax. Elytra arcuately and more or 
less vertically declivous behind; abdomen horizontal and rather 


a ee 


REVISION OF ELEODIINI 


BLAISDELL. 323 


strongly convex; anterior tibiae somewhat constricted at base; an- 
terior tarsi with the first joint slightly thickened at tip beneath, and 
there clothed with a small tuft of piceous spinules. 

Measurements—Female: Length, 16.2 mm.; width, 8 mm. 

Genital characters, male.—Not studied. 

Female.—Genital segment triangular and somewhat depressed, 
glabrous, very sparsely setose, and well chitinized. 

Valvula (Plate 5, fig. 21).—Dorsal plate suboblong, slightly widest 
at base, external border slightly convergent apically and _ slightly 
explanate; surface somewhat outwardly declivous, feebly convex 
but slightly concave before the apex, glabrous, with a few coarse, 
seattered, setigerous punctures, sete short; external border feebly 
arcuate or straight, passing arcuately into the apical border, the latter 
defined from the apical surface, angle not evident; internal border 
more or less feebly sinuous. : 

Apex fully chitinized, feebly convex above, scarcely excurved, 
rather broadly rounded at tip, with a number of moderately short 
setee over the base along the apical margin of the dorsal plate; in- 
ternal valvular membrane densely clothed with very short seta; fossa 
an excavation at the base of the external surface of the apex and 
beneath the apical margin of the dorsal plate. 

Appendage small and mammilliform, not conspicuous, with a tuft 
of moderately long sete at tip. 

Superior pudendal membrane longitudinally rugulose and attaining 
the apical third of the dorsal plate. 

Ventrolateral surfaces flattened centrally with the surface limes 
straight when viewed longitudinally, laterally moderately arcuate 
and sparsely setose; surface glabrous; submarginal groove distinct 
beneath the slightly explanate external border of the dorsal, plate 
and terminating in the fossa. Apex somewhat concave beneath and 
somewhat densely set with short sete. Internal margins of the 
valves contiguous at basal two-thirds; fissure apical, apices not con- 
tiguous. Inferior pudendal membrane not visible. 

Habitat.—Colorado (no special locality indicated by the labels). 

Number of specimens studied, 1 female, which was carefully com- 
pared with type and pronounced typical by Mr. Blanchard; the 
specimen is in his collection. 

Type in the LeConte collection. 

T ype-locality— Colorado. 

Salient type-characters—Somewhat shining. Head and thorax 
thickly and finely punctate, the latter with apex scarcely emarginate ; 
apical angles acute, not prominent; sides finely margined and strongly 
rounded, not sinuate posteriorly; base strongly truncate and scarcely 
narrower than the apex; basal angles obtuse. Elytra with the dor- 
sum slightly convex, apex strongly declivous, sides suddenly inflexed, 


824 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


with subseriately placed small granules, on each side and apex 
briefly piliferous. Antenne slightly incrassate externally. Length 
.60 (subaspera) (LeConte). 

Diagnostic characters—The present species was originally de- 
scribed by LeConte as subaspera; this name having been used by 
Solier at an earlier date. Doctor Horn renamed it in honor of Doctor 
LeConte. 

The single female specimen before me has been kindly loaned for 
study and pronounced by Mr. Blanchard as being exactly typical. 
In its general facies it is suggestive of planipennis. The elytra are 
distinctly flattened on the dorsum, and the sides of the pronotum are 
evenly rounded nearly to the basal angles where they become 
slightly subsinuate. 

Lecontei is much less shining than planipennis and the sculpturing 
is distinctly more asperate; the head and thorax are more coarsely 
punctate. Although the male is unknown to me, it is safe to as- 
sume that the secondary sexual characters are the same as in snowiéi. 

I believe that the elytra are more fortuitously depressed than nor- 
mally so and that the average specimens will have the elytra nor- 
mally convex. I surmise that I have examined such examples and 
erroneously referred them to snowii before I fully recognized the 
species under consideration. 

The genital characters in the female are the extreme for the sub- 
genus Blapylis, the valvular apex is fully chitinized, it is also setose 
and not produced, characters not observed in planipennis from which 
it is perfectly distinct and not at all closely related. 

From snowi?, which appears to be worthy of specific standing—if 
not, at least it would be a good race of decontec—it must be carefully 
differentiated. For a long time I have confused the two species; the 
sculpturing is quite similar in the two, but much less strongly and 
coasely granulate and more of the muricate type in snowi, where 
also the elytra are less broadly oval and more elongate and shining. 

In parvicollis and cordata and their races the pronotum is more or 
less strongly constricted at the base, less so in consobrina, where the 
body is more robust. 

Tn lecontei the marginal bead of the pronotum attains the base. 

General observations.—The mentum is moderate and rather para- 
bolic in outline, finely and not deeply punctate, very feebly convex, 
and obsoletely foveate. 

The prosternum is moderately prominent ventrally with the coxe, 
arcuately convex antero-posteriorly, and not in the least mucronate 
behind. 

The mesosternum is feebly arcuate and oblique, very slightly con- 
cave, 


— 


ee id ia as 


REVISION OF ELEODIINI—BLAISDELL, 325 


The metasternum laterally between the cox is as long as the width 
of a mesotibia at the apex. 

The intercoxal process of the abdomen is slightly transverse and 
about a third of its width broader than the metasternal salient. 

The post-coxal portion of the first abdominal segment is equal in 
length to the process and also to that of the third; the second is 
about twice as long as the fourth. 

The protarsi are about a fourth of their length shorter than a meso- 
tarsus; the mesotarsi are about a fifth of their length shorter than a 
metatarsus; the metatarsi are about three-eighths of their length 
shorter than a metatibia. 


ELEODES TENEBROSA Horn. 


Eleodes tenebrosa Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 816. 


Oblong-oval, about two and a third times longer than wide; elytral 
sculpturing consisting of very small shining tubercles arising from a 
very opaque base. 

Head twice as wide as long, feebly convex, moderately impressed 
laterally and frequently along the frontal suture, which is usually 
not evident, not coarsely and more or less densely punctate, punctures 
distinctly crowded laterally and on epistoma. Antenne moderate in 
length, slightly robust, outer four joints very feebly compressed and 
just noticeably wider, third joint equal to the next two taken together, 
fourth just the least longer than the fifth, the latter, sixth, and 
seventh subequal in length, eighth about as wide as long and subtri- 
angular, ninth subtrapezoidal, tenth suborbiculo-trapezoidal, eleventh 
frequently slightly smaller and short-ovate. 

Pronotum transversely oval, widest at the middle, a third to a half 
wider than long; disc moderately and quite evenly convex, rather 
coarsely and densely punctate, granulate at the sides; apex slightly 
emarginate, and more or less obsoletely margined; sides evenly and 
strongly arcuate from apex to the basal twelfth, then rather quickly 
sinuate and nearly straight and parallel to basal angles, more rarely 
evenly arcuate from angle to angle, marginal bead moderate, feebly 
reflexed and entire; base subtruncate in circular are and finely mar- 
gined, slightly wider than the apex, and not quite equal to the length; 
apical angles obtuse and more or less narrowly rounded; basal angles 
slightly prominent and quite rectangular, sometimes subobtuse. 

Propleure smooth and opaque, sparsely and quite evenly granulate, 
acetabular convexities frequently rugulose. 

Elytra oblong oval, about a half longer than wide, scarcely at all 
wider than the prothorax, widest at the middle; dase not emarginate; 
humeri quite broadly rounded, angle sometimes obtusely indicated ; 
sides evenly and not strongly arcuate, frequently subparallel, apex ob- 


326 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


tusely rounded and not very narrowly so, not in the least produced ; 
disc more or less feebly to modertaely convex oft the dorsum, sides 
evenly and moderately rounded, arcuately declivous behind; surface 
rather densely and irregularly covered with small, rounded, shining 
tubercles, arising from a very opaque base. 

Epipleure smooth, opaque, sparsely and evenly subasperately 
punctate, moderate in width, superior margin scarcely to feebly and 
broadly sinuate beneath the humeri, as viewed longitudinally. 

Sterna quite densely punctate. 

Parapleure quite densely and coarsely punctate. 

Abdomen somewhat shining, smooth, and rather densely punctate ; 
fifth segment frequently more or less rufous; intercoxal process at 
times quite rugose. 

Legs moderate and quite strongly sculptured; anterior femora 
mutie; spurs of the anterior tibiz quite similar in the sexes, the 
anterior slightly longer than the posterior; anterior tarsi dissimilar 
in the sexes, middle tarsi similar. . 

Male—Somewhat narrow. Antenne attaining the prothoracic 
base. Elytra arcuately declivous behind. Abdomen slightly oblique 
and quite moderately convex, broadly impressed at middle of the 
first two segments. Anterior tarsi with the first joint slightly thick- 
ened at tip beneath and invested with a moderately obtusely pointed 
tuft of golden, silken pubescence, which is inclosed by a few ordi- 
nary spinules; second joint very feebly thickened with tuft not evi- 
dent, or at most scarcely half as large as that of the first. 

Female.—Rather broadly oblong. Antenne not quite attaining the 
prothoracic base. Elytra arcuately and vertically declivous posteri- 
orly. Abdomen horizontal and rather strongly convex. Anterior 
tarsi with the first joint shghtly thickened at tip beneath, groove 
interrupted by ordinary piceous or ferruginous spinules. 

Measurements —M ales: Length, 14-16 mm.; width, 5.5-6.5 mm. 
Females: Length, 15-18.5 mm.; width, 7-7.5 mm. 

Genital characters, male-—KEdeagophore flattened oblong-ovate and 
not arched. 

Basale oblong oval; surface rather feebly convex, with the sides 
moderately arcuate. 

Apicale rather small, triangular, and about a half longer than 
wide; surface evenly convex, not grooved; sides somewhat arcuate in 
basal sixth, thence feebly and very broadly sinuate to apex, the latter 
subacute; base broadly lobed at middle and feebly sinuate laterally. 

Sternite transversely parabolic. Each lobe with the outer border 
more or less evenly arcuate; apical margin broadly and evenly 
rounded; internal margin feebly arcuate; surface very feebly con- 
vex, very sparsely punctate and setose, setee moderately long, slightly 


~ ae 


REVISION OF ELEODIINI—BLAISDELL. 327 


longer on apical margin, impunctate in basal fourth. Membrane not 
setose across the bottom of the sinus, the latter narrowly triangular. 

Female—Genital segment elongately triangular, superior surface 
plane. 

Valvula (Plate 1, fig. 9).—Dorsal plate elongately oblong; surface 
plane, glabrous, obsoletely punctate and sparsely setose; sides more 
or less feebly arcuate and subparallel; apical margin feebly arcuate 
or subtruncate, scarcely rounded internally over the apical lobe, 
angle evenly rounded. Apex rather long and slender, subacute and 
finely setose, with a few long sete at tip; external apical lobe not in 
the least evident. 

Appendage short mammilliform, with about, two to four long sete 
at tip. Fossa in basal and external surface of apex. 

Superior pudendal membrane reaching to the apical fourth of the 
dorsal plate and longitudinally rugulose. 

Basal prominences not evident, evenly rounded at base. 

Ventrolateral surfaces (Plate 1, fig. 10).—Body quite triangular 
and evenly convex from side to side, glabrous, surface lines straight as 
viewed longitudinally, very feebly punctate and setose about the apex. 

Submarginal groove moderate beneath the slightly explanate ex- 
ternal border of the dorsal plate and attaining the fossa. Internal 
margins of the valves contiguous in about basal three-fourths; genital 
fissure very narrow in apical fourth; the inferior pudendal mem- 
brane is not visible. 

Habitat—California (Owens Valley, Dr. George Horn; Bodie, 
elevation 8,475 feet, July, Wickham; Lundy, 7,000-8,000 feet, July) ; 
Nevada (western, Mr. Gabb; Verdi, April, Blaisdell). 

Number of specimens studied, 125. 

Type in the Horn collection. 

Type-locality—Owens Valley, California; collector, George Horn. 

Salient type-characters—Opaque. Head rather coarsely punc- 
tured. Thorax one-third broader than long, very coarsely punc- 
tured, becoming granular at the sides; sides strongly rounded, mar- 
gin entire, basal angles moderately prominent, rectangular. Elytra 
elongate oval, moderately convex, sides moderately rounded; basal 
angles obtuse; surface rather densely and irregularly covered with 
small, rounded, shining tubercles, arising from a very opaque base 
(Horn). 

Diagnostic characters—The salient type characters differentiate 
tenebrosa from all other species of the subgenus—in fact it is quite 
unique among its congeners by not having the elytra broader at the 
widest part than the thorax. 

Occasionally specimens are observed with the sides of the pro- 
notum evenly rounded from base to apex and in this character resem- 
ble snowii, but here the surface is more or less shining and the 


328 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


pronotal disc is less coarsely punctured. These remarks apply 
especially to the Arizona or elongate form of sndwii. 

I have seen a few specimens that were rather ovate in form, with 
the elytra showing a tendency to become less elongate and wider than 
the prothorax. Such variations are expected and must be recognized. 

General observations—The mentum is usually quite parabolic in ~ 
outline, sometimes subtriangular; surface very slightly convex, not 
coarsely but quite densely punctate laterally, where it is sometimes 
subfoveate; the punctures are not noticeably setigerous. 

The prosternum is variable, more or less prominent ventrally with 
the coxee, and moderately widened behind the axes of the acetabula, 
frequently grooved at the middle. In the more strongly developed 
individuals it is horizontal between the coxe and mucronate behind; 
in others, arcuate antero-posteriorly and with a more or less well- 
developed mucro at the middle of the posterior border; again, it is 
evenly arcuate and not at all mucronate. 

The mesosternum is vertically arcuate, or more obliquely so, deeply 
concave, appearing somewhat bilobed; the concavity varies in depth. 

The metasternum laterally between the coxe is as long as the 
width of a protibia at apex. 

The abdominal process is quadrate and about a fourth of its width 
wider than the metasternal salient, and equal in length to the post- 
coxal portion of the first segment; also about equal to the length of 
the second segment (male). 

In the male the second segment is one-half longer than the fourth, 
and the third about a third longer than the same segment. In the 
female the second is twice as long as the fourth and about a fourth 
longer than the third. 

The tarsi are moderate in length and somewhat stout. 

The protarsi are about two-fifths (male) to three-eighths (female) 
of their length shorter than a mesotarsus. 

The mesotarsi are about a seventh (male) to an eleventh (female) 
of their length shorter than a metatarsus. 

The metatarsi are about three-eighths (male) to two-thirds of their 
length shorter than a metatibia. 


ELEODES TENEBROSA var. NANA, new. 


As in tenebrosa, except smaller, more strongly and densely seulp- 
tured. Sides of the pronotum slightly tuberculate, margin more 
strongly sinuate before the basal angles. Elytral tubercles larger and 
more distinct, more evidently setigerous at the sides; humeri obtuse, 
scarcely rounded to subrectangular, with the epipleural margin often 
visible at the angle. 

Measurements —Males: Length, 10.5-12.5 mm.; width, 5-5.5 mm. 
Females: Length, 12 mm.; width, 6 mm. 


REVISION OF ELEODIINI—BLAISDELL. 329 


Genital characters as in tenebrosa. 

Habitat—Nevada (Carson City, July, Wickham; Verdi, April, 
Blaisdell) ; California (U. S. National Museum collection—no exact 
locality given). 

Diagnostic characters.—Nana is more coarsely sculptured; the tu- 
bercles of the elytra are denser, confluent, and submuricate, but most 
of them still remain bright and shining. The humeri are more often 
subangulate. This form inhabits the same region as the typical race. 


ELEODES INCULTA LeConte. 


Eleodes inculta LeContr, Proc. Acad. Nat. Sci. Phila., 1861, p. 352, male.— 
Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 315. 

Robust, oblong ovate, about twice as long as wide, black, often 
somewhat piceous, feebly shining to somewhat dull; elytra irregu- 
larly and subasperately muricately punctate. 

Head about twice as wide as long, feebly convex, more or less 
broadly impressed laterally and along the frontal suture, the latter 
more or less evident; irregularly and not very finely punctate, rather 
densely so laterally and on the epistoma. Antenne moderately 
long, somewhat slender, outer four joints feebly compressed and 
feebly incrassate, third jeint scarcely as long as the next two taken 
together, fourth just the least longer or subequal to the fifth, the 
latter, sixth, and seventh subequal, eighth just a little shorter and 
more or less subtriangular, ninth and tenth suborbicular in outline 
or very feebly transverse, eleventh short-ovate. 

Pronotum more or less transversely quadrate, a third to a half 
wider than long, widest at the middle; disc moderately and very 
evenly convex, distinctly, irregularly, and not coarsely punctate, 
punctures separated by an interval equal to or twice their own diame- 
ters, narrowly subgranulate laterally; apex broadly, evenly, but not 
strongly emarginate, finely and somewhat obsoletely margined; sides 
evenly, broadly, and not strongly arcuate, very feebly subsinuate 
before the basal angles, and less so behind the apical angles, marginal 
bead fine and entire; base very feebly and broadly rounded, sub- 
truncate, rather coarsely margined, one-sixth to one-third wider than 
the apex, equal to or a little greater than the length; apical angles 
subacute, slightly prominent anteriorly, and with a slight tendency 
to eversion; basal angles obtuse and not in the least rounded, just in 
the least prominent. 

Propleure smooth and shining, sparsely and distinctly submuri- 
cately punctate and rugulose. 

Elytra oval, less than a third longer than wide, widest at the mid- 
dle; base subtruncate. /7umeri more or less broadly rounded, epi- 
pleural margin not visible from above; sides evenly arcuate, apex 


330 BULLETIN 63,°UNITED STATES NATIONAL MUSEUM. 


not yery narrowly rounded; disc moderately to rather strongly 
convex on the dorsum, sides not broadly rounded, inflexed portions 
scarcely arcuate, arcuately declivous posteriorly; surface irregularly 
and quite densely, more or less distinctly subasperately and muri- 
cately punctate, sculpturing not strong, simple centrally and not 
distinctly granulate laterally and on the apex. 

Epipleure moderately wide, gradually narrowing from base to 
apex, superior margin not strong and almost straight beneath the 
humeri; surface smooth, impunctate toward base, obsoletely and 
sparsely punctate toward apex. 

Sterna.—Prosternum scabrously rugoso-punctate; meso- and meta- 
sterna more or less shining, glabrous and rugoso-punctate. 

Parapleure rather densely, more or less evenly, and somewhat 
coarsely punctate. 

Abdomen shining, sparsely and finely punctate, more or less — 
rugulose. 

Legs rather short and somewhat slender; anterior femora mutic 
and not noticeably sinuate; tibial spurs rather slender and moderate 
in length, similar in the sexes; anterior spurs of the protibie slightly 
longer than the posterior; anterior and middle tarsi dissimilar in 
the sexes. , 

Male.—Oblong-ovate, somewhat slender. Antennz reaching a little 
beyond the base of the prothorax. Elytra evenly and arcuately de- 
clivous posteriorly. Abdomen slightly oblique apically, moderately 
convex, segments one and two broadly impressed at middle. First 
three joints of the protarsi not noticeably thickened and clothed with 
golden pubescence beneath, surface of the pads flat; first two joints 
of the mesotarsi similarly pubescent, first jot in apical half, the 
second with a much smaller tuft at tip. 

Female.—Distinctly robust. Antenne about attaining the base of 
the prothorax. Elytra broadly oval, arcuately and more or less verti- 
‘ally declivous behind. Abdomen horizontal and strongly convex. 
First two joints of the anterior and middle tarsi a little more pubes- 
cent than usual beneath, grooves not evident. 

Measurements —M ales: Length, 13.5-14 mm.; width, 6.5-6.5 mm. 
Females: Length, 14-17 mm.; width, 7.5-8 mm. 

Genital characters, male-——Edeagophore flattened oblong-ovate, 
acutely pointed at apex and scarcely arched. 

Basale oblong-suboval, moderately convex, sides feebly arcuate and 
nearly parallel. 

Apicale triangular. Surface very moderately convex, without 
groove; sides briefly arcuate at base, thence broadly and more or less 
strongly sinuate, so that the apex appears more or less attenuated in 
apical half; base evenly rounded and scarcely lobed at middle. 


REVISION OF ELEODIINI—BLAISDELL. 831 


Sternite parabolic and slightly transverse. Each lobe triangular; 
external border slightly oblique, and more or less feebly sinuate in 
basal half, thence subtruncate and feebly arcuate to apex, the latter 
moderately rounded; internal border more or less sinuate; surface 
feebly convex, rather coarsely and densely punctate in apical three- 
fourths, sete rather long, denser on apical margin. 

Female.—Genital segment (Plate 5, fig. 13) triangular, about a 
third longer than wide; surface about plane, finely setose on apex. 

Valvula.—Dorsal plate oblong, elongate, about three times longer 
than wide; surface plane, more or less declivous laterally at base, 
smooth and shining, very sparsely and finely punctulate in apical 
half, sete very fine and not distinct; external border about straight, 
feebly sinuous and very feebly converging apically; apical margin 
more or less arcuate and rather distinct from apex; internal margin 
quite straight. Apex short and scarcely more prominent than the 
appendage, subacute, membranous, finely setose, with a few rather 
long sete at tip. 

Appendage short mammilliform, with a pencil of quite long sete 
at tip. Fossa nearly occupying the external surface of the apex. 

Basal prominences not evident. 

Superior pudendal membrane scarcely longitudinally rugulose and 
attaining the apical fourth of the dorsal plate. 

Ventrolateral surfaces —Body triangular, surface lines straight as 
viewed longitudinally,.smooth and shining, feebly punctate and 
setose in apical third; submarginal groove distinct beneath the 
slightly explanate external border of the dorsal plate. Internal 
margins of the valves contiguous in basal three-fifths, fissure in 
apical two-fifths, narrow and nearly closed. 

Habitat—Nearly all of the specimens before me were taken on 
Santa Rosa Island, which lies off the coast of Santa Barbara County, 
California. One specimen bears a “ Santa Barbara Co.” label and 
another simply a “ Cal.,” while in Mr. Fuchs’ collection is one labeled 
“Alameda County.” There are no differences noticeable in the speci- 
mens. I am not sure that this species is found on the mainland. 

Number of specimens studied, 19. 

Type a male in the LeConte collection. 

Type-locality.—Santa Rosa Island; collector, Mr. C. M. Bache. 

Salient type-characters——Subopaque. Head and thorax rather 
densely punctate; thorax with the sides strongly rounded, briefly 
sinuate behind; basal angles obtuse and slightly prominent; base 
broadly rounded. Elytra oval, apex attenuate and strongly de- 
clivous; humeri broadly rounded, dorsum deplanate, punctures small 
and granular, irregularly placed, granules on each side briefly pilifer- 
ous. Antenne a little incrassate at tip (LeConte). 


332 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Diagnostic characters —Inculta is a distinct species, with a shorter, 
more robust form, distinctly transverse prothofax (female), and 
feebly subasperate (subgranular) punctuation. The sides of the 
prothorax are scarcely at all sinuate before the basal angles—in some 
specimens not in the least and in others feebly so; the basal angles 
are obtuse, distinct, and with the appearance of being just a little 
prominent; the lateral bead is distinctly entire. 

I do not believe énculta to be a synonym of scabripennis. ‘The type 
of the latter is a female and was taken at Fort Tejon. Doctor Horn 
made so many wrong determinations in //eodes that I can not be- 
lieve him right in this instance. 

Mr. Blanchard has very carefully compared a female inculta (which 
is before me) with the female type of scabripennis. His com- 
ments are as follows: ‘“ Shorter and less convex than the type of 
scabripennis, thorax much broader. It seems nearer inculta of which 
the type is a male with base of the prothorax more constricted.” 
Later and in another sending I submitted a male specimen, which 
he unhesitatingly pronounced inculta. I sent the female first inten- 
tionally with a request to compare it with scabripennis. 

The form of the thorax in snow7i (male and female) and the male 
of inculta is quite similar, but the apical angles in the latter are 
more acute and prominent, and in every example studied the sides 
are just the least subsinuate behind the angles so as to give them an 
appearance of having a slight tendency to eversion. 

The form of the prothorax should readily separate znculta from 
hornii, neotome and consobrina—none have it so transverse in the 
female, nor the sides so feebly sinuate at base as a constant character. 
In consobrina the apical angles are as prominent, but scarcely as 
acute, the sides may be feebly sinuate behind the apical angles and 
more or less strongly sinuate before the base. 

General observations—The mentum is moderate in size, sometimes 
rather small in the male, subtruncate and parabolically rounded; 
surface feebly convex, scarcely subfoveate laterally, finely punctate, 
the punctures not distinctly defined nor noticeably setigerous. 

The prosternum is moderately prominent ventrally with the coxe, 
moderately arcuate antero-posteriorly between the coxe and usually 
with a small mucro behind at middle; at other times more feebly 
arcuate and more or less vertical behind, with the angle mucroid. 

The mesosternum is arcuately declivous and more or less deeply 
and not broadly concave. 

The metasternum laterally between the cox is as long as the width 
of a mesotibia at apex. 

The abdominal intercoxal salient is subquadrate (male) or slightly 
transverse (female) and a little (male) to a fourth (female) of its 
own width broader than the metasternal process. 


REVISION OF ELEODIINI—BLAISDELL. 333 


The post-coxal portion of the first abdominal segment is equal in 
length to the process, also to that of the third (male). 

In the male the second segment is a third of its length longer than 
the fourth; in the female the second is twice as long as the fourth 
and a little longer than the third. 

The tarsi are slender and moderate in length. 

The protarsi are about two-fifths (male) to a fourth (female) of 
their length shorter than a mesotarsus. 

The mesotarsi are about a seventh (male) to a fifth (female) of 
their length shorter than a metatarsus. 

The metatarsi are about a fourth (male) to two-thirds (female) 
of their length shorter than their respective metatibie. 


ELEODES CONSOBRINA LeConte. 


Eleodes consobrina LECONTE, Ann. Lyc. Nat. Hist. N. Y., V. 1851, p. 135.— 
Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 316. 
Bleodes veseyi LEContTrE, Proc. Acad. Nat. Sci. Phila., 1858, p. 187. 

Oblong-ovate, robust, elytra coarsely and submuricately punctate 
centrally and tuberculate laterally; surface dull or feebly shining. 

Head about twice as wide as long, more or less feebly convex, 
slightly impressed laterally and often along the frontal suture, which 
is generally evident as a more or less smooth line, sides of the frons 
not usually prominent, more or less evenly, rather densely and some- 
what finely punctate, punctures slightly denser laterally. Antenne 
rather stout, moderate in length, scarcely compressed, very feebly and 
gradually widened in outer four joints, third joint comparatively 
short and scarcely as long as the next two combined, fourth to the 
eighth, inclusive, quite equal, ninth and tenth feebly transversely 
oval, eleventh short-ovate. 

Pronotum somewhat transversely oblong-oval in outline, two- 
sevenths to a third wider than long, widest at the middle; disc mod- 
erately to rather strongly and evenly convex, quite densely, evenly, or 
irregularly punctate, punctures moderately coarse, becoming granu- 
late at the sides; apex evenly and moderately emarginate, obsoletely 
margined ; sides more or less strongly arcuate and constricted at basal 
twelfth or eighth, thence briefly straight or feebly oblique to the basal 
angles, marginal bead rather thin and fine, at times somewhat coarser 
and more or less entire and reflexed; base slightly rounded and not 
very coarsely beaded, equal to or a fourth wider than the apex and 
about equal to the length; apical angles subacute to feebly rounded 
and rather prominent anteriorly; basal angles minute, obtuse or rec- 
tangular, feebly prominent at times. 

Propleure opaque, not densely muricato-granulate, and more or less 
rugulose about the acetabular convexities, 


3384 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Elytra suboblong-oval, about a half longer than wide, widest at 
the middle; base rather truncate and more or Tess sinuate. Each 
side of the middle equal to or distinctly wider than the contiguous 
prothoracic base; hwmeri more or less obtusely rounded; sédes evenly 
and moderately arcuate, apex more or less feebly attenuate, not very 
narrowly rounded; disc moderately convex, not noticeably flattened, 
evenly and strongly rounded laterally, arcuately declivous posteriorly ; 
surface irregularly, densely and coarsely, slightly submuricately 
punctate on the dorsum, sides and apex densely sculptured with rec- 
linate granules or tubercles, rarely the punctures are feebly subseriate 
on the dorsum each side of the suture. 

Epipleure comparatively moderate in width, gradually narrowing 
from base to apex, superior margin nearly straight—at most but 
feebly sinuate beneath the humeri, somewhat decurving apically, 
rarely visible from above at the humeri; surface very sparsely sub- 
muricately punctulate. 

Sterna densely punctato-rugulose. 

Parapleure more or less shining and coarsely punctate. 

Abdomen more or less shining, more or less densely and coarsely 
punctate, more strongly so and rugulose to rugose about the coxe, 
less densely and more finely sculptured on the last two segments. 

Legs moderate in length, somewhat stout, and densely but not very 
coarsely sculptured; anterior femora mutic; tibial spurs similar in 
the sexes, the anterior of the protibiz slightly longer than the pos- 
terior; tarsi dissimilar in the sexes. 

Male.—Oblong-ovate, somewhat narrow. Antenne scarcely reach- 
ing beyond the prothoracic base. Elytra somewhat attenuate in 
apical fourth, arcuately and rather obliquely declivous behind. Ab- 
domen more or less oblique and moderately convex, more or less 
broadly impressed at middle on the first two segments. First two 
joints of the protarsi slightly thickened at tip beneath, and there 
clothed with blunt tufts of yellowish pubescence, the second smaller 
than the first; first joint of the mesotarsi with a similar but moderate 
tuft, obliterating the groove. 

Female.—Robust, ovate. Antenne scarcely attaining the protho- 
racic base. Elytra arcuately and quite vertically declivous behind. 
Abdomen horizontal and rather strongly convex. First joint of the 
protarsi slightly thickened at tip beneath, groove almost entire. 

Measurements.—Males: Length, 15-16.5 mm.; width, 6-6.5 mm. 
Females: Length, 16-19 mm.; width, 7.5-9 mm. 

Genital characters, male—Edeagophore flattened oblong-ovate, 
very slightly arched. 

Basale oblong, slightly more than twice as long as wide; sides 
feebly arcuate, subparallel; surface moderately convex. 


REVISION OF ELEODIINI-——BLAISDELL. 335 


Apicale triangular, moderately convex above, without groove; 
sides slightly prominent at base, thence feebly sinuate to tip, the 
latter subacute; base with a moderate and rounded lobe at middle, 
feebly arcuate laterally. 

Sternite transversely parabolic. Each lobe triangular; surface 
glabrous, feebly convex, impunctate in basal third, punctate else- 
where, more densely so about apex, setose, setae moderately longy ex- 
ternal border obliquely arcuate; apex narrowly rounded, and con- 
tinuously so with the internal border which becomes sinuate. Mem- 
brane not setose, sinus rather broad and triangulo-oval. 

Female.—Genital segment parabolo-triangular, rather depressed, 
dorsal surface quite plane and not noticeably setose, except at apex, 
glabrous. 

Valvula (Plate 4, figs. 9 and 10).—Dorsal plate oblong, quite hori- 
zontal, with the sides subparallel; surface plane or very slightly 
concave, obsoletely punctulate; internal and external borders slightly 
arcuate; apical border more or less evenly rounded, angle not in the 
least evident. Apex triangular, rather broad, very finely setose, 
setze at tip moderately long. Fossa at base of the external surface 
of the apex behind the dorsal plate. 

Appendage short mammilliform, with a pencil of quite long sete 
at tip. 

Superior pudendal membrane longitudinally rugulose and rather 
broadly exposed, attaining the apical fourth of the dorsal plate. 

Basal prominences very smal. 

Ventrolateral surfaces triangularly plane at center to base; surface 
lines quite straight as viewed longitudinally and moderately convex 
transversely; surface smooth and impunctate. Submarginal groove 
well developed beneath the distinctly explanate external margin of 
the dorsal plate, extending obliquely across the base of the apex as 
a shallow concavity; apex finely setose. 

Internal margins of the valves contiguous in basal three-fourths; 
genital fissure in apical fourth and rather narrowly fusiform. 
Inferior pudendal membrane not visible. 

Habitat—California (San Diego County; San Bernardino Moun- 
tains; Los Angeles County; Santa Barbara County; Fort Tejon, 
Fuchs and Hopping; San Francisco County; Shasta and Siskiyou 
counties). 

Number of specimens studied, 45. 

Type.—A male in the LeConte collection. 

Type-locality— Mountains about Santa Isabel, California. 

Salient type-characters—Oblong. Thorax rounded, apex emar- 
ginate, thickly punctate, sides granulate, margin reflexed, posterior 
angles minute. Elytra thickly and coarsely punctate, sides and pos- 
teriorly rough and reclinately granulate, apex attenuate (LeConte). 


336 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Diagnostic characters —It would appear from a careful study of 
LeConte’s description that the type (male) had the sides of the 
pronotum rounded nearer to the base than in the majority of the 
specimens before me, in consequence there is a less degree of con- 
striction and the angles smaller; such is the case in a specimen which 
has been compared with the type by Professor Fall, and in his col- 
lection, and which I take to be quite typical. The example was col- 
lected in the San Bernardino Mountains. 

In all of the specimens making up a small series taken at Fort 
Tejon by Fuchs and Hopping, the sides of the pronotum are more 
strongly constricted, straight, and parallel in about the basal eighth; 
the angles are therefore larger and rectangular, but some of the 
specimens approach Professor Fall’s specimen, so that this character 
is simply a variation and of no specific value. 

The Fort Tejon specimens no doubt belong to the form described 
by LeConte as veseyi, the type of which is a large male and remark- 
able for the attempt at strie of large punctures near the middle of 
the elytra (LeConte). Mr. Blanchard writes me that veseyi is so 
near the type of consobrina that he is surprised that LeConte should 
have given it a name; it is no doubt a heterotype and therefore a 
true synonym of consobrina. In veseyi the pronotum is more strongly 
constricted and the basal angles larger and subrectangular; therefore 
it is related to the specimens above mentioned from Fort Tejon. 

An interesting specimen from Lower California was referred to 
consobrina by Doctor Horn; it was destroyed with the Academy in 
the recent disaster before I had an opportunity to study it. 

It is very difficult oftentimes to satisfactorily separate this species 
from some forms of parvicollis; as a rule in consobrina the pronotum 
is more strongly and evenly convex from side to side, and not at 
all impressed or flattened within the lateral margins; in parvicollis, 
when the pronotal surface is evenly arcuate from side to side, it 
generally arises more suddenly and strongly from the margins. 

In consobrina the humer1 are more rounded and the superior 
epipleural margin not visible from above; the elytral sculpturing 
is also coarser and tuberculate. These characters are valuable when 
the pronotal ones are weak or exhibit an approach to parvicollis, as 
they do sometimes. The heterotypes of both species are at times no 
doubt amphitypical. 

In consobrina the marginal bead is not so reliable a character as in 
tenebrosa, inculta, and snowii, for instance; the species is always 
more robust. 

It is often necessary to examine the genital segment of the female 
to more satisfactorily place some particular specimen, and males 
from the same region can be placed with the females. 


*Proc. California Acad. Sciences, 2d ser., IV, Pt. 1, p. 350. 


REVISION OF ELEODIINI—-BLAISDELL. 337 


Specimens from the Farallone Islands have been determined as 
consobrina by Doctor Horn, but I prefer to consider them as a form 
of parvicollis, for the following reasons: 

The general facies is more like the latter species, although in some 
specimens the marginal bead is entire, the specimens are decidedly 
less convex, and the pronotal dise is feebly impressed laterally, the 
basal constriction is stronger, and the sculpturing is plainly more 
muricate laterally than tuberculate. Again, parvicollis is much more 
abundant along the coast on the mainland at points opposite the 
islands than is consobrina, so that in all probability when that body 
of land became separated from the mainland it was populated with 
the form of parvicollis existing at that time, and as this change 
occurred in comparatively recent times, the species has undergone but 
slight change since and is practically identical with that which is 
now found along the coast on the mainland. 

I have collected for several years about San Francisco Bay and 
have not taken a single specimen of consobrina, while I have found 
parvicollis abundant. Consobrina is surely more abundant south 
than north of San Francisco. 

In consobrina the gene are not produced as in blanchardii, and for 
remarks on scabripennis see p. 338. In fuchsii the prothorax is com- 
paratively smaller and less convex. . 

In some specimens (females from Los Angeles Cotinty) the elytra 
are densely and distinctly tuberculate throughout. 

General observations.—The mentum is triangulo-parabolic to para- 
bolic in outline; surface convex at middle and more or less foveate 
along the sides, rather densely but not distinctly punctate and not 
noticeably setose. 

The prosternum is not strongly prominent ventrally with the 
cox, coarsely sculptured, usually distinctly grooved along the mid- 
dle, horizontal; rarely feebly arcuate antero-posteriorly between the 
cox, strongly and horizontally mucronate behind: often vertically 
truncate posteriorly with the angle more or less well developed as a 
mucro, which may be deflexed or oblique. 

The mesosternum is arcuately and obliquely declivous, deeply and 
rather broadly concave, so that it appears somewhat bilobed. 

The metasternum laterally between the cox is as long as the width 
of a mesotibia at middle or base. 

The abdominal process is subquadrate (male) to moderately trans- 
verse (female) and a third (male) to a sixth (female) wider than 
the metasternal salient. 

In the male the post-coxal part of the first abdominal segment is 
equal in length to the process, and also to that of the second; the 
latter is twice as long as the fourth; the third is a half longer than 
the fourth. 

59780—Bull. 68—09——22 


338 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


In the female the second segment is equal in length to the process; 
the third is equal to that of the post-coxal part of the first and about 
a third longer than the fourth. 

The tarsi are moderate in length and more or less stout. 


Tarsal formula: 
Pro. Meso. Meta. Metatibia. 

Male.—24 34 4 6 

! 5 


Female.—2 23 34 4 


ELEODES SCABRIPENNIS LeConte. 


Eleodes scabripennis LECoNTE, Proc. Acad. Nat. Sci. Phila., 1859, p. 77, 
female.—Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 315. 


The original description is as follows: 

“Atra, ovata, subnitida, thorace latitudine paulo breviore, subquadrato, lateri- 
bus rotundatis, postice sensim paulo angustato, angulis posticis obtusis dense 
punctato, ad latera subasperato, elytris ovalibus, thorace latioribus, dorso parum 
convexis, postice valde declivibus, granulis parvis inordinatis dense exasperatis, 
et versus suturam punctatis, tibiis muticis, prosterno postice oblique submu- 
cronato, antennis extrorsum parum incrassatis. Long. .65.” 

Habitat—California (Fort Tejon). 

Diagnostic characters —Mr. Blanchard writes me that the type is 
a robust female, with the following remarks upon the same: Stout, 
convex. Thorax convex, wider than long, sides very briefly sinuate 
at hind angles, coarsely, densely punctate, more closely and asperately 
on sides, base margined; flanks opaque, rather closely and distinctly 
granulate. Elytra convex, sides strongly rounded, coarsely granu- 
late; epipleure broad, sparsely granulate, antenne rather stout, 
joints 4-7 subelongate, Sth scarcely longer than wide. 

LeConte compares it with veseyi, saying that it has the same 
sculpturing and size, but differmg by the thorax being much less 
rounded on the sides, by the posterior angles not being at all promi- 
nent. 

I have not seen a single specimen in the material at hand which I 
could refer to this species, and personally I have no idea of its habitus. 
Fortunately I can rely upon the observations of such careful workers 
as Mr. Blanchard and Professor Fall. 

Mr. Blanchard has compared all likely specimens with the type 
at Cambridge and has failed to refer any examples to the present 
species. For remarks upon inculta see p. 332. 

Professor Fall compared a series with the LeConte type and 
labeled a male example as scabripennis. It was collected at Santa 
Barbara, California. This specimen has the form of a male conso- 
brina, from which it differs no more than the males of that species 
differ among themselves. The elytral sculpturing is more like sharp- 
ened granules than tubercles, finer and dense, with punctures simple 
each side of the suture; the pronotal punctuation is denser than in 


REVISION OF ELEODIINI—BLAISDELL. 839 


consobrina. The specimen is distinctly consobrina-like in general, 
and not at all like ‘new/ta. 

If this specimen is to be considered as an exponent of scabripennis, 
I would have no hesitation in pronouncing it as a synonym of con- 
sobrina. 

One fact is evident, that no student besides LeConte ever had 
any true conception of this species. Doctor Horn did not, for he 
referred the Santa Barbara specimens (/ncu/ta 7) to this species. I 
know of only one authentic specimen, and that is the type. 

A specimen in Doctor Van Dyke’s collection was doubtfully re- 
ferred to scabripennis; it was collected at Indio, California. It is 
very opaque and sordid in appearance; the sculpturing is rather 
coarse and dense, but upon examination I found the gene produced, 
and it is therefore blanchardii, although rather strongly sculptured 
and larger (16.5 mm.-8 mm.) than any example of the series studied. 
It makes a good companion for Casey’s specimen referred to under 
blanchardii. 

Until more collecting has been done about Fort Tejon and Santa 
Barbara, I recommend that scabripennis be retained on our lists. 

LeConte writes that the prosternum is a little more prominent 
behind than in vesey?. 


ELEODES BLANCHARDII, new species. 


Oblong-ovate, somewhat robust at times, scarcely twice as long as 
wide, elytra coarsely punctate and subtuberculate laterally ; gene dis- 
tinctly produced; surface somewhat dull. 

Head scarcely twice as wide as long, feebly convex, more or less 
feebly and broadly impressed laterally, sometimes feebly so along 
the frontal suture, the latter more or less evident as a smooth line; 
rather thickly punctate, punctures rather coarse and somewhat 
granulate, denser laterally and on the epistoma. Anfennw moderate 
in length and stoutness, feebly compressed and slightly dilated in the 
outer four joints, third joint about equal in length to the next two 
combined, fourth just the least longer than the fifth, the latter to the 
eighth inclusive subequal in length and slightly longer than wide, 
the eighth sometimes subtriangular, ninth and tenth orbicular to 
slightly transverse in outline, eleventh truncate-ovate and about as 
long as wide. 

Pronotum somewhat transversely suboval, about a half wider than 
long, widest at the middle; disc moderately evenly convex, rather 
densely and more or less evenly punctate, punctures moderate in size, 
becoming granulate at the sides; apex subtruncate in circular are, 
obsoletely beaded ; sides evenly and rather strongly rounded, more or 
less constricted or sinuate at basal ninth, thence straight or oblique 


340 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


to angles, marginal bead more or less reflexed and somewhat entire; 
base feebly rounded and feebly beaded, scarcely a seventh wider than 
the apex and equal to the length; apical angles obtuse, sometimes 
distinct and at others feebly rounded; basal angles subrectangular 
and not at all prominent. 

Propleurw more or less coarsely and rather sparsely muricato- 
granulate, more or less rugulose on the acetabular convexities. 

Elytra oval to somewhat oblong-oval, about a third longer than 
wide and widest at the middle; base subtruncate, wider than the con- 
tiguous prothoracic base; humeri more or less obtusely rounded; sides 
evenly arcuate, apex not broadly rounded; disc more or less mod- 
erately convex on the dorsum, rather strongly, evenly, and not very 
broadly rounded at the sides, arcuately declivous posteriorly; surface 
coarsely punctate, punctures irregularly placed, with a tendency to 
coalesce in twos and threes, rather simple and with a feeble corroded 
appearance on either side of the suture, becoming muricato-tubercu- 
late laterally and on the apex. 

Epipleure moderate in width, gradually narrowing from base to 
apex, superior margin rather strong, scarcely sinuate beneath the 
humeri, where it is sometimes slightly visible at the angle from above; 
surface usually impunctate and smooth. 

Sterna densely and rather scabrously sculptured. 

Parapleure coarsely and rather densely punctate. 

Abdomen more or less shining, quite coarsely and densely rugoso- 
punctate, especially about the coxe; last two segments more finely, 
less densely punctate, and not rugose. 

Legs moderate in length and stoutness; anterior femora mutic; 
anterior tibial spurs rather small and similar in the sexes, the an- 
terior slightly longer and just noticeably stouter than the posterior; 
anterior and middle tarsi dissimilar in the sexes. 

Male.—Oblong-ovate, somewhat narrow. Antenne attaining the 
prothoracie base. Elytra arcuately and-somewhat obliquely decli- 
yous behind. Abdomen slightly oblique, moderately convex, broadly 
and rather strongly impressed at middle of the first two segments. 
Anterior tarsi with the first two joints not noticeably thickened at 
tip beneath, each with a small tuft of yellowish pubescence, tufts 
subacute; first joint of the middle tarsi bearing a small tuft. 

Female.—Ovate, somewhat robust. Antenne not attaining the 
prothoracic base. Elytra arcuately and almost vertically declivous 
posteriorly. Abdomen horizontal and rather strongly convex. An- 


terior tarsi with the first joint slightly thickened at tip beneath, . 


groove scarcely interrupted by the apico-marginal spinules. 
Measurements.—Males: Length, 15 mm.; width,6.3 mm. Females: 
Length, 14.5-17 mm.; width, 7.3-7.5 mm, 


REVISION OF ELEODIINI—BLAISDELL. 341 


Genital characters, male—Edeagophore elongately oblong-ovate 
and slightly arched. 

Basale elongate oblong, slightly narrowing apically ; surface evenly 
convex and the sides feebly arcuate. 

Apicale triangular; surface moderately convex, without groove ; 
sides quite strongly sinuate in apical two-thirds, apex appearing 
attenuate and acute; base obtusely lobed at the middle and sinuwate 
laterally. 

Sternite transversely parabolic in outline. Each lobe subtriangu- 
lar; surface feebly convex and glabrous, punctate and setose in apical 
three-fourths, punctures rather dense, especially about apex, sete 
moderate in length, longer about apical margin; external margin 
more or less arcuate to apex, the latter introrse and rather narrowly 
rounded ; internal margin quite straight. Membrane not setose across 
the bottom of the sinus, the latter rather deep and triangular. 

Female.—Genital segment triangular, obtuse at apex and moder- 
ately depressed; surface nearly plane, well chitinized, and sparsely 
setose. 

Valvula (Plate 6, fig. 1). Dorsal plate oblong, four times as long 
as wide; surface glabrous and shining, slightly concave, very sparsely 
punctate and setose in apical moiety, sete. fine; sides parallel and 
more or less feebly arcuate or sinuous; apical margin evenly rounded. 
Apex short and more or less chitinized, finely and rather densely 
setose, sete slightly longer at tip. 

Appendage short mammilliform with pencil of moderately long 
sete at tip; fossa in the external surface of apex and not covered by 
apical margin of the dorsal plate, sete just noticeably longer about its 
margins. 

Superior pudendal membrane longitudinally rugulose and attain- 
ing the apical fourth of the dorsal plate. 

Basal prominences scarcely evident. 

Ventrolateral surfaces flattened centrally to base, triangular, sur- 
face lines straight, viewed longitudinally; laterally moderately 
rounded. Submarginal groove well developed beneath the slightly 
explanate external border of the dorsal plate, and attaining the fossa, 
surface scarcely at all concave before the apex. Internal margins 
of the valves contiguous in basal two-thirds; genital fissure narrow 
in apical third, inferior pudendal membrane not visible. 

Habitat.—California (San Diego County). I have taken this 
species upon the Bolean Mountain, which borders Warner’s ranch. 
It probably occurs throughout the Cuyamaca Range, also at Poway 
(elevation, 700 feet). Examples are before me from the Blanchard 
collection, and others received from D. W. Coquillett a number of 
years ago, all, however, from the above-named county. 


342 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Number of specimens studied, 9. 

Types in my own and Mr. Blanchard’s collections. 

Type-locality—San Diego County, California. 

Salient type-characters.—The buccal processes of the gene are pro- 
duced and anteriorly prominent. Pronotum moderately convex; 
apex subtruncate in circular arc; sides rather strongly rounded, con- 
stricted at basal ninth, thence straight to the basal angles, the latter 
subrectangular; apical angles obtuse and distinct. Elytra about a 
third longer than wide; humeri obtuse and rounded, epipleural 
margin just visible from above; disc coarsely punctate, punctures 
somewhat eroded, noticeably coalescing in twos and threes, simple 
about the suture, muricato-tuberculate laterally and about apex; the 
most peripheral are distinct and shining tubercles. 

Diagnostic characters—Smaller and similarly sculptured to con- 
sobrina, to which it is closely related, and easily separated by the 
unusually developed buccal process of the gene that are anteriorly 
produced and somewhat acute. 

An example before me, and kindly given me for study by Colonel 
Casey, apparently belongs here. It has the thorax very coarsely and 
densely punctate. There is also a tendency for the punctures to 
coalesce. In fact, it is more strongly sculptured throughout than 
any other specimen of blanchardii at hand. A female taken on the 
Bolcan Mountain approaches it and connects it with the typical speci- 
mens. Casey’s specimen has the buccal processes very slightly promi- 
nent and not narrowed. It is without doubt heterotypical of the 
present species; in form it agrees with a male in Mr. Blanchard’s 
collection, 

It should be noted that the elytra are shorter than in consobrina. 

I take great pleasure in remembering the kind and painstaking aid 
accorded me by Mr. Frederick Blanchard. 

General observations—The mentum is comparatively large, but 
variable in size, slightly transverse and parabolic in outline, some- 
times less strongly rounded at the sides; surface rather broadly 
convex at the center and narrowly subfoveate within the lateral mar- 
gins; not noticeably setose. 

Pro- and mesosternum as in consobrina. The metasternum laterally 
between the cox is as long as the width of a mesotibia at the middle. 

The abdominal process is subquadrate, and a fifth (female) to a 
third (male) of its width broader than the metasternal salient, also 
equal to the post-coxal part of the same segment, equal to the length 
of the third (female). 

In the male the second segment is twice as long as the fourth, the 
third being a third of its own length longer than the fourth. 

In the female the second is twice as long as the fourth. 

The tarsi are of moderate length and not as stout as in consobrina. 


REVISION OF ELEODIINI—BLAISDELL. 343 


Tarsal formula: 


Pro. Meso. Meta. Metatibia, 
Male,—2 24 3 5 
Female.—2 24 24 5 


ELEODES FUCHSII, new species. 


Ovate, subopaque, prothorax comparatively small. Elytra sub- 
asperately granulate laterally and on apex. //ead about twice as 
wide as long, more or less feebly convex, impressed laterally and 
usually along the frontal suture, the latter more or less bisinuate and 
fine, sides not very prominent, more or less evenly punctate, pune- 
tures moderately fine and not very densely placed. Antenne rather 
long, feebly compressed and slightly widened in the outer four joints, 
third joint scarcely as long as the next two combined, the fourth to 
the eighth inclusive subequal in length, the eighth slightly widened, 
ninth as wide as long and subtriangular, tenth orbicular or trian- 
gulo-orbicular, eleventh ovate. 

Pronotum scarcely to moderately transverse, a fourth to a half 
wider than long, widest at about the middle; disc moderately and 
evenly convex, rather densely and somewhat finely punctate, becoming 
more or less granulate along the sides; apea slightly and evenly 
emarginate or truncate in circular arc, obsoletely margined; sides 
rather strongly arcuate, less so behind the middle, where they appear 
somewhat oblique to the rather strong constriction at basal seventh, 
which may not be very suddenly or at other times rather gradually 
formed, thence to the angles more or less straight and parallel; mar- 
ginal bead rather fine and somewhat obsolete at basal constriction ; 
base slightly rounded, finely to obsoletely margined, very slightly 
wider than the apex and about equal to the length; apical angles dis- 
tinct and subacute, or slightly obtuse; basal angles subrectangular or 
obtuse and not in the least rounded. 

Propleure smooth and subopaque, sparsely muricato-granulate and 
more or less rugulose on the acetabular convexities. 

Elytra oval, about a third longer than wide, widest at the middle; 
base equal to or distinctly wider than the contiguous prothoracic base ; 
humeri evenly or obtusely rounded; sides broadly and moderately 
arcuate, moderately attenuate at apex, the latter not broadly rounded ; 
disc moderately convex, not depressed, quite evenly and not very 
broadly rounded laterally, arcuately declivous posteriorly ; surface 
with the punctures rather coarse, densely and irregularly placed, at 
times vaguely seriate when viewed longitudinally, punctures simple 
on each side of the suture, asperately granulate laterally and on the 
apex, minutely setigerous. 

Epipleure moderate in width, gradually narrowing from base to 
apex, superior margin straight or very feebly sinuate beneath the 


344 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


humeri, where it is at times somewhat visible from above; surface 
smooth, opaque, sparsely and quite obsoletely punctate, 

Sterna irregularly and densely punctato-rugulose. 

Parapleure rather coarsely but not densely punctate. 

Abdomen more or less glabrous, variable in dullness, rather sparsely 
and somewhat finely punctate, more coarsely, densely, and rugulosely 
sculptured about the coxee, last two segments more finely so. 

Legs moderate in length and stoutness; anterior femora mutic and 
more or less sinuate in outer fifth; tibial spurs similar in the sexes and 
rather small, the anterior slightly longer than the posterior; tarsi 
dissimilar in the sexes. 

Male.—Moderately narrow. Antenne reaching a little beyond the 
base of the prothorax. Elytra a little narrowed posteriorly, arcuately 
and rather obliquely declivous apically. Abdomen slightly oblique 
and moderately convex, impressed on the first two segments at mid- 
dle. First two joints of the protarsi slightly thickened at tip beneath 
with pubescent tufts upon the same; the tufts are truncate, that of 
the first moderate in size and about twice as large as that of the 
second. First joint of the middle tarsi with a very small tuft at tip; 
all the tufts golden yellow. 

Female.—Robust. Antenne just slightly reaching beyond the base 
of the prothorax. Elytra arcuately but scarcely vertically declivous 
posteriorly. Abdomen horizontal and rather strongly convex. An- 
terior and middle tarsi simple beneath, grooves distinct. 

Measurements.—Males: Length, 13-16 mm.; width 6-7 mm. Fe- 
males: Length 14.5-17 mm.; width 7-8.2 mm. 

Genital characters, male—KEdeagophore somewhat flattened ob- 
long-ovate, moderately elongate and slightly arched. 

Basale oblong, sides feebly arcuate, surface moderately convex. 

Apicale triangular, rather strongly convex and without a groove; 
sides rather strongly sinuate so that the apex appears produced, the 
latter acute; base with a small rounded median lobe and feebly sin- 
uate laterally. . 

Sternite small, slightly transversely oblong. Each lobe small and 
subtriangular; surface almost impunctate in basal third, thence to 
apex sparsely to densely punctate at apical margin, setose, sete 
small, increasing in length to apex where they are moderately long; 
sides sinuate and somewhat oblique in basal two-thirds, thence to apex 
truneately subarcuate; apex introrse and subacute; internal margin 
straight or feebly sinuate. Membrane not setose, sinus triangulo- 
oblong. 

Female.—Genital segment trapezo-triangular, dorsal surface very 
slightly ogival, very feebly setose and moderately depressed. 

Valvula—Dorsal plate slightly oblique, being externally declivous, 
elongately oblong, sides subparallel and moderately explanate exter- 


REVISION OF ELEODIINI—BLAISDELL. 345 


nally; surface scarcely concave, very sparsely punctate and _setose, 
sete very small; external and internal borders straight; apical mar- 
gin oblique externally and arcuate internally, angle broadly rounded, 
apex small and short, triangular and finely setose, without longer 
sete at tip; fossa at middle of the external surface. 

Appendage small and short mammuilliform, with a pencil of moder- 
ately long sete at tip. 

Superior pudendal membrane more or less longitudinally rugulose 
and attaining the apical fourth of the dorsal plate. 

Basal prominences not evident. 

Ventrolateral surfaces flattened centrally to base, rounded laterally ; 
surface smooth and not noticeably setose; sides feebly concave later- 
ally before the apices, and the submarginal groove broad beneath 
the.explanate external border of the dorsal plate. Internal mar- 
gins of the valves contiguous in basal two-thirds; genital fissure in 
apical third, moderately wide and fusiform. Inferior pudendal mem- 
brane not visible. 

Habitat—California (Tulare County, at Colony Mills, elevation 
5,415 feet, Giant Forest ; Marble Fork, Kaweah River, elevation 5,200 
feet, Kaweah to Progress Springs, elevation, 1,000-1,740 feet. Col- 
lected by Charles Fuchs and Ralph Hopping). 

Number of specimens studied, 90. 

Sexitypes in my own collection; co-types in the collections of Van 
Dyke, Fuchs, Hopping, and my own. 

Type-locality—Colony Mills, Tulare County, California. 

Salient type-characters—Pronotum comparatively small, appar- 
ently subcordate, the sides rather strongly arcuate anteriorly, behind 
the middle less arcuate and very slightly converging, constricted in 
basal seventh, thence quite straight to the basal angles, which are 
subrectangular; disc rather finely and densely punctate, becoming 
granular laterally; apical angles distinct and subacute; apex feebly 
and evenly emarginate. 

Elytra about a third longer than wide, in the female broadly oval 
and apparently inflated in contrast to the small prothorax; dise mod- 
erately convex on the dorsum, punctures rather coarse, densely and 
irregularly placed, Simple about the suture and exasperately granu- 
late laterally and about the apex. : 

Diagnostic characters.—This species is variable in form, typically 
the females are subcychriform; the heterotypes resembling conso- 
brina on the one hand and parvicollis on the other. 

In typical specimens the prothorax is comparatively small and 
apparently subcordate, the elytra broadly oval in the female, giving 
the effect of inflation, when there is none the dorsum is moderately 
convex; the humeri are broadly rounded in both sexes. Such 
examples resemble consobrina., 


346 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


In other specimens the humeri are obtusely rounded and the su- 
perior epipleural margin is often visible from “above, the thorax 
wider and more evenly rounded at the sides, and the general form 
resembles parvicollis. The integuments are more or less opaque. 

Fuchsii appears to be intermediate between consobrina and parvi- 
collis. I at first associated it with the former and then with the lat- 
ter asa race, but could not satisfy myself as to which species it is 
most closely ‘télnieds and therefore determined on the intermediate 
course and give it specific standing. 

It can be recognized from consobrina by the more finely ae much 
less densely sculptured abdomen and more cordate prothorax; from 
parvicollis by the narrower prothorax and the more sonata 
habitus. 

From neotome it is separated by the much less convex form and 
less strongly sculptured abdomen. 

Producta has a much more transverse prothorax, the humeri 
strongly prominent with the superior epipleural margin very dis- 
tinctly visible from above, the elytra very evidently flattened, form 
oblong- oval and broad at base. The males are more difficult to sepa- 
rate, the sides of the pronotal disc are usually distinctly flattened and 
granulate; in fuchsi the pronotal disc is not usually depressed at the 
sides and quite strongly and evenly convex. 

Hornii is more depressed, with very oval elytra, the humeri being 
obsolete. 

General observations —The mentum is usually parabolie in out- 
line, although at times somewhat triangular; the surface is rather 
narrowly convex at middle from apex to base and more or less 
broadly foveate laterally, finely scabrous, punctures not distinctly 
defined and not noticeably setigerous. 

The prosternum is quite gradually prominent from ‘the anterior 
margin so that the intercoxal process is not strongly nor suddenly 
protuberant ventrally; at times it is evenly arcuate antero-posteriorly 
or feebly arcuate between the coxe and subtruncate behind, with the 
mucro more or less produced or small and more or less deflexed. 

Mesosternum more or less vertically arcuate and with a varying 
concavity. 

The metasternum laterally between the coxe is as long as the width 
of a mesotibia at middle. 

The abdominal process is more or less quadrate and a third (male) 
to a fourth (female) of its width broader than the metasternal 
salient, as long as the second segment in the male and to the post- 
coxal part of the first segment, as well as the third, in the female. 

In the male the post-coxal part of the first segment is equal to the 
length of the third, the latter about a fourth longer than the fourth. 
In the female the second is twice as long as the fourth. 


REVISION OF ELEODITINI—BLAISDELL. 347 


The tarsi are moderate in length and not very short; there is evi- 
dent variation in the individuals of the series before me. 
Tarsal formula: 


Pro. Meso. Meta, Metatibia, 
Male,—2}4 3h 34 6 
Female,—2 23 3h 54 


ELEODES NEOTOMA2, new species. 


Ovate, moderately robust, shining, and smooth, strongly convex; 
Elytra scarcely asperately muricately punctate. 

Head twice as wide as long, feebly convex, more or less impressed 
laterally and along the frontal suture, the latter usually evident as a 
glabrous line, quite evenly and not very densely punctate, punctures 
moderate, sides scarcely at all prominent. Antenne moderate in 
Jength, rather stout, scarcely compressed or widened in outer four 
joints, third hardly as long as the next two taken together, fourth 
to the eighth, inclusive, subequal and rather short, the eighth just 
the least subtriangular, ninth and tenth scarcely transversely oval, 
eleventh short-ovate. 

Pronotum transversely oval, about a third wider than long, widest 
at the middle; dise evenly and strongly convex, irregularly and 
somewhat densely punctate, punctures dense and granulate laterally 
along the margins; apex not strongly emarginate and more or less 
obsoletely margined; sides evenly and moderately strongly arcuate, 
constricted at basal tenth, thence parallel or feebly oblique to the 
basal angles, margin fine and entire; base slightly rounded and rather 
finely margined, slightly wider than the apex and about equal to the 
length; apical angles subacute, not rounded and rather prominent 
anteriorly; basal angles more or less rectangular, sometimes feebly 
prominent. 

Propleure sparsely muricato-granulate and more or less rugulose 
on the acetabular convexities. 

Elytra rather broadly oval, scarcely a third longer than wide, 
widest at the middle; dase not at all wider than the contiguous 
prothoracie base; humeri rounded and quite continuously so with the 
strongly arcuate sides, apex not narrowly rounded nor produced; 
dise rather strongly and evenly convex on the dorsum, evenly and 
somewhat broadly, more strongly, rounded laterally, arcuately de- 
clivous posteriorly; surface densely, coarsely and not deeply, evenly 
and diffusely submuricately punctate on the dorsum, slightly denser 
and muricato-granulate laterally and on apex. 

Epipleure somewhat wide, gradually narrowing from base to 
apex, superior margin very broadly and feebly sinuate beneath the 
humeri, not visible from above; surface opaque, sparsely and obso- 
letely submuricately punctate at base, becoming more strongly muri- 
cate apically. 


348 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Sterna densely punctato-rugose. 

Parapleure coarsely and rather densely punctate. 

Abdomen shining, coarsely and more or less strongly punctato- 
rugulose, more densely and strongly so about the coxee, last two seg- 
ments more finely and sparsely punctate, not usually rugulose. 

Legs moderate in length and rather stout, densely and somewhat 
coarsely sculptured; anterior femora mutic and deeply sinuate in 
outer fifth; tibial spurs rather small and similar in the sexes, the 
anterior of the protibiz slightly longer than the posterior; tarsi dis- 
similar in the sexes. 

Male.—Slhghtly robust, but somewhat narrow. Antenne scarcely 
reaching beyond the base of the prothorax. Elytra arcuately decliy- 
ous posteriorly. Abdomen rather strongly oblique, moderately con- 
vex, first two segments very broadly and slightly flattened at middle. 
First and second joints of the anterior tarsi slightly thickened at tips 
beneath and with somewhat flattened tufts of golden pubescence; first 
joint of the middle tarsi with smaller tufts. 

Female.—Robust. Antenne scarcely attaining the prothoracic 
base. Elytra strongly, arcuately and vertically declivous behind. 
Abdomen moderately oblique, rather strongly convex. First joint 
of the anterior tarsi thickened at tip beneath, marginal spinules 
rather dense, groove scarcely interrupted at me 

Measurements.—Males: Length, 13-15.5 mm.; width, 6.2-7 mm. 
Females: Length, 14.5-15.5 mm.; width, 7.5-8 mm. 

Genital characters, male——Edeagophore fusiform oblong-ovate and 
feebly arched. 

Basale about two and a half times longer than wide, dorsal sur- 
face evenly and not strongly convex, sides feebly arcuate. 

Apicale triangular; surface quite strongly convex, without groove; 
sides broadly and feebly sinuate; tip subacute; base broadly rounded. 

Sternite rather short and somewhat transverse. Each lobe tri- 
angular and rather small; surface feebly convex externally, and 
declivous internally, smooth in basal third, finely and not very 
densely punctate, setose, setee gradually increasing in length to apex 
where they are moderately long; external border oblique in basal 
half, thence arcuate to apex, the latter rounded but not broadly so; 
internal margin oblique and more or less straight. Membrane not 
setose across the bottom of the sinus, the latter triangular. 

Female——Genital segment distinctly triangular, surface very 
slightly ogival, feebly-setose, rather depressed and well chitinized. 

Valvula (Plate 4, fig. 2).—Dorsal plate elongately oblong-tri- 
angular, sides more or less converging to apex; surface slightly out- 
wardly declivous, feebly concave, glabrous and shining, very sparsely 
punctate, each puncture with a minute seta, few apically; external 
border straight; apical margin arcuate to subacutely rounded; in- 


REVISION OF ELEODIINI—BLAISDELL. 349 


ternal border sinuate to feebly arcuate. Apex small and triangular, 
finely setose, tip apparently without longer sete, fossa at the base of 
the external surface. 

Appendage short mammilliform, with tuft of moderately long 
sete at tip. 

Superior pudendal membrane not ragulose and attaining the apical 
fourth of the dorsal plate. 

Basal prominences very small. 

Ventrolateral surfaces flattened centrally to base and convex later- 
ally; each lateral surface smooth, sparsely and finely punctate, setose, 
sete minute, slightly concave before the apex. Submarginal groove 
small beneath the external border of the dorsal plate, which is not 
explanate, apex finely setose, Internal margins of the valves con- 
tiguous in basal two-thirds; genital fissure in apical third, not very 
narrow and fusiform. Inferior pudendal membrane not visible. 

Habitat—California (San Diego County; Port Harford, San Luis 
Obispo County). 

I have taken this species from old stumps at Poway, elevation 700 
feet; from wood-rats’ nests on the hills bordering Mission Valley, 
near San Diego, elevation 300 feet. 

Number of specimens studied, 14. 

Sexitypes and co-types in my own collection. 

Type-locality.— Mission Valley, near San Diego. 

Salient type-characters.—Strongly convex, shining. Elytra scarcely 
asperately sculptured. Pronotum strongly convex, apex emarginate, 
sides rather strongly arcuate and constricted in basal tenth, apical 
angles not at all rounded and somewhat prominent anteriorly, basal 
angles rectangular. Elytra with the humeri rounded, and quite con- 
tinuously so with the strongly arcuate sides; punctures rather 
coarse, not deep, rather simple centrally and muricato-granulate 
laterally. Abdomen very strongly and densely sculptured, last two 
segments much less so. 

Diagnostic characters.—One of the most convex species of the sub- 
genus Blapylis, and can not be confounded with any species, except 
inculta, hornii, consobrina, and parvicollis. 

Professor Fall has very carefully compared it with types in the 
LeConte collection and found it not to correspond with any named 
species. 

Inculta is more robust and with the pronotum more transverse, 
especially in the female, the integuments are duller. The prothorax 
is not distinctly constricted at base and the abdomen is finely sculp- 
tured. 

Neotome is no doubt closely related to consobrina, but can readily 
be separated by its greater convexity, the circularly oval elytra, not 


350 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


oblong-oval as in the latter species; there is also much less difference 
in general form in the sexes than is observed in other species. 

A single female taken at Port Harford evidently belongs here; the 
elytra are broader at base and the humeri more obtusely rounded. 

Hornii is more depressed than convex and the antenne longer, the 
abdomen is not strongly sculptured nor the pronotum strongly con- 
stricted at base. 

In ‘parvicollis the pronotum is broader, the abdomen never as 
strongly sculptured, although in producta it is at times nearly so, 
but here the elytra are distinctly flattened and the humeri promi- 
nent. 

The strongly convex pronotum and elytra of neotome are very 
striking characteristics. The abdomen is rather oblique in the 
female, an unusual character for that sex. 

General observations —The mentum is rather small, usually para- 
bolically rounded, at times subtriangular: surface rather densely 
punctate, scarcely convex or foveate and not noticeably setose. 

The prosternum is very densely sculptured and scabrous, moder- 
ately prominent ventrally with the coxe, grooved along the middle, 
subhorizontal, compressed behind and quite vertically truncate, angle 
mucroid and more or less deflexed. 

The mesosternum is rather obliquely and arcuately declivous and 
more or less deeply concave. 

The metasternum laterally between the coxe is as long as the 
width of a mesotibia at middle. 

The abdominal process is quadrate (male) or slightly transverse 
(female) and a fifth (male) or a third (female) of its width wider 
than the metasternal salient. 

In the male the second abdominal segment is equal in length to 
that of the process and twice as long as the fourth: the third is 
equal to the post-coxal part of the first. 

In the female the post-coxal part of the first segment is equal in 
length to the process; the second about a half longer than the third 
and a fourth longer than the fourth segment. 

The tarsi are moderate in length and rather stout. 

The relative lengths of the tarsi were constant in the series studied. 

Tarsal formula: 


Pro. Meso. Meta. Metatibia. 
Male.—2} EBB 3 5 
Female,—2} 2% 3 5 


ELEODES HORNII, new species. 


Oval to ovate, about twice as long as wide, more or less shining, 
subglabrous; elytra feebly submuricately punctate laterally, and 
gradually widening from the base to middle; humeri obsolete. 


REVISION OF ELEODITNI—BLAISDELL. 351 


Head about twice as wide as long, slightly convex, feebly impressed 
laterally, frontal suture not evident, distinctly, ratherly coarsely, 
and quite densely punctate. Antenne quite long, moderate in stout- 
ness, outer three joints slightly compressed and feebly dilated, third 
joint equal to the next two taken together, fourth to the seventh in- 
clusive subequal, eighth scarcely shorter and slightly triangular, ninth 
and tenth triangulo-orbicular in outline, eleventh subovate. 

Pronotum widest at the middle, about a half wider than long; 
dise moderately convex, distinctly and quite densely punctate, more 
densely and narrowly punctato-granulate laterally along the margins, 
the punctures are not very strongly marked at the center and rather 
small; apex feebly emarginate or truncate in circular are and obso- 
letely margined ; sides evenly and moderately arcuate, moderately and 
not suddenly sinuate in basal seventh, becoming rather briefly straight 
to the angles, marginal bead rather fine, slightly reflexed and entire; 
base slightly rounded, obsoletely margined, and about equal to the 
apex, and also to the length; apical angles obtuse and more or 
less narrowly rounded and not prominent; basal angles minutely 
prominent. 

Propleure rather sparsely muricato-granulate and more or less 
irregularly rugulose. 

Elytra distinctly oval, widest at the middle, scarcely a half longer 
than wide; base about equal—not wider—to the contiguous pro- 
thoracic base; Awmeri obsolete or subobsolete; sides evenly and 
moderately arcuate, gradually widening from the base to the middle, 
thence more or less gradually narrowing to apex, the latter not 
broadly rounded; disc moderately convex and somewhat depressed on 
the dorsum, rather strongly and not very broadly rounded laterally, 
arcuately declivous posteriorly; swrface punctate, punctures distinct, 
rather densely and irregularly placed, rather coarse, simple on the 
dorsum, becoming submuricate laterally with a tendency to coalesce 
transversely in twos or fours. 

Epipleure moderate in width, gradually narrowing from base to 
apex, superior margin quite strong, especially beneath the humeri, 
where it is broadly and feebly sinuate; swrface subopaque and more 
or less obsoletely punctate. 

Sterna densely, subscabrously, and rather finely punctato-rugulose. 

Parapleure distinctly and rather coarsely punctate. 

Abdomen irregularly and sparsely punctate, quite densely so and 
rugose on the first segment, elsewhere rugulose, except on the fifth 
segment, where the punctures are denser at the periphery. 

Legs moderate in length and somewhat slender. Anterior femora 
mutic; tibial spurs rather small, the anterior of the protibie slightly 
thicker and longer than the posterior. Tarsi dissimilar in the sexes. 


352 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Male—Somewhat narrow, antenn reaching beyond the base of 
the prothorax. Elytra slightly wider than the prothorax, evenly 
arcuately declivous posteriorly, abdomen distinctly oblique, moder- 
ately convex, feebly and broadly flattened on first two segments. 
Anterior tarsi with the first joint slightly thickened at tip beneath. 
(In the type the tarsi are entirely denuded of spinules, but the first 
two joints in all probability have pubescent tufts on the tips beneath, 
the first larger than the second; the first joint of the middle tarsi 
although not noticeably thickened at tip, undoubtedly bears a small 
tuft of golden hairs.) Such is the case in the Calaveras specimens, 
and besides the third joint of the protarsi and the second of the meso- 
tarsi have smaller tufts at tips. 

Female—Moderately robust, antenne attaining the prothoracic 
base. Elytra rather broadly oval and distinctly wider than the pro- 
thorax, arcuately and vertically declivous posteriorly. Abdomen 
horizontal and rather strongly convex. Anterior tarsi with the first 
joint slightly thickened at tip beneath, grooves entire. 

Measurements —Male: Length, 13 mm.; width, 5.5 mm. Female: 
Length, 13.5 mm.; width, 6.5 mm. 

Genital characters, male-—Edeagophore oblong-ovate, somewhat 
flattened and quite strongly arched. 

Basale oblong, about twice as long as wide, rather strongly convex ; 
sides moderately arcuate. 

Apicale triangular, moderately convex above, without groove; sides 
moderately sinuate, apex subacute and apparently produced; base 
with a moderate, rounded median lobe and sinuate laterally. 

Sternite rather short and transverse, triangulo-parabolic in out- 
line. Each lobe subtriangular; surface feebly convex, glabrous, 
rather finely and quite densely punctate toward apex, setose, sete 
moderate and rather fine, not very long; external border oblique and 
feebly sinuate in basal half, thence obliquely truncate to apex, or 
feebly arcuate so as to appear subangulate at middle; apex subacute 
and distinct; internal margin straight or feebly sinuate and oblique. 
Membrane not setose across the bottom of the sinus, the latter tri- 
angular. 

Female.—Genital segment (Plate 5, fig. 15) obtusely triangular, 
moderately depressed, dorsal surface of each valve more or less 
slightly declivous externally, apex setose. 

Valvula.—Dorsal plate oblong, glabrous, almost impunctate; sur- 
face more or less distinctly concave; external border slightly arcuate, 
the internal more or less sinuate; apical margin more or less evenly 
rounded. Apex short and rather thick, finely setose, sete at tip 
rather long. Fossa at the middle of the external surface of the apex, 
the sete rather long around its margin. 


REVISION OF ELEODIINI—BLAISDELL. 353 


Appendage short mammilliform, with a pencil of rather long sete 
at tip. 

Superior pudendal membrane apparently not rugulose and rather 
broadly exposed, reaching about to the apical fifth of the dorsal plate. 

Basal prominences not evident. 

Ventrolateral surfaces slightly triangular from base to middle, 
sides quite evenly convex and with a few rather coarse punctures. 
Submarginal groove moderate beneath the feebly expanded external 
margin of the dorsal plate, more concave at base of the apex laterally. 
Internal margins of the valves contiguous in basal two-thirds, geni- 
tal fissure fusiform in apical third, and the inferior pudendal mem- 
brane is not visible. 

Habitat.—California (Plumas County, elevation, 1,200 feet; Moke- 
lumne Hill, Calaveras County, elevation, 1,300 feet.—Blaisdell). 

Number of specimens studied, 5 (4 males, 1 female). 

Sexitvpes in my own collection. 

Type-locality —Plumas County, California ; collector, Dr. Walther 
Horn, of Berlin, to whom I dedicate the species. 

Salient type-characters—Subglabrous and shining. Pronotum 
with dise moderately convex, rather finely, distinctly, and quite 
densely punctate, punctato-granulate along the margins laterally, 
punctures not strongly marked at center; sides evenly and moderately 
arcuate, not suddenly sinuate at basal seventh, thence rather briefly 
straight to the basal angles. Elytra gradually widening from each 
extremity to middle, humeri obsolete, surface with simple punctures 
at center of the dorsum, becoming submuricate laterally, with a ten- 
dency to coalesce transversely in twos and fours. 

Diagnostic characters —By its smaller size and more graceful form, 
smooth integuments, narrower and distinctly oval elytra, rather pro- 
portionally larger prothorax, which is somewhat remote from the 
elytra, and distinctly less convex form, hornii is sufficiently distinct 
from consobrina although derived from the same ancestral stock. 

Neotome compared with hornii is larger and strongly convex; the 
male is much less narrow. 

In fuchsti the integuments are opaque and the humeri are more 
distinct, although often broadly rounded; the elytra are broader and 
much more strongly convex—even somewhat inflated (female). 

In parvicollis the humeri are generally strongly evident and the 
prothorax distinctly constricted near the base. Parvicollis var. con- 
stricta more nearly approaches horn/i in the degree of convexity, but 
the sculpturing is different—coarser and roughly muricato-granu- 
late—the humeri are distinct, and the elytra in the males are more 
oblong oval and not narrowing somewhat equally each way from the 
middle. 


59TSO—Bull, G8—09 23 


354 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


In scabrosa the coarse sculpturing as well as the setose propleuree 
will serve to differentiate it from hornii. ~ 

Three males out of the five specimens before me were collected at 
Mokelumne Hill, Calaveras County, and are no doubt heterotypical ; 
they are subopaque, larger, the elytra are a little more strongly sculp- 
iured than in the types; the prothorax is a little more strongly con- 
stricted at the base and the basal angles are rectangular. 

The narrow elytral base, with the sides very ovaidentig widening 
to the middle, and the obsolete humeri (see Plate 12, fig. 33) give 
hornii a characteristic facies, the prothorax appearing somewhat 
remote from the elytra. 

General observations —The mentum is rather small and subpara- 
bolic in outline; the surface is distinctly convex at the middle and 
subfoveate laterally; the punctures ¢ are fine and ae dense laterally, 
not noticeably setose. 

The prosternum in the types before me is not suddenly protuberant 
ventrally with the cox, although rather strongly and gradually so, 
evenly arcuate antero-posteriorly, and not in the least mucronate 
behind. 

Mesosternum arcuately declivous, feebly and broadly concave. 

The metasternum laterally between the coxe is as long as the width 
of a mesotibia at middle. 

The abdominal process is subquadrate and about a fourth of its 
width broader than the metasternal salient, equal to the length of the 
post-coxal portion of the same segment, and also to the second. 

In the male the third segment is about a fourth of its length 
longer than the fourth. In the female the third is shghtly shorter 
than the second and about a fifth of its length shorter than the fourth. 

The tarsi are moderate in length and rather slender. The protarsi 
of the male are noticeably longer than in the female, and about a 
fourth (male) or two-fifths (female) of their length shorter than a 
mesotarsus. 

The mesotarsi are about a fifth (male) or about two-sevenths of 
their length shorter than a metatarsus. 

The metatarsi about a half (male) or a third (female) of their 
length shorter than a metatibia. 

Tarsal formula: 


Pro. Meso. Meta. Metatibia. 
Male.—2 24 4 
Female.—1 4 24 3 ft 


ELEODES PARVICOLLIS Eschscholtz. 


Bleodes parvicollis Escuscnorrz, Zool, Atlas, IIT, 1833, p. 11.—MANNER- 
HEIM, Bull. Soc. Nat. Moscow, XVI, 1843, p. 271.—LEContr, Proc. 
Acad. Nat. Sci. Phil., 1858, p. 182.—Horn, Trans. Amer.’ Phil. Soc., 
LV, 1800, p. Slt. 


REVISION OF ELEODIINI——BLAISDELL. 355 


var. planata Escuscnoirz, Zool, Atlas, Ill, 1833, p. 11.—MANNeRHEIM, 
Bull. Soc, Nat. Moscow, XVI, 1843, p. 270.—Hlorn, Trans. Amer, Phil. 
Soe, XLV, 1870, p. 817. 

var. producta MANNERHEIM, Bull, Soc. Nat. Moscow, XVI, 1845, p. 271— 
Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 317. 

var. constricta LeConter, Proc. Acad. Nat. Sci. VPhila., 1858, p. 187. 

Subovate, about twice as long as wide, more or less shining, thorax 
rather coarsely but not confluently punctate, evenly rounded atythe 
sides and distinctly constricted at the base; elytra oval, one-fourth to 
one-third longer than wide, coarsely punctate at middle, submuricato- 
granulate laterally and on apex. 

Head twice as wide as long, feebly convex, more or less impressed 
laterally, sometimes more or less so along the frontal suture, which 
is more or less obsoletely indicated; irregularly and rather coarsely 
punctate, punctures sparse on vertex and center of frons, small im- 
punctate areas frequently present, denser on the sides and epistoma. 
Antenne somewhat long and rather slender, feebly compressed and 
slightly widened in outer four joints, third joint hardly as long as 
the next two taken together, fourth scarcely longer than the fifth, 
the latter to the eighth inclusive subequal, the eighth subtriangular, 
ninth triangulo-oval, tenth just the least transversely oval, eleventh 
short-ovate and as long as wide. 

Pronotum somewhat transversely oblong-oval, two-fifths to a half 
wider than long, widest at about the middle; disc evenly and moder- 
ately convex, somewhat arcuately declivous at the apical angles, quite 
densely, irregularly, and coarsely punctate, punctures not confluent, 
becoming granulate at the sides; apex evenly but not deeply emar- 
ginate, and more or less obsoletely and finely margined ; sides evenly 
and broadly rounded, strongly constricted at about basal sixth, thence 
more or less straight and parallel to the basal angles, marginal bead 
thin and fine, feebly reflexed, becoming a mere line on the constriction ; 
base truncate, rather coarsely or indistinctly margined, equal to or 
slightly wider than the apex and about equal to the length; apical 
angles obtuse and not prominent, frequently more or less feebly 


~ rounded; basal angles more or less rectangular. 


Propleure smooth, very sparsely and finely punctato-granulate, 
more or less feebly rugulose. 

Elytra oval, widest at the middle, a fourth to a third longer than 
wide; base subtruncate, distinctly wider than the contiguous pro- 
‘thoracic base; humeri rounded, more or less obtusely so, not promi- 
nent; sides evenly arcuate, apex not very narrowly rounded; disc 
moderately convex, not distinctly flattened on the dorsum, strongly 
and not very broadly rounded laterally, inflexed portion but feebly 
convex, arcuately declivous posteriorly, coarsely punctate, punctures 
quite simple on either side of the suture and without any sign of order, 


356 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


submuricato-granulate, subasperately and feebly rugosely so later- 
ally and on apex. ~ 

Epipleure somewhat wide, distinctly dilated beneath the humeri, 
thence gradually narrowing to apex; superior margins feeble, a little 
stronger at the humeri, where it is arcuate and more or less visible 
from above, thence feebly and broadly sinuate for a fourth of its 
length; surface smooth, very sparsely and finely punctate. 

Sterna quite densely but not coarsely rugoso-punctate. 

Parapleure more or less coarsely punctate. 

Abdomen smooth and shining, sparsely and not coarsely punctate, 
more or less rugulose; quite densely sculptured about the coxee, last 
two segments less strongly so. 

Legs comparatively slender, and about moderate in length. An- 
terior femora mutic; anterior tibial spurs quite similar in the sexes, 
the anterior slightly longer and just the least thicker than the pos- 
terior; anterior and middle tarsi dissimilar in the sexes. 

Male—Somewhat narrow, subovate. Antenne reaching a little 
beyond the base of the prothorax. Elytra somewhat feebly convex 
on the dorsum and rather narrowly rounded at the sides, arcuately 
and slightly obliquely declivous behind, somewhat distinctly nar- 
rowed apically. Abdomen more or less oblique, moderately convex, 
first two segments more or less broadly flattened or impressed at 
middle. Anterior tibiae moderately constricted at base. Anterior 
tarsi with the first two joints feebly thickened at tips beneath, each 
with a moderately small truncate tuft of golden pubescence; first two 
joints of the middle tarsi with very small tufts at tips beneath, grooves 
feebly interrupted. 

Female.—Robust and ovate. Antenne scarcely reaching beyond 
the base of the prothorax. Elytra more or less evenly and arcu- 
ately declivous, or vertically so posteriorly. Abdomen rather strongly 
convex and horizontal. Anterior tarsi with the first joimt slightly 
thickened at tip beneath, grooves distinct. 

Two forms may be recognized: 


Forma typica.—Moderate in size, surface somewhat shining, shorter 
ovate. Elytra submuricato-granulate laterally and on the apex. In 
the males the elytra are rather obliquely narrowed in apical third; 
females have the elytra moderately convex on the dorsum, 

Measurements—Males: Length, 13-14 mm.; width, 6-6.5 mm, 
Females: Length, 13.5-16 mm.; width, 7-8 mm. 

Forma farallonica.—Larger, somewhat dull, slightly longer ovate, 
thorax a little more transverse. Elytra more granulate laterally. In 
the females the elytra are scarcely more convex than in the male. 

Measurements—Males: Length, 16-16.5 mm.; width, 7-7.5 mm. 
Females: Length, 13-17.5 mm.; width, 7-8.5 mm. 


a REVISION OF ELEODIINI—BLAISDELL. 357 


Genital characters, male—Edeagophore elongately oblong-ovate, 
flattened and very feebly arched. 

Basale elongate, oblong; surface moderately convex, sides feebly 
arcuate. 

A picale rather small, triangular, moderately and rather evenly con- 
vex above, without groove; sides broadly and moderately sinuate, 
apex acute and somewhat slender; base subacutely lobed at the middle 
and sinuate laterally. 

Sternite transversely parabolic in outline. Each lobe triangular; 
external border quite straight or feebly arcuate in basal two-thirds, 
thence subtruncate or feebly arcuate to apex, the latter obtuse; in- 
ternal margin straight and oblique; surface feebly convex, very 
sparsely punctate and setose, sete rather short, moderately long about 
the apical margin. Membrane not setose across the bottom of the 
sinus, the latter deep and triangular. 

Female.—Genital segment not broadly triangular, less than a half 
longer than wide, dorsal surface of each valve slightly obliquely de- 
clivous externally, feebly setose. 

Valvula (Plate 5, fig. 4).—Dorsal plate elongate oblong, slightly 
narrowing from base to apex; surface smooth, feebly convex, very 
sparsely and feebly setose in apical fourth; external border moder- 
ately arcuate, scarcely at all expanded; apical margin not defined 
from the apex, scarcely wider than the apex at base, angle obsolete ; 
internal border broadly and feebly sinuate. Apex small and more or 


Jess membrano-chitinous, feebly setose. 


Appendage small mammilliform with a pencil of quite long sete 
at tip. Fossa small and at the middle of the external surface of the 
apex, 

Basal prominences scarcely evident. 

Superior pudendal membrane longitudinally rugulose, and attain- 
ing the apical fourth of the dorsal plate. 

Ventrolateral surfaces—Body triangular, surface lines straight 
when viewed longitudinally, quite evenly convex laterally. Sub- 
marginal groove small, slightly widened apically beneath the scarcely 
explanate external border of the dorsal plate. Internal margins of 
the valves contiguous in basal two-thirds; genital fissure narrowly 
fusiform in apical third, and the inferior pudendal membrane is not 
visible. | 

Habitat—California (region about San Francisco Bay; San 
Joaquin and Sacramento valleys; hills along the coast in the central 
part of the State), Farallone Islands, 

Number of specimens ‘studied, 375. 

Type in the Eschscholtz collection ? 

Type-locality.—Near San Francisco. 


358 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Salient ty pe-characters.—Antenne filiform. Thorax rounded, base 
narrowed, punctate. Elytral dorsum with larger and slightly im- 
pressed punctures, sides granulate (Eschscholtz). 

Diagnostic characters.—Eschscholtz says that the thorax is a little 
broader than long, the greatest width being in front of the middle, 
sides strongly rounded and smaller behind; the apex is faintly 
emarginate; the humeri are prominent and the sharp epipleural mar- 
gin can be seen from above. The sides of the elytra are rugosely 
granulate. 

I have examples before me which have been identified by Casey, 
Fall, and Blanchard. 

A female compared by Mr. Blanchard with the specimen received 
by LeConte with others from Europe ® as typical parvicollis Esch- 
scholtz, is pronounced by him as “an exact copy” of LeConte’s 
typical specimen. Professor Fall’s and Colonel Casey’s specimens, 
while a little larger, present no notable differences. 

My typical specimen does resemble a large female scabrosa, but it 
is without the hairs upon the sides of the prothorax. It is well 
to note that my series of typical parvicoltis was taken from ledges 
upon the hills about San Francisco, and not about the sand dunes 
where scabrosa abounds. 

I do not agree with Doctor Horn in considering the large aggregate 
of specimens referable to parvicollis as one species without the recog- 
nition of well-marked races; in so doing I consider that it would be 
a retardation to our knowledge of these forms, and I shall in this 
paper consider planata, producta, and constricta as valid races. 

I have studied an immense series and feel warranted in so doing. 
My disposal of scabrosa will probably be a surprise to many, but it 
appears out of place when associated with parvicollis. 

In parvicollis the pronotum is rather less convex and the base more 
strongly constricted than in consobrina, where the sides of the elytra 
are more coarsely and tuberculately sculptured. 

Producta is oblong, flattened on the elytral disc, sides parallel with 
prominent humeri, which are strongly margined by the superior 
epipleural margin. 

Planata has the elytra less flattened, humeri less prominent, with 
the elytral sides evenly arcuate and more gradually narrowed behind. 

Parvicollis is shorter and more ovate. 

Producta inhabits the mountain regions chiefly and planata the 
valleys and foothills where there are oak groves and where smaller 
examples of producta are also found. Constricta occurs chiefly in the 
middle altitudes of the Sierra Nevadas as far north at least as Mount 
Shasta. These races overlap in their distributional areas; so do other 
races and species. 


@Horn’s Rev. Ten., p. 317. 


REVISION OF ELEODIINI—BLAISDELL. 359 


General observations—The mentum is variable, small, and para- 
bolic (male) or comparatively large and trapezoidal or triangulo- 
trapezoidal; surface very feebly convex at middle, and broadly, very 
feebly subfoveate laterally, finely but not distinctly punctate, not 
noticeably setose. 

The prosternum is not strongly protuberant with the coxw, and 
quite evenly arcuate antero-posteriorly between the coxe, sometimes 
more or less vertical behind; rarely submucronate. Mesosternum 
arcuately declivous and more or less concave. 

The metasternum laterally between the coxe is as long as the width 
of a mesotibia at apex. 

The abdominal process is quadrate (male) or slightly transverse 
(female) and about equal to (male) or a fourth (female) of its width 
broader than the metasternal salient. 

In the male the post-coxal part of the first abdominal segment is 
about equal in length to the second segment; the third is equal to 
that of the process and about a third longer than the fourth. 

In the female the post-coxal part of the first segment is about equal 
in length to the process, also to that of the third; the second is twice 
as long as the fourth. 

The tarsi are variable in length and stoutness; although moderate in 
length they are noticeably longer and slender in some males and 
females, and stouter in others, and this appears to be independent of 
size in the specimen. 

Tarsal formula: 


Pro. Meso. Meta. Metatibia. 
Male.—2} 3 Bh 5 
Female.—2 23 3 AG 


ELEODES PARVICOLLIS var. PLANATA Eschscholtz. 


Subfusiform-ovate to ovate, integuments feebly shining, twice as 
long as wide. Antenne long. Elytra depressed and granulate 
laterally. 

Head twice as wide as long, feebly convex, rather strongly and 
broadly impressed laterally, also more or less along the frontal suture, 
the latter rather feebly evident, supra-antennal convexities quite 
strong; surface rather densely, not very coarsely punctate, punctures 
denser laterally and on the epistoma. Antenne rather long, moder- 
ately slender, outer four joints slightly compressed and feebly dilated, 
third joint a little shorter than the next two combined, fourth ap- 
parently just the least longer than the fifth, the latter to the eighth 
inclusive equal in length, the eighth subtriangular and longer than 
wide, ninth triangular and as wide as long, tenth triangulo-oval, 
eleventh subovate and as long as wide. 


360 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Pronotum somewhat transversely oblong-oval, Be: at the middle, 
and more than a half wider than long; disc evenly and moderately 
convex, more or less flattened along the lateral margins, quite densely, 
more or less irregularly, and rather coarsely punctate, punctures not 
confluent, granulate on the impressed sides; apes evenly but not 
deeply emarginate, more or less obsoletely and finely margined; sides 
evenly and broadly rounded, strongly constricted at about basal 
eighth, thence somewhat oblique, straight, or sinuate to angle, mar- 
ginal bead thin and rather distinctly reflexed, becoming a mere line 
on the constriction; base truncate or feebly rounded, not strongly 
margined, a fifth to a sixth wider than the apex and a little greater 
than the length; apical angles obtuse and not prominent; basal angles 
slightly prominent. 

Propleure as in parvicollis. 

Elytra oval, widest at the middle, a third to a fifth longer than 
wide; base subtruncate, equal to or distinctly wider than the con- 
tiguous base of the prothorax; humeri obtusely rounded, sometimes 
prominent ; s¢des evenly arcuate, apex somewhat attenuate, not broadly 
rounded; disc depressed and moderately convex, strongly but not 
broadly rounded laterally, inflexed sides oblique and feebly convex, 
arcuately declivous posteriorly ; surface coarsely and irregularly pune- 
tate, somewhat foveolately so, punctures quite simple about the suture, 
series evident centrally when viewed longitudinally, not coarsely 
muricato-granulate laterally and about apex. 

E'pipleure distinctly dilated beneath the humeri, where the supe- 
rior margin is usually visible as an arcuate line; otherwise as in 
parvicollis. 

Legs comparatively slightly thicker than in parvicollis. 

Otherwise as in parvicollis. 

Male—FElytra at base scarcely wider than the contiguous pro- 
thoracic base; humeri scarcely prominent, sides thence evenly arcuate, 
apically gradually convergent so as to appear somewhat attenuate, 
the dise being gradually arcuately declivious behind. | Anterior tarsi 
with the first three joints bearing pubescent tufts beneath, the tuft on 
third joint sometimes obsolete; middle tarsi with the first two joints 
bearing small tufts at tips beneath. Otherwise as in parvicollis. 

Female—Elytra broadly oval; humeri prominent and rounded; 
base comparatively rather broad; sides evenly arcuate. 

Otherwise as in parvicollis. 

Measurements.—Males: Length, 15-16 mm.; width, 6.5-7.5 mm, 
Females: Length, 16-16.5 mm.; width, 7.8-8.5 mm. 

Edeagophore and sternite as in parvi- 


Genital characters, male. 
collis. 

Female.—Dorsal plate sometimes feebly concave and very sparsely 
setose. Apex chitinous and setose, sete a little longer than usual and 


REVISION OF ELEODIINI—BLAISDELL. 861 


abruptly terminating where the apical margin of the dorsal plate 
should be. Pencil of hairs at tip of appendage quite long. Other 
wise as in parvicollis. 

Tabitat.—California (Russian settlement, Fort Ross, Eschscholtz). 
My series is without definite locality data and were probably collected 
by Mr. G. W. Dunn. 

Number of specimens studied, 61. , 

Type in the Eschscholtz collection ? 

Type-locality.—Russian settlement, Fort Ross, California. 

Salient type-characters.—Thorax transverse, densely punctate ; sides 
rounded; base coaretate. Elytra depressed on the dorsum and fove- 
olato-punctate. Antenne subfiliform (Eschscholtz). 

Diagnostic characters —KEschscholtz says that the thorax is much 
broader than long, strongly rounded on the sides and with the nar- 
row margins more or less strongly reflexed. Elytra with the dorsum 
distinctly flattened (but convex) and not markedly declivous behind; 
side margins of the humeri very distinct; coarsely punctured at mid- 
dle, seriate and granulate laterally. 

The type must have been a male. My series, although not exactly 
typical, consists of both sexes. Mr. Blanchard has kindly made the 
necessary comparisons in Cambridge. and writes that a male (now 
before me) “is not quite typical, being less convex.” It must be 
noted, in order that the above descriptive remarks be not considered 
inconsistent, that the elytra are depressed and at the same time moder- 
ately convex. 

Typical parvicollis resembles scabrosa more than it does planata; 
the latter has the sides of the pronotum impressed, the elytra are more 
distinctly oval, and more gradually declivous behind in the male, 
and not as roughly sculptured; there is more or less evidence of a 
serial arrangement of the punctures on the dorsum; there the punc- 
tures are simple and the strial punctures are arranged in a straight 
line as usual, but greatly obscured by the very irregularly diffused in- 
terstitial punctures, both being of the same size. 

I can not discover any signs of a serial arrangement in the typical 
parvicollis, where the sculpturing is coarser and more scabrous. 

Producta is distinctly oblong, with the sides of the elytra parallel, 
the dorsum distinctly flattened and with the humeri very prominent 
in both sexes. 

Of course the extremes are more difficult to separate, but I consider 
them no more so than in some parts of the subgenus ./e/aneleodes. 

Two males before me are very coarsely and densely sculptured. 
One from Trinity County is especially so; the elytral punctures on 
the dorsum show evidence of serial arrangement of the interstitial 
punctures; both series are of the same size and submuricate, laterally 
the punctures are muricato-tuberculate, approaching consobrina and 


362 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


hornii, from which it is separated by the epipleure being dilated 
beneath the humeri and the superior margin distinctly visible from 
above, there forming the sharp arcuate margin of the rounded humeri. 
A Humboldt County specimen is similar, but less strongly developed. 

General observations —The mentum, pro- and mesosternum, are 
as in parvicollis. 

The metasternum laterally between the cox is as long as the width 
of a mesotibia at the middle. 

The abdominal process is subquadrate and a third (male) to a 
fourth (female) of its width broader than the metasternal salient. 

In the male the post-coxal part of the first abdominal segment is 
equal in length to that of the process, also to that of the third; the 
second is one-half longer than the fourth. 

In the female the second segment is equal in length to the process; 
the third to the post-coxal part of the first and a third longer than the 
fourth. 

The tarsi are variable in thickness, but moderate in length. Usually 
more slender in the male. 

Tarsal formula: 


Pro. Meso. Meta. Metatibia. 
Male.—24 3 4 5 
Female,—2 3 3 5 


ELEODES PARVICOLLIS var. PRODUCTA Mannerheim. 


Distinctly oblong to oblong-ovate, thorax transverse; elytra de- 
pressed, sides parallel, humeri prominent and obtusely rounded. 

IIead twice as wide as long, feebly convex, slightly impressed 
laterally and along the frontal suture, the latter more or less obsolete, 
densely, evenly, or irregularly and somewhat coarsely punctate, punc- 
tures denser laterally. Antennw somewhat long, stout, very feebly 
compressed and scarcely dilated in outer four joints, third joint about 
as long as the next two combined, fourth just the least longer and 
stouter than the fifth, the latter to the eighth, inclusive, equal in 
length, the eighth not noticeably wider than the preceding two, ninth 
triangulo-oval, tenth feebly transversely oval, tenth ovate. 

Pronotum transversely oblong-oval, widest at the middle, a half 
to three-fifths wider than long; disc moderately convex, feebly im- 
pressed laterally so as to appear somewhat dilated, rather coarsely 
and somewhat densely punctate, denser and somewhat finely granu- 
late laterally on the impressed area; apea more or less feebly and 
broadly emarginate, more or less obsoletely beaded; sides broadly 
and quite strongly arcuate to basal eighth, thence constricto-sinuate 
to the angles, at times quite abruptly and tubularly constricted, with 
sides parallel to the angle, bead thin and reflexed, a mere line on the 
basal construction ; base truncate or very feebly rounded, and usually 


—_ ae 


Ee = 


REVISION OF ELEODIINI—BLAISDELL. 363 


obsoletely beaded, a fifth wider than the apex and greater than the 
length ; apical angles somewhat distinct or obtuse and not prominent ; 
basal angles rectangular or minutely prominent. 

Propleure smooth, not densely granulato-muricate and more or 
Jess rugulose. 

Elytra oblong-ovate, arcuately attenuate from the middle pos- 
teriorly, about a third longer than wide; base broad and truncate, 
distinctly wider than the contiguous base of the prothorax; humeri 
obtusely rounded and more or less prominent; s/des parallel in basal 
one-half, thence arcuate to apex, somewhat attenuate in apical third 
or fourth, apex moderately rounded; disc distinctly flattened, more 
or less moderately convex, and arcuately declivous posteriorly, areu- 
ately, but not broadly, rounded laterally; surface densely, somewhat 
coarsely punctate, punctures very irregular, at times granuliferous or 
simple on the dorsum, more or less muricato-granulate or distinctly 
subrugose laterally and on apex. 

Epiple ure distinctly dilated beneath the humeri where the superior 
margin is coarse and arcuately prominent, as viewed from above. 
Otherwise as in parvicollis. 

Abdomen rather densely and somewhat coarsely punctate, usually 
distinctly rugulose. 

Legs stout and comparatively moderate in length. Anterior tibial 
spurs rather stout, the anterior more so than the posterior. Other- 
wise as-in parvicollis. 

Male.—Oblong, somewhat narrow, antenne just reaching beyond 
the base of the prothorax. First two joints of the protarsi mod- 
erately thickened at tip beneath and there clothed with small tufts 
of piceo-flavate pubescence; the first joint of the mesotarsi has a small 
tuft also. Otherwise as in parvicollis. 

Female—Oblong to ovate, somewhat broad. Antenne scarcely 
attaining the prothoracic base. Otherwise as in parvicollis. 

Measurements.—Males: Length, 15-17.5 mm.; width, 7-7.5 mm. 
Females; Length, 14.5-18.5 mm.; width, 7.5-8 mm. 

Genital characters, male-—As in parvicollis. 

Female.—The genital segment as in planata, apparently more equi- 
laterally triangular. The apex of each valve is distinctly chitinous, 
rounded at tip, with a slight tendency to become concave beneath. 
These characters foreshadow those observed in Metablapylis. 

Tabitat.—California (Giant Forest, Tulare County, Fuchs and 
Hopping; Yosemite, Mariposa County; Tuolumne County; Calaveras 
County, Big Trees, elevation, 4,702 feet; Amador and Eldorado 
counties). 

Number of specimens studied, 300. 

Type in the Mannerheim collection ? 


364 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Type-locality California; no special locality is mentioned by 
Mannerheim. Collector, D. Blaschke. 

Salient type-characters—Oblong. Thorax with the sides rounded, 
thinly margined, base narrowed and equal to the apex; dorsum 
thickly punctate and densely granuliferous at the sides. Elytra 
oblong-ovate, attenuate behind the middle, dorsum gently punctate, 
the punctures granuliferous, on each side asperate, humeri rounded 
and anteriorly prominent and somewhat produced (Mannerheim). 

Diagnostic characters —According to Mannerheim it differs from 
parvicollis in being much larger and narrower, the thorax longer 
and broader; elytra at the periphery asperate, chiefly distinctly 
subrugose. 

Examples compared with specimens in the LeConte collection by 
Mr. Blanchard were unhesitatingly referred to the present distinetly 
marked race. 

Mannerheim’s type was undoubtedly more asperately sculptured 
on the elytra than the majority of specimens before me, but there are 
many that exactly agree with his description. 

These more strongly sculptured examples must be separated from 
the var. constricta; in the latter the head and thorax are much more 
coarsely and less densely punctured, the elytra are quite strongly 
asperate. 

Any student ought not to have any trouble in recognizing this dis- 
tinctly oblong variety, the elytra flattened, with the base broad, the 
humeri prominent and distinctly margined by the epipleure. 

A fact already brought to the reader’s notice is that all //leodes— 
with very few exceptions—vary in form, and the heterotypes may 
simulate (be amphitypical of) other species so closely as to often be 
very difficult of recognition. 

For example, a specimen before me could easily be mistaken for 
consobrina, but the broader thorax with impressed sides and more 
prominent humeri renders the diagnosis certain. 

Smaller and less characteristic forms will be found in the lower 
levels westward even as far as the coast. 

General observations —The mentum is usually distinctly parabolic 
in outline and variable in size, the surface is very broadly and obso- 
letely foveate laterally, centrally more or less feebly convex, finely 
sculptured, and not noticeably setose. 

Prosternum not very prominent ventrally with the cox: often 
arcuate antero-posteriorly, or feebly convex between the coxe and 
vertically truncate behind; rarely distinctly mucronate. 

Mesosternum variable, sometimes deeply excavated at middle and 
lobed laterally; usually more or less broadly concave. 

The metasternum laterally between the cox is as long as the width 
of a mesotibia at middle. 


REVISION OF ELEODIINI-——BLAISDELL, 365 


The abdominal process is quadrate (male) or transverse (female) 
and equal (male) to or a sixth (female) of its width broader than 
the metasternal salient. 

In the male the post-coxal part of the first abdominal segment is 
equal in length to that of the process; the second, twice as long as 
the fourth; the third, a third longer than the fourth. In the female 
the post-coxal part of the first is equal to the second; the third to the 
length of the process and a half longer than the fourth. 

The tarsi are somewhat short and very stout. The anterior and 
posterior are apparently slightly compressed. The basal joint of each 
is noticeably stout, the others gradually smaller, so that each looks 
somewhat attenuate. 

Tarsal formula: 


Pro. Meso, Meta. Metatibia. 
Male.— 2} 3 3% 5} 
Female.— 2 3 34 54 


ELEODES PARVICOLLIS var. CONSTRICTA LeConte. 


Oblong-ovate, twice as long as wide; surface dull to shining, head 
and thorax more or less coarsely punctate. Elytra flattened on the 
dorsum, rugoso-punctate, laterally sculptured with setigerous recli- 
nate tubercles; humeri strong and more or less anteriorly prominent ; 
punctures throughout minutely setigerous. 

Head twice as wide as long, frons feebly convex, impressed later- 
ally and along the frontal suture, which is usually not evident; sur- 
face coarsely and densely punctate, impunctate areas frequently ob- 
served about the middle, laterally the punctures are closely crowded. 
Antenne somewhat long, third joint scarcely as long as the next two 
combined, fourth to the eighth, inclusive, subequal in length, the 
eighth more or less triangular, ninth and tenth orbicular in outline, 
or the former orbiculo-triangular and the latter feebly transversely 
oval, eleventh short-ovate. 

Pronotum transversely oblong-oval, widest at the middle, a little to 
a third wider than long; disc moderately and evenly convex, scarcely 
impressed laterally, coarsely and more or less irregularly punctate, 
impunctate areas frequently observed about the middle, intervals flat 
and smooth, laterally densely rugoso-granulate along the margins; 
apex feebly emarginate to subtryncate, obsoletely margined; sides 
evenly and strongly arcuate to basal seventh, there strongly con- 
stricted with sides parallel to base, sometimes less strongly arcuate or 
very feebly oblique behind the middle to constriction, margin fine, 
thin, and reflexed, a mere-line on the constriction; base very feebly 
rounded or truncate, obsoletely margined, a fourth or less wider than 
the apex, and about equal to the length; apical angles distinct and 


366 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


obtuse, not prominent, sometimes feebly rounded; basal angles rec- 
tangular. ~ 

Propleurw sparsely and evenly muricato-granuiate, more or less 
rugulose, especially about the acetabular convexities. 

Elytra oblong ovate, widest at the middle, a fifth to a fourth longer 
than wide; dase subtruncate and distinctly wider than the contigu- 
ous prothoracic base; humeri obtusely rounded and more or less 
prominent; sides evenly arcuate, apex rather narrowly rounded; 
disc more or less depressed, rather feebly to somewhat strongly con- 
vex, rather narrowly to somewhat broadly rounded at the sides, arcu- 
ately declivous behind; surface more or less rugoso-punctate cen- 
trally along the suture, punctures sometimes rather simple, more or 
less strongly muricato-tuberculate laterally and on apex. 

Epipleure with the superior margin arcuately margining the 
humeri and more or less distinctly visible from above; otherwise as 
in parvicollis. 

Abdomen somewhat thickly punctate and rugulose, most so about 
the coxe, punctures moderate in size. 

Otherwise as in parvicollis. 

Male—Somewhat narrow; antenne just slightly reaching beyond 
the prothoracic base. Elytra arcuately or also slightly obliquely 
declivous behind and ordinarily attenuate in apical fourth; anterior 
tibiz not noticeably constricted at base; first two joints of the 
anterior tarsi feebly thickened at tips beneath with quite small tufts 
of brownish yellow pubescence; that on the first joint of the meso- 
tarsi somewhat obsolete. Otherwise as in parvicollis. 

Female.—More or less robust; antennze about attaining the pro- 
thoracic base. Otherwise as in parvicollis. 

Measurements.—Males: Length, 12.5-15 mm.; width, 5.9-6.5 mm. 
Females: Length, 13.5-15.5 mm.; width, 6.5-8 mm. 

Genital characters, male——<As in parvicollis. 

Female.—The dorsal plate is not defined at apical margin from the 
apex, the latter subchitinous and subacute at tip; also somewhat de- 
flexed, so that the junction with dorsal plate appears somewhat promi- 
nent. Otherwise as in parvicollis. 

Habitat—California (Sacramento, LeConte; Tallac, Riverton, 
Strawberry Valley, 6,000 feet elevation, Eldorado County; Big Trees, 
elevation 4,702 feet, Calaveras County). 

Number of specimens studied, 53. 

Type in the LeConte collection. 

Type-locality.— Sacramento, California; collector, Mr. Wittick. 

Salient type-characters—Somewhat shining, prothorax with the 
sides strongly rounded, tubulately constricted at base, finely mar- 
gined ; dorsum thickly and laterally rugoso-punctate. Elytra slightly 
convex, sides greatly rounded; humeri a little produced, rounded, and 


— 


REVISION. OF ELEODIINI—BLAISDELL. 367 


margined; dorsum densely subseriately punctate, laterally and on 
apex scabrous from reclinate granules. Antenne slightly incrassate 
externally. Length .52. (LeConte.) 

Diagnostic characters —This race is remarkable on account of the 
coarse.and dense elytral sculpturing, somewhat like that observed in 
cordata, but with the muricate character predominating; the head 
and thorax is more coarsely punctured than in parvicollis oryits 
races above considered; in size the punctures approach those in 
cordata, but the intervals are perfectly flat and smooth and small 
impunctate areas are usually present. 

The humeri are prominent and the elytral dise flattened on the 
dorsum in the males especially. It is interesting to note that the 
females are apt to be robust and in form quite like cordata, so much 
so that most of them have heretofore been placed with that species; 
the broader prothorax with sides broadly rounded separates con- 
stricta; in rotundipennis the prothorax is rugoso-punctate. 

I have carefully examined the genital segments, and find the dorsal 
plate not at all angulate at the apical margin, but possessing the char- 
acteristics of the parvicollis group. 

It would be very instructive and an advance in our knowledge if 
it were possible to determine why the heterotypes of many species 


or races mimic other more or less closely related species. Is it ata- 


vism, hybridism, or environment / 

If LeConte’s description of constricta be compared with the one 
given above, they will be found to agree in most points. 

A specimen before me, and which has been compared with the type 
by Mr. Blanchard, is pronounced by him as undoubtedly that form 
described by LeConte, from a specimen collected near Sacramento, 
The thorax is “ ante basin tubulatim constricto.” 

It is possible that I am making a mistake in applying LeConte’s 
name to this mountain form. I have not seen any specimens from 
the vicinity of Sacramento. It is possible, of course, that the rivers 
may have carried individuals down from the higher altitudes, and 
in this way populated this particular region and elsewhere, for that 
matter. 

I will quote LeConte’s remarks upon constricta: 

Related to HE. producta, but the base (prothoracic), though not narrower 
than the apex, is much more constricted than in that species, so that the sides 
become impressed. The elytra are broader, more rounded on the sides, the 
humeri are less broadly produced and conspicuously margined, and the sides 
are more rough with small, reclinate tubercles. This species agrees with the 
description of EB. parvicollis, given by Mannerheim,? except that the base of the 
thorax is not narrower than the apex; Eschscholtz does not, in his description, 
mention that the sides of the thorax are scabrous, but adds that the base of 


* Bull. Mosc., 1843, p. 271. 


368 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


the thorax is nearly as wide as the widest portion of the thorax, whereby the 
latter appears small. A specimen from the St. Petersbu® Museum, sent me as 
a type, is not allied at all to this species, but rather to LZ. cordata. 

It is well to mention here that the typical parvicollis resembles a 
large scabrosa, and the latter can easily be confused with cordata. 
A collector with any diagnostic acumen at all will not make such 
mistakes. 

To satisfactorily work out the races of parvicollis has given me 
more trouble and consumed more time than any other whole sub- 
genus, and while I feel that I might be wrong in some of my conclu- 
sions, I have Mr. Blanchard’s careful comparisons to support my 
opinions, which are, besides, based upon the study of large series of 
specimens and much careful dissection of material. 

General observations —The mentum, pro- and mesosterna as in 
parvicollis. The metasternum laterally between the coxe is as long 
as the width of a mesotibia at apical third. 

The abdominal process is subquadrate and about a third of its 
width broader than the metasternal salient. 

In the male the process is equal in length to the post-coxal part of 
the first segment and equal to the second; the latter is a fourth longer 
than the third, which is slightly longer than the fourth. 

In the female the post-coxal part of the first segment of the. 
abdomen is equal in length to the third, the latter equal to that of 
the process; the second is twice as long as the fourth. 

The tarsi are slender and moderate in length. 

Tarsal formula: 


Pro. Meso. Meta. Metatibia. 
Male.—2 _ 5. © err see Teas 
Female.—14 24 3 44 


ELEODES HOPPINGII, new species. 


Oblong-oval, about twice as long as wide, somewhat depressed, 
surface rather dull, punctures of the head and thorax scarcely coarse ; 
elytral sculpturing rather fine and dense, muricato-granulate lat- 
erally, punctures throughout finely setigerous. 

Head \ess than twice as wide as long, rather evenly convex, feebly 
impressed laterally, obsoletely so along the frontal suture, the latter 
not evident; surface densely punctured. Antenne short, attaining 
the posterior fourth of the prothorax, moderate in stoutness, feebly 
compressed and slightly widened in the outer four joints, third joint 
equal to the next two combined, fourth to the eighth inclusive sub- 
equal in length and scarcely longer than wide, the eighth triangular, 
ninth and tenth feebly transversely oval, eleventh short-ovate. 

Pronotum transversely oblong-oval, widest at the middle, less to 
ubout a half wider than long; dése scarcely moderately and evenly 


REVISION OF ELEODIINI—BLAISDELL. 369 


convex, densely and somewhat finely punctate, finely granulate lat- 
erally; apex feebly and evenly emarginate, obsoletely margined ; 
sides broadly, evenly, and rather strongly rounded to basal twelfth, 
thence sinuate to the basal angles, finely beaded almost to the base, 
the latter very feebly rounded, not distinctly margined, a seventh to 
a fifth wider than the apex and about equal to the length; apical 
angles obtuse and not rounded; basal angles subrectangular.  » 

Propleurew sparsely and distinctly granulate and more or less 
rugulose. 

Elytra oval, about a third longer than wide, widest at the middle; 
base subtruncate and a little wider’than the contiguous prothoracic 
base; humeri obtusely rounded and not prominent; sides evenly 
arcuate, gradually converging in apical fourth, apex rather narrowly 
rounded ; dise rather feebly and evenly convex, somewhat flattened on 
the dorsum, rather narrowly, evenly, and somewhat strongly rounded 
laterally, arcuately declivous behind; surface densely and rather 
finely sculptured, punctures about simple centrally along the suture, 
muricato-granulate laterally and on apex. 

Epipleure moderately wide, not dilated beneath the humeri, 
superior margin not visible from above; surface very sparsely and 
finely punctate. 

Sterna not coarsely punctato-rugulose. 

Parapleure densely and not distinctly punctate. 

Abdomen more or less shining, quite densely punctate and rugu- 
lose, most so about the coxe. 

Legs short and moderate in stoutness. Anterior femora mutic; 
anterior tibial spurs similar in the sexes, the anterior a little stouter 
than the posterior; anterior tarsi dissimilar in the sexes. 

Male—Somewhat narrow. Elytra arcuately declivous posteriorly. 

Abdomen feebly oblique, moderately convex and impressed at middle 
of the first two segments. First two joints of the protarsi not notice- 
ably thickened at tips beneath, each bearing a minute, brownish, and 
rather pointed tuft of pubescence; first joint of the mesotarsi without 
tuft, groove distinct. 
Elytra arcuately and vertically de- 
clivous behind. Abdomen horizontal and rather strongly convex. 
First joint of the protarsi noticeably thickened at tip beneath, groove 
quite distinct. 

Measurements.—Males: Length, 12.5-13 mm.; width, 5.5 mm. Fe- 
males: Length, 13 mm.; width, 6.2 mm. 

Genital characters, male—Edeagophore elongately oblong-ovate, 
acute at tip and slightly arched. 

Basale oblong, two and a half times longer than wide, evenly con- 
vex, sides parallel and slightly arcuate, 

59780—Bull. 683—09——24 


370 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Apicale triangular, a little longer than wide; surface moderately 
convex, without groove; sides strongly and broadly sinuate, apex 
appearing attenuate; base obtusely lobed at middle, feebly and 
broadly sinuate laterally. 

Sternite transverse; each lobe triangular; surface feebly convex, 
not very densely punctate, setose; setee moderate in length. gradually 
longer about apex; external border rather evenly arcuate and oblique, 
apex introrsely placed and obtuse; internal border more or less sinu- 
ate and oblique. Membrane not setose across the bottem of the sinus, 
the latter triangular. 

Female.—Genital segment somewhat depressed, about equilaterally 
triangular, dorsal surface feebly ogival, feebly setose. 

Valvula—Dorsal plate oblong, three times longer than wide, sides 
just the least converging apically; surface more or less feebly con- 
cave, glabrous, very sparsely and finely punctured, sete short; ex- 
ternal border feebly arcuate; internal border more or less sinuous; 
apical margin more or less arcuately rounded and only apparently 
defined from the semi-chitinous apex; the latter small, short and 
scarcely more prominent posteriorly than the appendage, blunt and 
more or less rounded, sparsely and finely setose with longer hairs, tip 
with a pencil of moderately long hairs. Fossa large, in the external 
surface of the apex. 

Appendage short triangulo-conical, with a pencil of long hairs at 
tip. 

Superior pudendal membrane longitudinally rugulose, sttaining 
the apical fourth of the dorsal plate. 

Basal prominences not evident. 

Ventrolateral surfaces plane centrally to base, surface lines straight 
when viewed longitudinally, laterally rounded; surface glabrous, with 
a few coarse punctures scattered about, continuous with the semi- 
chitinous apex, not at all concave before the apex, the latter finely 
setose. Submarginal groove feeble at base, but well developed in 
apical two-thirds beneath the feebly dilated external edge of the 
dorsal plate. Internal margins of the valves contiguous in basal 
two-thirds; fissure narrow and in apical third, about closed, and the 
inferior membrane is not visible. 

Habitat.—California (Eldorado County, on the way to the summit 
above Riverton, July). 

Number of specimens studied, 3. 

Sexitypes in my own collection. 

T ype-locality—Western slope of the Sierra Nevada Mountains, 
above Riverton, Eldorado County, California. 

Salient type-characters.—See general description. 

Diagnostic characters.—This species differs from parvicollis and 
its races in the more oval and somewhat depressed form, finer and 


REVISION OF ELEODIINI—BLAISDELL. 371 


denser sculpturing. The prothorax, while it retains the same form 
observed in parvicollis, is more broadly rounded and more briefly 
sinuate at the base, the marginal line nearly attains the angle. Con- 
stricta, which occurs in the same region, is much more strongly and 
coarsely sculptured, the prothorax is strongly constricted at base. 

The wider pronotum, smaller and less robust form, distinguishes 
hoppingti from consobrina. 

The pubescent tufts on the protarsi are smaller than in any other 
species of Blapylis known to me. The antenne are also shorter than 
in any other closely related species. 

The female before me was compared with types in Cambridge by 
Professor Fall, and found to be quite different from anything there. 

I have named it in honor of my friend, Ralph Hopping, in recogni- 
tion of many favors. 

General observations.—The mentum is moderate and parabolic in 
outline, slightly triangular in the males; surface finely and not dis- 
tinctly punctate, broadly and feebly foveate laterally. 

The prosternum is not strongly protuberant ventrally with the 
coxw, arcuately rounded antero-posteriorly between the same, and 
not in the least mucronate. 

The mesosternum is arcuately declivous and feebly concave. 

The metasternum laterally between the cox is as long as the 
width of a mesotibia just before the apex. 

The abdominal process is subquadrate (male) or slightly trans- 
verse (female), and about a fourth of its width broader than the 
metasternal salient; it is also equal to the length of the post-coxal 
part of the same abdominal segment; it is also equal to the length 
of the second segment, the latter being twice as long as the fourth; 
the third is one-half longer than the fourth. It is quite unusual for 
these relative proportions of parts to correspond in the two sexes. 

The tarsi are slender and of moderate length. 

Tarsal formula : 


Pro. Meso. Meta. Metatibia. 


Male.— 13 2} 23 + 
Female.— 1} 2 2} 34 


ELEODES CLAVICORNIS Eschscholtz. 


Eleodes clavicornis Escuscuo.itz, Zool. Atlas, IL], 1833, p. 11.—MANNER- 
HEIM, Bull. Soc. Nat. Moscow, XVI, 18438, p. 269.—LeConTer, Proc. Acad. 
Nat. Sci. Phila., 1858, p. 182.—Horn, Trans, Amer, Phil. Soc., XIV, 
1870, p. 315. 

Eleodes impressicollis RoHEMAN, Freg. Eugen. Resa, Ins., 1858, p. 90. 


Oblong-ovate, about twice as long as wide, small, pronotal dise 
not noticeably transverse and with moderately long erect setw along 


372 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


the sides, also upon the propleura. Punctures setigerous distinetly 
throughout. ° 

Head twice as wide as long, feebly convex, more or less impressed 
laterally and along the frontal suture, the latter more or less obsolete ; 
surface irregularly and sparsely punctate at middle, also on the 
vertex, more densely and coarsely so laterally and on the epistoma. 
Antenne rather long, quite slender, outer four joints feebly com- 
pressed and distinctly and gradually incrassate, third joint scarcely 
as long as the next two taken together, fourth just the least longer 
than the fifth, the latter to the seventh inclusive subequal in length, 
the seventh slightly broadened and subtriangular, eighth just the least — 
shorter and more or less triangular, ninth triangulo-oval and feebly 
transverse, tenth transversely oval, eleventh transversely suboval. 

Pronotum slightly transversely suboval, widest at the middle, about 
or a little more than a third wider than long; disc moderately convex, 
distinetly arcuately declivous laterally and at the apical angles, 
sparsely, distinctly, and very irregularly punctate, with scattered 
impunctate areas, punctures coarser and denser laterally on the de- 
clivities, where each bears a moderately long erect seta; apex sub- 
truncate in circular are, finely and more or less obsoletely margined ; 
sides more or less evenly and moderately arcuate, somewhat straight 
and convergent posteriorly, feebly or not at all sinuate before the 
basal angles, marginal bead fine and entire; base very feebly rounded 
and finely beaded, about a fifth to a fourth wider than the apex and 
equal to or slightly greater than the length; apical angles obtuse, not 
in the least prominent, and more or less feebly rounded; basal angles 
obtuse or minutely rectangular and just the least prominent. 

Propleure opaque, sparsely punctate, each puncture with a moder- 
ately long, erect, and brownish seta. 

Elytra oval, not over one-half longer than wide, widest at the mid- 
dle; base subtruncate and about equal to the contiguous prothoracic 
base; humeri rounded; sides evenly arcuate, apex not very narrowly 
rounded and not in the least produced; disc almost evenly convex 
from side to side, dorsum scarcely flattened, arcuately declivous 
posteriorly ; swrface punctate, punctures coarse and impressed, rather 
dense and scarcely arranged in series, at the sides and apex the 
punctures are more irregular, submuricate, each with a rather short 
suberect seta, which become less evident on the dorsum. 

Epipleurw rather wide, gradually narrowing from base to apex, 
superior margin rather feebly and broadly sinuate beneath the 
humeri and not visible from above; surface smooth, sparsely and 
coarsely punctate. 

Sterna obsoletely punctate and rugulose, sparsely pubescent, hairs 
short and erect. 


REVISION OF ELEODITNI—BLAISDELL. 378 


Parapleure somewhat sparsely punctate, hairs short. 

Abdomen horizontal, glabrous and shining, very sparsely punctate, 
hairs short. 

Legs comparatively moderate in length and rather slender. 
Anterior femora mutic; protibial spurs about similar in the sexes, the 
anterior a little longer than the posterior; tarsi dissimilar in the 
sexes. . 

Male.—Body rather slender. Antenne reaching a short distance 
beyond the base of the prothorax. Elytra evenly arcuately decli- 
vous posteriorly. Abdomen moderately convex, first two segments 
distinctly impressed at middle. Protibial spurs subequal, slender, 
and acute. Protarsi with the first two joints very feebly thickened 
at tips beneath, each bearing a small tuft of silken pubescence that 
obliterates the groove. 

Female.—Body rather robust. Antenne reaching just slightly 
beyond the prothoracic base. Elytra rather broadly oval, arcuately 
and almost vertically declivous posteriorly. Abdomen quite strongly 
convex. Anterior spurs of the protibie a little longer than the pos- 
terior, both slightly thickened and acute. Anterior tarsi with the 
first joint slightly thickened at tip, groove interrupted by a few paler 
spinules, that are transversely arranged on tip. 

Measurements —Males: Length, 10.2-12.5 mm.; width, 4.2-5 mm. 
Females: Length, 12-13 mm.; width, 5.2-6 mm. 

Genital characters, male—KEdeagophore elongately oval and 
scarcely arched. 

Basale oblong, not three times as long as wide; surface evenly con- 
vex, with sides feebly arcuate. 

Apicale subequilaterally triangular; surface rather strongly con- 
vex, without groove; sides distinctly sinuate in apical two-thirds, 
arcuate in basal third; apex more or less attenuate and acute; base 
broadly lobed at middle and feebly sinuate laterally. 

Sternite transverse. Each lobe triangular, with the external bor- 
der more or less sinuate in basal half, thence more or less obliquely 
truncate to apex, the latter narrowly rounded; internal border more 
or less straight; surface feebly convex, sparsely punctate in apical 
half, setose, sete quite long, still longer on apical margin. Mem- 
brane not setose across the bottom of the sinus, the latter rather 
broad. 

Female.—Genital segment elongately triangular, superior surface 
plane. 

Valvula (Plate 4, fig. 11).—Dorsal plate narrow and elongately 
oblong; surface plane, glabrous, very sparsely and obsoletely pune- 
tate, setose; sides subparallel; apical margin evenly rounded and 
clearly defined from apex, the latter small and finely setose; external 
lobe obsolete. 


374 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Appendage very short mammilliform, with three or four long hairs 
at tip; fossa on the basal and external surface of apex, small. 

Superior pudendal membrane long, attaining the apical sixth of 
the dorsal plate and longitudinally rugulose. 

Basal prominences not evident, evenly rounded about base. 

Ventrolateral surfaces.—Body quite evenly convex from side to 
side, surface lines straight when viewed longitudinally, feebly con- 
cave laterally before the apex; apex finely setose. Submarginal 
groove distinct and moderate beneath the very narrowly prominent 
external margin of the dorsal plate. Internal margins of the valves 
contiguous in basal third; fissure rather long and quite narrowly 
fusiform; inferior pudendal membrane not visible. 

Habitat.—California (common about the sand dunes under vege- 
tation, along the coast of central California; Doctor Horn reports 
it as extending northward to the head of the Sacramento Valley; as 
far south as Los Angeles County). 

Number of specimens studied, 600. 

Type in the Eschscholtz collection. 

Type-locality.—California near San Francisco. 

Salient type-characters—Oblong-ovate. Antenne slender and 
subclavate. Pronotal disc not noticeably transverse, arcuately de- 
clivous laterally, with moderately long erect sete on the declivous 
portions; apical angles obtuse, feebly rounded; sides rather evenly 
rounded, rather straight and converging posteriorly, just the least 
sinuate before the basal angles. Elytra coarsely punctate, with slight 
evidence of a serial arrangement when viewed longitudinally, 
sides more irregularly and submuricately punctate. The punctures 
are setigerous throughout. 

Diagnostic characters.—Clawecornis is a very distinct species and 
quickly recognized by its small size and setose sides of the pronotal 
disc. No other species of the subgenus Blapylis has the propleure 
so distinctly pubescent. Scabrosa is the most closely related species, 
and differs in its distinctly larger and transverse pronotum, the discal 
sides of which are not in the least declivous, although set with very 
short erect sete. Clavicornis has the elytra rather coarsely and 
densely punctate, the punctures are impressed, the sculpturing is 
quite like that observed in scabrosa. 

The punctures are noticeably setigerous throughout. The antenne 
are usually feebly clavate. 

The species named by Boheman as impressicollis is simply speci- 
mens which have two thoracic fovee. Such accidental variations 
are common (see p. 27): sometimes only one fovea is present; this 
variation is frequently observed in other species. 


REVISION OF ELEODITNI—BLAISDELL. 875 


General observations —The mentum is small and more acute at 
apex than in other species; the surface is rather strongly convex at 
middle, finely and not strongly punctured, not noticeably setose. 

The prosternum is variable. Feebly widened or subparallel be- 
hind the coxw, more or less arcuate antero-posteriorly, frequently 
subtruncate behind or feebly mucronate; surface usually distinctly 
grooved at middle. 

Mesosternum vertically arcuate or obliquely declivous, and more or 
less distinctly concave. 

The metasternum laterally between the coxe is as long as the width 
of a metatibia at apex. 

The abdominal process is quadrate (male) or slightly transverse 
(female) and a fifth (male) toa fourth (female) of its width broader 
than the metasternal salient, subequal in length to the second segment 
(male) or equal to that of the post-coxal portion of the first segment 
(female). 

In the male the post-coxal part of the first segment is a little shorter 
than the process and about a half longer than the fourth. 

In the female the second segment is about twice as long as the 
fourth. 

The protarsi are about two-fifths (male) or a half (female) of 
their length shorter than a mesotarsus. 

The mesotarsi are about a seventh (male) of their length shorter 
or subequal (female) to a metatarsus. 

The metatarsi are about a fourth (male) or about two-thirds 
(female) of their length shorter than a metatibia. 


ELEODES SCABROSA Eschscholtz. 


Bleodes scabrosa Escuscnortz, Zool. Atlas, III, 18338, p. 11.—MANNERHEIM, 
Bull. Soc. Nat. Moscow, XVI, 1843, p. 272.—LeConTer, Proc. Acad. Nat. 
Sci. Phila., 1858, p. 182.—Horn, Trans. Amer. Phil. Soc., XIV, 1870, 
p. 317, 


Suboblong-ovate, somewhat robust, shining, about twice as long as 
wide; pronotum transverse and with very short erect sete laterally 
on the dise and propleure. 

ITead twice as wide as long, feebly convex, usually impressed lat- 
erally and along the frontal suture, the latter more or less evident. 
coarsely and irregularly punctate, frequently with impunctate areas 
on the frons and vertex, punctures crowded laterally and on the epis- 
toma. Antenne moderate in length and stoutness, outer three or four 
joints slightly compressed and noticeably dilated, third joint about 
equal in length to the next two combined, fourth and fifth subequal, 
sixth and seventh apparently just the least shorter and subequal, 
eighth a little shorter, broader, and subtriangular, ninth and tenth 
transversely oval, eleventh ovate, apparently broader than long. 


876 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Pronotum transverse, widest at the middle, about a half wider than 
long; disc moderately and evenly convex, not déclivous laterally, 
coarsely and irregularly punctate, impunctate areas usually present, 
laterally the punctures are denser, becoming granulate and scabrous, 
each puncture with an erect and very short seta for a short distance 
from the side margins; apea truncate or feebly emarginate in cir- 
cular arc, obsoletely beaded; s¢des very evenly and rather strongly 
arcuate to basal eighth, there feebly constricted and sinuate, thence 
parallel or feebly convergent to the basal angles, marginal bead fine 
and usually entire; base feebly rounded, marginal bead not evident, 
slightly wider than the apex, equal to or slightly greater than the 
length; apical angles obtuse and scarcely rounded; basal angles 
obtuse or subrectangular, not at all prominent. 

Propleure smooth, very sparsely punctate, punctures submuricate, 
each with a moderately short and erect seta, more or less regulose. 

Elytra oval, about a third longer than wide, widest at the middle; 
base subtruncate, wider than the contiguous prothoracic base; hwmeri 
obtusely rounded or subangulate; sides evenly arcuate, apex rather 
narrowly to somewhat broadly rounded and not in the least produced ; 
disc moderately and evenly convex on the dorsum, strongly but not 
very broadly rounded laterally, arcuately declivous behind; surface 
punctate, punctures coarse, moderately impressed, diffused, without 
order, sometimes a slight serial arrangement evident when viewed 
longitudinally, punctures simpler on the dorsum, denser and muricato- 
granulate with a tendency to coelscence in transverse rug laterally 
and on apex; the punctures are minutely setigerous. 

Epipleure shining, rather wide, gradually narrowing from base to 
apex, superior margin more or less broadly sinuate beneath the 
humeri; surface submuricately punctate. 4 

Sterna densely punctate and somewhat rugulose. 

Parapleure coarsely and more or less densely punctate. 

Abdomen smooth and shining, evenly and sparsely punctate, first 
segment on and about the process densely sculptured; fifth segment 
more densely punctate at the periphery; each puncture with a short 
seta. 

Legs moderate in length, somewhat stout; anterior femora mutic;: 
anterior tibial spurs quite similar in the sexes, the anterior slightly 
longer and stouter than the posterior, both acute; tarsi dissimilar in 
the sexes. 

Male.—Somewhat robust. Antenne scarcely reaching beyond the 
prothoracic base. Elytra arcuately and feebly obliquely declivous 
posteriorly. Abdomen feebly oblique and moderately convex, first 
two segments more or less feebly impressed at middle. Anterior tarsi 
with the first joint shghtly thickened at tip beneath and bearing a 


REVISION OF ERLEODITNI—BLAISDELL. 3877 


small subacute tuft of yellowish pubescence enveloped by a few ordi- 
nary spinules, groove distinct on the second joint. 

Female.—Robust. Antenne not quite attaining the prothoracic 
base. Elytra rather broadly oval, arcuately and vertically declivous 
behind. Abdomen horizontal and rather strongly convex. Anterior 
tarsi with the first joint transversely thickened ‘at tip beneath, and 
with a transverse line of piceous spinules upon the same. 

Measurements.—M ales: Length, 10-12 mm.; width, 4.5-5.8 mm. 
Females: Length, 12-13 mm.; width, 6—-6.8 mm. 

Genital characters, male.—Edeagophore flattened oblong-ovate and 
feebly arched. 

Basale oblong, about two and one-fourth times longer than wide, 
surface evenly convex, the sides parallel and very feebly arcuate. 

Apicale triangular, surface evenly and not strongly convex, without 
groove; sides arcuate in basal third, thence broadly and quite strongly 
sinuate to apex, the latter subacute; base lobed at middle, slightly 
sinuate laterally. 

Sternite parabolic in outline. Each lobe triangular; external bor- 
der rather evenly and not strongly arcuate; internal border quite 
straight; apex somewhat introrsely placed and rather narrowly 
rounded; surface very feebly convex, glabrous and shining, sparsely 
punctate and setose in apical half, setae moderate in length, longer 
about apex, membrane not setose across the bottom of the sinus, the 
latter deep and triangular. 

Female.—Genital segment triangular, elongate, about twice as long 
us wide, surface plane, not conspicuously setose. 

Valvula (Plate 5, fig. 22).—Dorsal plate very elongate, oblong, 
about four times longer than wide; surface plane, glabrous, very 
sparsely punctate and setose, sete fine and inconspicuous; sides par- 
allel and nearly straight; apical margin parabolically rounded. 
Apex very small, slightly longer than the appendage, quite narrow 
and finely setose. 

Appendage mammilliform, about half as large as the apex, with a 
pencil of about five hairs at tip; fossa at base of the external surface 
of the apex. 

Superior pudendal membrane reaching to the apical fourth of the 
_ dorsal plate and longitudinally rugulose. 

Basal prominences not evident. 

Ventrolateral surfaces feebly convex from side to side, glabrous, 
sparsely setose in apical moiety. Each valve very feebly and broadly 
concave apically, submarginal groove broad beneath the laterally 
prominent external border of the dorsal plate, and quite evanescent 
before the fossa, which is distinctly lateral in position. Internal 
margins of the valves contiguous in basal two-thirds; fissure apical 
and rather narrow. Inferior pudendal membrane not visible. 


378 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Habitat—Plentiful about sand dunes along the coast of California, 
northward to Oregon. 

Number of specimens studied, 400. 

Type in the Eschscholtz collection ? 

Ty pe-locality—San Francisco, California. 

Salient type-characters.—Thorax transverse, densely punctate, sides 
rounded and scabrous. Elytral punctures on- the dorsum simple, 
granulate laterally. Antenne clavate (Eschscholtz). 

Diagnostic characters—The following characters seem to show 
conclusively that scabrosa is more closely related to clavicornis than 
to parvicollis with which it has for a long time been associated by the 
more recent authorities: the sculpturing is quite similar in the two 
species, the setose sides of the pronotal dise and propleuree, clavate 
antennie, elongate and narrow dorsal plates of the valves of the geni- 
tal segment in the female characters not observed in parvicollis; the 
sculpturing is also much coarser than is observed in the latter species. 

Scabrosa differs from clavicornis in having the antenn shorter, 
a coarser and denser cephalic and pronotal punctuation, a much 
broader pronotum, which is not so declivous at the sides and more 
strongly sinuate before the basal angles, the very short discal sete, 
those of the propleure being noticeably shorter; the male is also 
more robust. 

The present species is smaller than the average individuals of 
parvicollis. So, in view of these differential facts I am forced to 
restore this species of Eschscholtz’s to specific standing. 

The elytral sculpturing is quite similar to that observed in cordata, 
but in the latter species the pronotal sides are more strongly rounded 
anteriorly and more suddenly narrowed behind, becoming subangu- 
late at times; the pronotal punctuation is much coarser and the inter- 
vals convex, the pronotum is more cordate in the one and more trans- 
versely oval in the other. I am indebted to Professor Fall for com- 
parisons made with the LeContian specimens at Cambridge. 

General observations.—In scabrosa the mentum is larger and more 
broadly rounded at apex than in clavicornis (compare figs. 5 and 6 
of Plate 8); usually quite parabolic in outline, sometimes subtri- 
angular; surface convex along the median line and shghtly sub- 
foveate laterally, moderately punctate, setee not noticeable. 

The prosternum is quite prominent ventrally with the coxe, 
rounded antero-posteriorly between the same and not mucronate. 

The metasternum laterally between the cox is as long as the 
width of a mesotibia at apex. 

The intercoxal process of the abdomen is quadrate (male) or more 
or less transverse (female), and a fourth to a third of its width 
broader than the metasternal salient; the post-coxal portion of the 


REVISION OF ELEODIINI—BLAISDELL. 379 


first segment is slightly shorter than the process; the second segment 
is about twice as long as the fourth. 

In the male the third segment is about equal in length to that of the 
process and about a third of its own length shorter than the second. 

In the female the second segment is about equal in length to the 
process, and the third is about a fourth of its length shorter than the 
second. 

The protarsi are abeut three-eighths (male) or a third (female) of 
their length shorter than a mesotarsus. 

The. mesotarsi are about four-elevenths (male) or a fourth of 
their length shorter than a metatarsus. 

A metatarsus is about a third (male) or three-fifths of its length 
shorter than its metatibia. 


ELEODES CORDATA Eschscholtz. 


Bleodes cordata Escuscnortz, Zool. Atlas, IIT, 18383, p. 12.—MANNERHEIM, 
Bull. Soc. Nat. Moscow, XVI, 1848, p. 272.—-LEConTE, Proc. Acad, Nat. 
Sci. Phila., 1858, p. 182.—Horn, Trans. Amer. Phil. Soc., XIV, 1870, 
p.. 317. 

Hleodes tuberculata Escuscnourz, Zool. Atlas, III, 1883, p. 12.—MANNER- 
HEIM, Bull. Soc. Nat. Moscow, XVI, 18438, p. 274. 

Fleodes intricata MANNERHEIM, Bull. Soc. Nat. Moscow, XVI, 1843, p. 273. 

Bleodes stricta LeContr, Reports of Explor. and Surveys, 47th Parallel, 
XII, Appendix No. 1, 1857, p. 50. 

var. rotundipennis, LECoNTE, Reports of Explor. and Surveys, 47th Parallel, 
XII, Appendix No. 1, 1857, p. 50. 

Moderately robust, subovate, more or less shining, about twice as 
long as wide, prothorax strongly constricted at base, pronotum con- 
fluently and very coarsely punctate; elytra rugosely muricato-tuber- 
culate. 

Head about twice as wide as long, feebly convex, lateral impres- 
sions not strongly marked, frontal suture not evident, very coarsely 
and confluently punctate. Antenne moderate and somewhat stout, 
outer four joints very feebly compressed and very slightly widened, 
third joint equal in length to the next two combined, fourth scarcely 
longer than the fifth, the latter to the eighth inclusive subequal, 
eighth subtriangular, ninth transversely oval and frequently slightly 
robust, tenth transversely oval, eleventh short ovate and usually 
slightly smaller than either of the two preceding joints. 

Pronotum subcordate, about a half wider than long, widest at 
about the middle; disc moderately convex, more or less declivous at 
apical and basal angles, coarsely, densely, and more or less confluently 
punctate, interstices more or less convex; apea feebly emarginate 
to truncate, obsoletely margined; sides more strongly rounded in 
front, narrowing rather suddenly behind the middle, causing them 
to appear somewhat angulate, frequently more evenly arcuate to the 


380 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


constriction, which is usually at basal sixth, thence straight and 
parallel to the basal angles; the fine margin is ¥ery feebly. reflexed 
and obsolete on the constriction; base truncate in circular are, finely 
or obsoletely margined, about equal to the length and slightly to a 
fourth wider than the apex; apical angles obtuse, not in the least 
prominent anteriorly and more or less rounded; basal angles rec- 
tangular and not prominent. 

Propleurw more or less sparsely granulate and rugulose. 

Elytra rather broadly oval, about a fourth longer than wide, widest 
at the middle; base truncate and wider than the contiguous pro- 
thoracic base; humeri rounded, scarcely at all angulate; sides evenly 
arcuate, apex obtuse and moderately rounded; disc moderately convex 
on the dorsum, rather strongly and evenly rounded laterally, strongly 
and arcuately declivous posteriorly, generally quite vertically so; 
surface densely, irregularly, and rugosely muricato-tuberculate, less 
strongly sculptured along the suture; punctures minutely setigerous. 

Epipleure moderate in width, slightly dilated beneath the humeri, 
superior margin feebly and very broadly sinuate in basal fourth and 
scarcely visible at humeri when viewed from above; surface smooth, 
sparsely and rather coarsely punctate. 

Sterna more or less densely punctato-rugose, punctures setigerous, 
noticeably so on the mesosternum; set short, black, and rather stiff. 

Parapleure densely punctate. . 

Abdomen smooth, sparsely and distinetly punctate; first segment 
punctato-rugose, the fifth rather densely punctured. 

Legs moderate in length and thickness and quite strongly seulp- 
tured, the punctures bearing short black sete. Anterior femora 
mutic in the sexes. Anterior tibial spurs quite similar, the anterior 
slightly longer than the posterior. Anterior and middle tarsi dis- 
similar in the sexes. 

Male—Moderately robust, somewhat narrow, antenne reaching to 
or just beyond the base of the prothorax. Elytra about a fourth 
longer than wide. Abdomen moderately convex, slightly oblique, 
broadly impressed at middle of the first segment. Spurs of the pro- 
tibiz quite slender and acute. First two joints of the protarsi clothed 
with flattened tufts of golden pubescence beneath; first joint of the 
middle tarsi with a well-developed and similar tuft at apical third 
beneath; pubescent joints not noticeably thickened at-tip. 

Female.—Robust. Antenne about reaching to the prothoracic base. 
Elytra broadly oval and about a fifth longer than wide. Abdomen 
quite strongly convex. Spurs of the protibix slightly thickened and 
acute. First joint of the protarsi slightly thickened at apex beneath, 
groove more or less distinct. 

Measurements —Males: Length, 13-15.2 mm.; width, 6—-6.8 mm. 
Females: Length, 12-17 mm.; width, 6.8-9 mm. 


REVISION OF ELEODIINI—BLAISDELL. 381 


Three forms may be recognized according to the character of the 
sculpturing : 

Forma sublevis.—Similar in form to the typical form, but smoother, 
with the sculpturing somewhat obsolete. 

Forma typica—Strongly and roughly sculptured; elytra rugosely 
punctate, the interstices between the punctures elevated and convex, 
and more or less confluent transversely. ‘ ; 

Forma intermedia.—Sculpturing more strongly muricato-tubercu- 
late, with the intervals between the punctures less convex and conse- 
quently less rugose, approaching pimelioides. 

Genital characters, male-—Edeagophore stoutly fusiform, feebly 
arched. 

_ Basale oblong, about twice as long as wide; surface evenly convex; 
sides feebly arcuate, and the apex quite deeply and triangularly 
emarginate. 

Apicale triangular, rather narrow, usually distinctly narrower at 
base than the basale at apex; surface strongly convex, without 


groove; sides nearly straight or feebly arcuate, slightly prominent at 


base; apex acute; base acutely lobed at middle, laterally feebly 
sinuate. 

Sternite subparabolic. Each lobe triangular; external border 
more or less straight or feebly and broadly sinuate in basal two- 
thirds, thence quite evenly arcuate to apical margin, angle nar- 
rowly rounded; internal margin feebly sinuate; surface very feebly 
convex, punctate and setose in apical half, sete moderate in length, 
not dense, longer toward and on apical margin. Membrane not 
setose across the bottom of the sinus, the latter more or less triangular 
and apparently not closed by the membrane. 

Female——Genital segment triangulo-trapezoidal, surface of the 
valves slightly declivous, setose. 

Valvula (Plate 4, fig. 6).—Dorsal plate oblong, feebly narrowed 
from base to apex. Surface plane or feebly concave, smooth, punc- 
tures scattered, each with a rather long flying seta; borders quite 
straight; apical margin oblique, angle obtuse and feebly rounded; 
internally not distinctly defined from the apex, the latter rather short 
and membranous, rounded at tip and clothed with moderately long 
setae. 

Appendage mammilliform, with a pencil of quite long hairs at 
tip. Fossa rather large at base of the external apical surface. 

Basal prominences not evident. 

Superior pudendal membrane longitudinally rugulose, and attain- 


ing the middle of the dorsal plate. 


Ventrolateral surfaces.—Body triangular, surface lines straight 
when viewed longitudinally, quite evenly convex laterally, sparsely 
setose in apical half. setae moderate and more or less flying. Sub- 


382 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


marginal groove well developed beneath the slightly expanded ex- 
ternal border of the dorsal plate and terminating at the fossa beneath 
the angle. Internal margins of the valves contiguous in basal two- 
thirds; fissure narrowly fusiform in apical third, and the inferior 
pnudendal membrane is not visible. 

Habitat——California. Forma sublaevis, about San Francisco Bay; 
Forma typica, central California, along the coast; San Joaquin and 
Sacramento valleys; Calaveras County; Tulare County, Ralph Hop- 
ping; Los Angeles County, Doctor Van Dyke; Forma intermedia, 
northern California, southern Oregon, western slopes of the Sierra 
Nevada Mountains. 

Professor Wickham in his “ Coleoptera of Colorado” says that 
Professor Snow has reported one specimen from Colorado. It is in 
all probability a variation of pémelioides. 

Number of specimens studied, 500. 

Type in the Eschscholtz collection ? 

T ype-locality — California.” 

Salient type-characters.—Ovate, robust, shining. Head and pro- 
notum coarsely and confluently punctate; thorax subcordate, apex 
feebly emarginate, angles obtuse; sides more strongly rounded in 
front, narrowing rather suddenly behind the middle, causing them to 
appear somewhat angulate; strongly constricted at base; angles 
rectangular. Elytra oval; humeri much rounded and obtuse; dise 
arcuately and vertically declivous posteriorly; surface densely and 
rugosely muricato-tuberculate, less strongly so along the suture. 

Diagnostic characters—Cordata differs from all other species in 
the subgenus Blapylis, except pimelioides, in having the sides of the 
prothorax more or less subangulate, This character is especially 
marked in the males. The prothorax is more evenly rounded behind 
the middle in most females, but always strongly constricted at base, 
giving it with the above characters a subcordate appearance. The 
marginal bead does not reach the base. 

The coarse sculpturing will cause it to be frequently confused with 
scabrosa, but the setose propleure of the latter will readily separate 
the two, besides the pronotal sculpturing is quite different in the 
two species. Clavicornis is much less strongly sculptured, and the 
sides of the pronotal dise are distinctly setose along, with the pro- 
pleure. 

In pimelioides and brunnipes the elytra are sculptured with rows 
of small and rounded tubercles; in the latter species the legs are 
brown or reddish. 

In studying a large series of cordata it is very evident that the 
sculpturing varies toward pimelioides, so that in the heterotypes it is 
quite difficult to tell where one species leaves off and the other begins: 


REVISION OF ELEODIINI—BLAISDELL. 883 


cordata acquires a rugoseness of sculpturing and pimelioides a tuber- 
culate form of elytral markings. 

They are to be considered as two divergent forms derived from a 
common ancestral ramus. 

Specimens from Oregon have the prothorax rounded on the sides, 
as in parvicollis and scabrosa (see rotundipennis). 

Specimens from southern California collected by Doctor Van Dyke 
are remarkable among those of the large series before me on account 
of their elongate form, measuring: Length, 15 mm.; width, 6.5 mm.; 
elytra more elongately oval (forma elongata). These specimens 
approach an unique male from Amador County, in which the form is 
elongate oblong, thorax as in cordata, antenne long and stout, and 
the elytra are distinctly tuberculate and not rugose. The whole 
habitus is that of a stout, oblong cordata (forma oblonga). Length, 
16 mm.; width, 7 mm. These are remarkable variations from the 
monotonous short, robust, normal form. 

Mannerheim’s types of éntricata were taken about Fort Ross, Cali- 
fornia, by Doctor Fischer, and I can not see that it differs in any 
essential characters to give it even varietal standing. Many females 
have the “ thorace transverso, lateribus valde rotundatis.” Ina large 
series of cordata a variation in the length and stoutness of the an- 
tenne and legs, independent of sex, may frequently be observed. 
The humeri vary considerably also, never prominent, but frequently 
subangulate and always more or less rounded. 

Tuberculata Eschscholtz is to be disposed of in the same way as 
intricata. For sticta, see p. 384. 

General observations—The mentum is usually more or less trian- 
gulo-trapezoidal and rather small, finely sculptured, feebly foveate 
laterally, and very slightly convex at middle. 

The prosternum is variable, usually rather prominent ventrally 
with the cox, and quite strongly arcuate antero-posteriorly, without 
a mucro; frequently a small mucro is present at middle of the pos- 
terior edge; rarely horizontal, subtruncate behind, and more or less 
feebly mucronate. 

The mesosternum is vertically arcuate and more or less broadly 
and deeply concave. 

The metasternum laterally between the cox is as long as the width 
of a mesotibia at middle. 

The abdominal process is subquadrate, equal in width (female) or 
a third (male) of its width broader than the metasternal salient. 

In the male the post-coxal part of the first segment is equal in 


length to the third; the second to that of the process and twice as long 
as the fourth and about a fifth longer than the third. 

In the female the post-coxal part of the first segment is equal in 
length to that of the process and also to the second, the latter being 


384 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


about twice as long as the fourth; the third is about one-half longer 


than the fourth. 
The tarsi are usually moderate in length and stoutness. 
Tarsal formula: 


Pro. Meso. Meta. Metatibia. 
Male.—2 24 3 43 
Female.—1# 24 3 4} 


ELEODES CORDATA var. ROTUNDIPENNIS LeConte. 


Pronotal sides evenly and broadly rounded (as in parvicollis), 
basal constriction short and the angles rectangular. 

Otherwise as in cordata. 

LeConte was very much in doubt as to the propriety of separating 
this. form even as a race. The constancy of form in the prothorax, 
which, taken in conjunction with the coarse elytral sculpturing, make 
it appear at first sight as a northern modification of scabrosa; this 
is not so; the propleure are without distinct sete, and besides 
the pronotal punctuation is quite different, being as in cordata; 
this is in fact the only superficial character which separates and pre- 
vents it from being naturally associated with the parvicollis section. 
I consider it a good race, and the series before me shows conclusively 
that it varies more toward typical cordata than pimelioides. 

Habitat—Oregon (Koebele); British Columbia (Victoria and 

North Bend, Hubbard and Schwarz) ; Washington (H. K. Morrison, 
Easton and ( )lympia). 

Number of specimens studied, 5 

Type in the cia collection. 

Ty pe-locality —* Oregon.” 

I have taken this variety at Oregon City, near Portland, Oregon. 

Stricta LeConte, according to the author, has the “ thorax less sud- 
denly but very much constricted at base, fully one-half wider than 
long; elytra more coarsely and less densely punctured,” and is to be 
taken as a synonym of the present race. 


ELEODES PIMELIOIDES Mannerheim. 


Eleodes pimelioides MANNERHEIM, Bull. Soc. Nat. Moscow, XVI, 1848, p. 
274: Mag. Zool., XIII, 1843, no. 129, fig. — LeConTe, Proc. Acad. 
Nat. Sci. Phila., 1858, p. 182.—Horn, Trans. Amer. Phil. Soc., XIV, 
1870, p. 318. 
Eleodes viator LeContr, Proc. Acad. Nat. Sci. Phila., 1858, p. 188. 
Eleodes subligata LeConte, Reports of Explor. and Surveys, 47th Parallel, 
XII, Appendix no. 1, 1857, p. 50. 
var. brunnipes Casey, Ann. N. Y. Acad. Sci., V, Nov., 1890, p. 402. 
Moderately robust, ovate, feebly shining to opaque, about twice as 
long as wide; prothorax more or less strongly constricted at base, 
dongely rugoso-punctate; elytra sculptured with small tubercles, 


which may be rounded or reclinate and more or less piliferous, 


REVISION OF ELEODIINI—BLAISDELL. 885 


Head densely punctate, antenne somewhat slender, ninth joint tri- 
angulo-orbicular to transversely oval, tenth more or less transversely 
oval. 

Pronotum subcordate to transversely suboval, widest near the mid- 
dle, a fourth to scarcely a half wider than long; sides evenly and 
quite strongly arcuate to basal seventh, or subangulate at middle, 
rounded in front and quite rapidly converging posteriorly and gin- 
uate at basal fourth, thence in each instance quite straight and par- 
allel to the basal angles; base equal to the length or in some males 
shorter than the length; apical angles obtuse, frequently not in the 
least rounded, at other times more or less so. 

Elytra quite broadly oval to subquadrate, widest at or behind the 
middle, a fourth to a third longer than wide; disc more or less 
deplanate on the dorsum, strongly, arcuately, and vertically declivous 
posteriorly; surface densely tuberculate, tubercles apparently ar- 
ranged in rows on the dorsum or irregular throughout; each bears a 
very short, black seta near apex; when arranged in rows there are 
very small muricate punctures scattered sparsely and irregularly 
between, always less distinct along the suture centrally; the tubercles 
are more or less rounded and shining, the interstices between more 
or less opaque. Otherwise as in cordata. 

Male.—First two joints of the protarsi with tuft of yellowish 
pubescence near tip beneath; that of the second joint is rather small; 
tuft on the first joint of the mesotarsi quite small. Tufts somewhat 
long and truncate at tips. Otherwise as in cordata. 

Female.—First joint of the anterior tarsi distinctly thickened at 
tip beneath. Otherwise as in cordata. 

Measurements—Males: Length, 12-14.5 mm.; width, 6-7 mm. 
Females: Length, 13-15.2 mm.; width, 6.5-8 mm. 

Genital characters, male.—As in cordata. 

Female.—Genital segment triangular, surface quite plane and 
slightly setose. 

Valvula (Plate 5, fig. 5).—Dorsal plate oblong, feebly or scarcely 
narrowed apically, slightly explanate externally; apical margin 
nearly transverse to feebly oblique, inwardly not defined from the 
surface of the apex, angle obtuse and more or less feebly rounded. 

Ventrolateral surfaces—Submarginal groove distinct and well 
defined beneath the expanded external border of the dorsal plate. 
Otherwise as in cordata. 

Habitat—California (Shasta, Siskiyou, Butte, Lake, and Hum- 
boldt counties) ; Oregon (The Dalles, Hubbard and Schwarz; Clacka- 
mas County, Charles Fuchs); Utah (Alta, Wasatch in July, Amer- 
ican Fork Canyon in July, Hubbard and Schwarz; Park City); Ne- 
vada (Elko, H. F. Wickham); Washington (Easton, C. V. ‘Riley; 


597TSO— Bull. 683—09——25 


386 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Pullman, Palouse, A. M. Warren; Olympia, E. C. Van Dyke; Walla 
Walla, Charles Fuchs; Everett, July, Wickham® and Van Dyke) ; 
Idaho (Camas Prairie, C. V. ier: Cceur d’Alene, June, H. F. Wick- 
ham); Montana (Helena, August, Hubbard and Schwarz) ; Colo- 
rado (Central City, Garland, C. V. Riley; Buena Vista, July, Hub- 
bard and Schwarz; Golden, Glenwood, Horse Fly Peak, Idaho 
Springs, Aspen, Fort Collins, Leavenworth Valley (9,000—10,000 
feet), Roaring Fork, Monument Gulch to Montrose, Little Willow 
Creek, Canyon of Big Blue, Montrose and adjacent mountains, H. F. 
Wickham in the “ Coleoptera of Colorado”); Wyoming (Natural 
Park, July) ; British Columbia (Vancouver Island, collection of Dr. 
EK. C. Van Dyke). 

Number of specimens studied, 90 

Type in the Mannerheim collection. 

Type-locality.—* California.” 

Salient type-character—Opaque. Thorax with the sides rounded, 
densely rugoso-punctate. Elytra subquadrate, apex strongly decli- 
vous, dorsum deplanate, densely granulate, granules reclinate, rough 
and piliferous (Mannerheim). 

Diagnostic characters.—In general form like cordata, from which 
it differs in having the elytra sculptured with small rounded tuber- 
cles. The form of the tubercles differs, in some specimens distinctly 
rounded, in others reclinate and often more or less muricate. In 
some males there is evidence of rows 

The series from Walla Walla, Wachiti atom; have the elytra depla- 
nate and subquadrate; these characters are less marked in the other 
specimens. In the race brunnipes the legs are brownish and more 
distinctly piliferous, at least in those specimens from Nevada. 

Viator, described by LeConte from Fort Bridger, is a male, with 
the elytral tubercles large and extending almost to the suture. 
Colonel Casey also considers that this form does not differ from 
ordinary males of pimelioides. 

Subligata is one of the forms associated with cordata by LeConte; 
he obtained specimens from Oregon. Doctor Horn considered it a 
synonym of the present species, and which is undoubtedly correct. 

For remarks upon variation in sculpturing see cordata. 

The legs are variable, usually more slender in the males and 
stouter in the females. The thorax and humeri vary as in cordata. 


ELEODES PIMELIOIDES var. BRUNNIPES Casey. 


Rather robust, moderately convex, coarsely, densely sculptured and 
dull, black; legs dark brown throughout. 

Head large, fully two-thirds as wide as the prothorax, coarsely, 
extremely. densely punctate and scabrous. Antenne longer than the 


REVISION OF ELEODITNI—BLAISDELL. 387 


head and prothorax, rather slender, third joint about four times as 
long as wide. 

Prothorax scarcely a third wider than long; disc moderately, 
evenly convex throughout, very coarsely, deeply, and confluently 
punctate; apex subtruncate; sides strongly angulate at the middle, 
thence very feebly arcuate to the apex and broadly sinuate to the 
base; base subtruncate and nearly equal to the apex in width; 
apical angles obtuse, not distinctly rounded, not at all prominent; 
basal angles right and not rounded. 

Elytra at base nearly a third wider than the contiguous protho- 
racic base, very slightly more than twice as long as the latter, ab- 
ruptly and obtusely rounded behind when viewed vertically; sides 
strongly arcuate behind, gradually convergent and straighter thence 
to the humeri, which are slightly obtuse but scarcely at all rounded; 
disc widest behind the middle, where it is from a third to two-fifths 
wider than the prothorax, feebly convex on the dorsum, strongly so 
laterally; surface coarsely, densely, asperately punctate, the asperi- 
ties arranged without trace of order. 

Legs moderate in length, somewhat slender. 

Male and female secondary sexual characters as in pimelioides. 

Measurements.—Length, 9.2-9.5 mm.; width, 4.34.6 mm, <A 
female taken at Verdi, Nevada, has a length of 15 mm. ; width, 7.5 mm. 

Genital characters, male—As in pimelioides. 

Female.—In the specimens referred to this species and which are 
not typical, the genital segment is triangular. 

Dorsal plate moderately narrow and elongately oblong; apical bor- 
der feebly oblique with the angle rounded. Apex rather short; fossa 
shallow. 

Appendage subspheroidally mammilliform. 

Habitat—Idaho (Casey); Wyoming (Casey; Green River, H. F. 
Wickham); Colorado (Buena Vista, July, elevation, 7,900—8,000 
feet, Wickham). Professor Wickham says that this race is not rare 
about the roots of gooseberry bushes at Buena Vista (“ Coleoptera of 
Colorado”). Nevada (Verdi, in April). 

Number of specimens studied, 8. 

Type in Colonel Casey’s collection. 

Ty pe-locality.—Idaho. 

Diagnostic characters—Colonel Casey’s specimens are small, and 
to me their most conspicuous difference from pimelioides is the color 
of the legs, which are brownish and sometimes quite reddish; in all 
probability this color is due to the failure of normal pigmentation. 
This same character is observed in tenebrosa (var. nana) and in 
Caseyt. 


388 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


The individuals of cordata and pimelioides vary greatly, and the 
present race appears to be but a well-marked heterotype; this rela- 
tion is quite clearly demonstrated in the large series before me. 

In examples of brunnipes from Pocatello, Idaho, and which agree 
quite well with the description, the sete are longer and more evident 
throughout. 

Otherwise as in pimelioides. 


ELEODES CASEYI, new species. 


Ovate, about twice as long as wide, legs reddish-brown, distinctly 
pubescent throughout. Pronotum rather coarsely, densely, and more 
or less rugosely punctate; elytra rather finely and densely granulate, 
granules submuricate laterally. 

Head about twice as wide as long, feebly convex, more or less im- 
pressed laterally and along the frontal suture, very densely and 
scarcely coarsely punctate. Antennw long, very feebly compressed 
and scarcely at all dilated in outer three or four joints, third joint 
scarcely as long as the next two combined, fourth to the eighth inclu- 
sive subequal, the eighth subtriangular, ninth somewhat triangulo- 
orbicular, tenth orbicular, eleventh ovate. 

Pronotum transversely oblong-oval, widest at the middle, about a 
third wider than long; disc moderately convex, declivous about the 
apical angles, very densely, rather coarsely, and more or less con- 
fluently rugoso-punctate, granulate laterally; apex truncate in cir- 
ear are, marginal bead nearly obsolete; sides evenly and moderately 
strongly arcuate, feebly sinuate in basal eighth and not noticeably 
constricted, marginal bead rather fine and almost entire; base trun- 
vate in circular arc, obsoletely beaded, equal in width to the apex 
‘and also to the length; apical angles obtuse and slightly rounded and 
not in the least prominent; basal angles nearly obtuse and distinet. 

Propleure opaque and densely granulate. 

Elytra oval, a fifth to a third longer than wide, widest at the 
middle; base subtrunecate, distinctly wider than the contiguous pro- 
thoracic base; hwmeri obtusely rounded and not in the least promi- 
nent; sides evenly arcuate, apex rather broadly rounded and not in 
the least produced; disc moderately convex on the dorsum, strongly 
rounded laterally, arcuately and more or less vertically declivous . 
posteriorly ; surface densely and irregularly granulate, granules more 
or less shining, simple and smaller along the suture, submuricate lat- 
erally and on apex, where they are larger and more discrete. 

Epipleure moderately narrow, not dilated beneath the humeri and 
gradually narrowing from base to apex, superior margins nearly 
straight beneath humeri when viewed longitudinally and not visible 
from above; surface smooth, finely and sparsely punctate. 


REVISION OF ELEODITNI—BLAISDELL. * 889 


Sterna finely punctato-rugulose. 

Parapleure densely punctate. 

Abdomen somewhat shining and densely punctate, punctures 
moderate. 

Legs rather short and moderate in stoutness, quite densely pune- 
tured; anterior femora mutic; anterior tibial spurs comparatively 
small, the anterior slightly longer and a little stouter than the pos- 
terior; similar in the sexes. Anterior tarsi dissimilar in the sexes. 
- Male——Somewhat narrow. Antenne reaching beyond the pro- 
thoracic base. Abdomen distinctly oblique, moderately convex and 
distinctly and broadly flattened at middle of the first two segments. 
Protarsi with the first two joints scarcely thickened at tips beneath, 
each with a very small but distinct tuft of pubescence, tufts subacute, 
grooves interrupted. Middle tarsi simple. 

Female.—Broader ovate. Antenne attaining the prothoracic base. 
Elytra rather broadly oval. Abdomen horizontal and rather strongly 
convex. First joint of the protarsi not noticeably thickened at tip 
beneath, groove entire. 

Measurements—Male: Length, 10 mm.; width, 4.5 mm. Female: 
Length, 13 mm.; width, 6.2 mm. 

Genital characters, male—KEdeagophore somewhat oblong-ovate, 
apicale greatly and suddenly narrowed in apical two-thirds. 

Basale elongate oval, moderately convex with the sides moderately 
arcuate. 

Apicale subtriangular in outline, suddenly narrowed at basal and 
middle thirds so that the apical two-thirds appear greatly narrowed, 
convex above, with a linear groove extending from near the tip to 
the base; sides very prominent in basal third, subangulate, thence 
strongly sinuate to the tip, the latter flattened, narrowly rounded, and 
somewhat reflexed at tip; base lobed at middle and feebly sinuate 
laterally. 

Sternite moderately transverse. Each lobe subtriangular, with the 
external border sinuate in basal half, thence arcuate to apex, the 
latter rather prominent, obtuse, and rather introrse; internal border 
more or less oblique and sinuous; surface feebly convex, rather 
coarsely punctate in apical three-fourths, punctures rather dense 
toward apex, sete moderately long especially about the apical mar- 
gin. Membrane not setose across the bottom of the sinus, the latter 
triangular. 

Female—Genital segment (Plate 6, fig. 3) rather equilaterally 
triangular, surface somewhat convex from side to side and feebly 
setose. 

Valvula—Dorsal plate somewhat oblong-triangular, sides feebly 
converging apically; surface glabrous, more or less concave, with a 
few scattered and fine punctures, each with a short and fine seta; 


390 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


external border feebly arcuate: apical border briefly truncate and 
internally angulate with the internal border, which is briefly and 
obliquely truncate, thence more or less sinuous to base. Apex short 
and somewhat deflexed, narrowly rounded at tip, finely setose, with 
few longer hairs. 

Superior longitudinal membrane longitudinally rugulose, reaching 
about to the apex of the dorsal plate. 

Basal prominences scarcely evident. 

Ventrolateral surfaces.—Body triangular, flattened, surface lines 
straight when viewed longitudinally and convex laterally; surface 
glabrous, rather coarsely and sparsely punctate. Submarginal groove 
strongly developed beneath the explanate external border of the 
dorsal plate, narrow at base, thence widening broadly, so as to in- 
volve the whole side before the fossa, at which it terminates. Ap- 
pendage showing distinctly beneath the apical border of the dorsal 
plate, apex finely setose. Margin of the valves contiguous in basal 
four-fifths, fissure in apical fifth and narrow. Inferior membrane 
not visible. 

Habitat—Nevada (Verdi, Blaisdell); California (Bodie, eleva- 
tion, 8,475 feet, July 1-7, Wickham). 

Number of specimens studied, 2. 

Sexitypes: Male in Wickham’s, female in my collection. 

Ty pe-localities—V erdi, Nevada, and Bodie, California. 

Diagnostic characters.—Caseyi, by its pubescence, will naturally 
be associated with Tricheleodes, and by being Amphidora-like in 
facies it will most likely be referred to hirsuta. In color and pubes- 
cence it reminds one of Stenotrichus. 

It is aberrant, and, for the present at least, will have to remain as 
representative of a group in the present subgenus, analogous in this 
respect to tibialis. Unfortunately the types are all the examples I 
have seen, and this fact accounts for the obscurity of the relationships. 

The grooved apicale of the edeagophore in the male and the pubes- 
cence in both sexes indicates an apparent approach at least toward 
Tricheleodes; on the other hand, the dorsal plate is more like that 
observed in Blapylis; the pudendal membrane is also unusually long. 
A larger series must be studied before any definite conclusion can be 
arrived at. The two first joints of the protarsi have pubescent tufts 
such as are observed in Blapylis, but not in 7'richeleodes. 

General observations —The mentum is rather small and subtri- 
angulo-trapezoidal in outline; the surface is more or less foveate lat- 
erally and densely punctured. 

In both types the prosternum is but feebly prominent ventrally 
with the coxe, slightly convex antero-posteriorly between the coxe, 
and with a subacute, small but distinct mucro behind, dorsad to which 
the posterior edge is vertically truncate. 


REVISION OF ELEODIINI-——BLAISDELL. 391 


The mesosternum is strongly prominent ventrally, arcuately and 
vertically declivous, but feebly concave. 

The metasternum laterally between the coxv is as long as the width 
of a mesotibia at middle. 

The abdominal intercoxal process is quadrate (male) or transverse 
(female) and a fourth (male) to a third (female) of its own width 
broader than the metasternal salient. 

The abdominal process is equal in length to the second segment ; 
the post-coxal part of the first is equal to the length of the third. 
In the male the third is but slightly longer than the fourth; in the 
female the third is about one-half longer than the fourth. 

The profemora are moderately clavate; the mesofemora feebly 
widened, with the superior and inferior surface lines slightly arcuate; 
the metafemora are subequal in width throughout, with the superior 
and inferior surface lines parallel. 

The tibial grooves reach nearly to the femoral base on all of the 
femora, but more or less evanescent internally on the meso- and 
metafemora; their margins are feebly developed and subasperate. 

The tibix are without tarsal grooves; the articular cavities are 
closed. The protibie are slightly carinate externally in basal half. 

The tarsi are comparatively moderate in length and stoutness. 

Tarsal formula: 


Pro. Meso. Meta. Metatibia. 
Male.—1} ee ey 34 
Female.—1} 24 23 4 


Subgenus METABLAPYLIS, new. 


The mentum has the middle lobe very small and the lateral lobes 
fully exposed (Plate 6, fig. 16). These characters appear unique in 
the genus /Jeodes, although in some parts the middle lobe is reduced 
in size, but never so completely and persistently. 

The anterior tarsi are similar in the sexes, simple beneath, the 
grooves entire. 

The anterior femora are more or less sinuate beneath in apical fifth, 
never dentate, although in nigrina the rudiments are present. The 
anterior tibiz are more or less constricted at base, and recalls similar 
characters in Blapylis and planipennis (Discogenia): this appears 
to be an atavistic character, and bears an inverse relation to the 
femoral teeth; it is at times feebly indicated in nearly all of the 
subgenera. 

Schwarzii forms an exception, and in the strict sense does not 
belong here; the first joint of the protarsi is distinctly and unmis- 
takably thickened at tip beneath, and can only be considered as a tem- 
porary constituent of the present subgenus until more can be learned 
of its relationships, and this depends upon more extensive collecting. 


392 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


It is also at variance in Blapylis, and rather than create another sub- 
genus it is best to do as I have done and retain it ‘As a section in the 
present aggregate. 

Tibialis, planipennis, and schwarzti are all disturbing elements in 
the subgenera in which they are placed, and their true position in all 
probability will only be known when the Mexican species shall have 
been carefully studied. 

The genital characters give the essential definition to the present 
subgenus: 

Male.—Apicale of the edeagophore more or less equilaterally tri- 
angular, more or less depressed, apex not in the least attenuate, the 
dorsal surface with a broad central membrano-chitinous area. In 
schwarzii a well-developed groove is present and the apex attenuate. 

Female.— Genital segment more or less triangular; apex chitinous, 
flattened, and moderately produced, but narrow and evenly rounded 
at tip, convex above and concave beneath. Dorsal plates of the 
valves oblong with the external edge converging apically, angle not 
evident. Appendage small and mammilliform, penicillate. Superior 
pudendal membrane rather broadly exposed and reaching to the 
apical fourth of the dorsal plate. Genital fissure nearly closed. 

In schwarzii the female characters are as discordant here as else- 
where. 

Distribution.—Of the three species and two races constituting the 
present subgeneric aggregate, nigrina has the most extensive distribu- 
tion, occurring in Oregon, Washington, Northern California, Nevada, 
Arizona, Colorado, New Mexico, Nebraska, Kansas, Idaho, and the 
Dakotas. It is to be inferred that it occurs in the intervening States 
not mentioned above, but representatives are not at hand. 

Dissimilis appears to be abundant in Arizona. It also occurs in 
New Mexico. 

Nevadensis, in Nevada, southward into Arizona and the Colorado 
Desert of eastern southern California. 

Perlonga is only known to me from Wyoming. 

Schwarzii occurs in Washington. It will be observed that J/eta- 
blapylis only occurs in California within its northern and eastern 
confines. Closely related species are undoubtedly plentiful to the 
southward in Mexico. 

Genealogy.—Very little can be expressed in regard to the origin of 
the species enumerated above. Too little is known of schwarzi; 
nigrina and dissimilis without doubt arose from a common ancestral 
trunk, and each again is diverging into well-marked races, as repre- 
sented by perlonga and nevadensis, while many no doubt are yet to be 
discovered. 

NVevadensis exhibits evidences of several incipient races. 


REVISION OF ELEODIINI—BLAISDELL. 393 


A synoptical tabulation of the genital characters will prove in- 
structive : 


Males.—Apicale equilaterally triangular and depressed, central 
area of the dorsal surface feebly concave and membrano- 
chitinous; apex not in the least attenuate______._-__________- 


nigrina, 
joel perlonga, 
dissimilis, 
ie nevadensis, 
Apicale triangular, not depressed, with a definite median longitudinal groove; 
Mi UES ee ee __-schwearzii. 
Females.—Valvular apex chitinous, flattened and moderately (nigrina. 
produced, tip rounded, convex above and concave beneath ; | var. perlonga. 
superior pudendal membrane longitudinally rugulose, reach- | dissimilis. 
ing at least to the apical fourth of the dorsal plate_--_--~-~- var. nevadensis. 
Valvular apex apparently chitinous, short, triangular, somewhat deflexed ; 
superior pudendal membrane reaching to the apical third of the dorsal plate, 
and not rugulose in the examples studied_____________________~_-- schwarzii. 


ANALYTICAL KEY TO SPECIES OF SUBGENUS METABLAPYLIS. 


First joint of the anterior tarsi not thickened at tip beneath, groove distinct 
and entire. 
Seulpturing submuricate on the elytral sides and apex. 
OS RET 0: Ce Se ae ee nigrina, 
Surface very bright and shining—males frequently very elongate. 
var. perlonga. 
Sculpturing smooth, not at all subasperate. 
Surface luster more or less shining; form fusiform-ovate to ovate; 
ventral surface not distinctly pubescent______-_--_-------~- dissimilis. 
Surface subopaque and alutaceous; both sexes typically of the same 
form, elongately cylindrical; ventral surface with rather long, sparse, 


SO GML Seka aes a led Ey oe SE a aa Rs ee var. nevadensis. 
First joint of the protarsi in both sexes, thickened at tip beneath, groove in- 
EN, ER Be as a a ae Os ae oe ee es schicarzii. 


ELEODES NIGRINA LeConte. 


Eleodes nigrina, LEContTE, Proc. Acad. Nat. Sci. Phila., 1858, p. 1S6.—Horn, 
Trans. Amer. Phil. Soc., XIV, 1870, p. 313. 

Elongately oblong-ovate to ovate, upiform, usually about three 
times longer than wide, subopaque; elytra muricato-granulate later- 
ally and on the apex. 

Head \ess than twice as wide as long, feebly convex, more or less 
impressed laterally and very vaguely along the frontal suture, the 
latter not evident; surface scarcely coarsely, irregularly, and densely 
punctate, punctures denser laterally and on the epistoma, where they 
bear minute sete. Antenne moderate in length and equal in the 
sexes, somewhat stout, very feebly compressed, and more or less 
slightly dilated in outer four joints; the third equal to the next two 
combined; fourth and fifth equal in length and apparently just the 
least shorter than the sixth or seventh, which are subequal; eighth 
about as wide as long and subtriangular; ninth and tenth more or 
less transversely oval; eleventh ovate, truncate at tip. 


894 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Pronotum subquadrate, widest at or just in front of the middle, an 
eighth to a fifth wider than long, less than twice af wide as the head; 
disc moderately to quite strongly convex, arcuately rounded laterally 
and at the apical angles; surface almost finely, densely, and irregu- 
larly punctate, becoming very densely granulate laterally; apex 
truncate to feebly emarginate in circular are and more or less obso- 
letely margined; sides evenly, broadly, but not strongly rounded, 
behind the middle slightly converging and straight or feebly sub- 
sinuate as viewed from above, but quite evenly rounded from apex 
to base as viewed from the side, marginal bead fine, entire, feebly 
limiting the disc from the propleure, the former not in the least 
dilated laterally; base subtruncate to feebly arcuate, occasionally 
broadly and feebly sinuate at middle, rather finely beaded, and a little 
wider than the apex; apical angles obtuse, not rounded nor promi- 
nent anteriorly; basal angles obtuse to subrectangular, never promi- 
nent, 

Propleure distinctly convex, moderately finely, more or less evenly 
and not very densely, granulately punctate, frequently rugulose. 

Elytra elongate oval, usually widest at the middle, sometimes be- 
hind the same, scarcely twice as long as wide; base subtruncate, 
scarcely to slightly wider than the contiguous base of the prothorax; 
humeri rounded, but not broadly so; sides more or less evenly arcuate, 
not strongly so, apex hardly produced and not broadly rounded; dise 
moderately convex, frequently somewhat depressed on the dorsum, 
rather obliquely attenuate in apical fourth and more or less arcuately 
declivous behind; swvface punctate, punctures somewhat fine, usually 
arranged without order, frequently more or less striate, almost simple 
about the suture on the dorsum, muricato-granulate laterally and on 
apex; when the punctures are serially arranged, those of the striz 
are closely placed and frequently almost larger than the interstitial 
punctures, which are more distantly spaced and quite regular, almost 
serial; laterally the punctures are always confused and minutely 
setigerous. 

Epipleure moderate in width, very feebly widened beneath the 
humeri, thence gradually narrowing to apex; superior margin very 
broadly and slightly sinuate in basal fourth, as viewed longitudinally ; 
surface sparsely, irregularly, and finely punctate, at times obso- 
letely so. 

Sterna quite densely punctate and rugose, sometimes strongly so. 

Parapleure irregularly and at times densely punctate. 

Abdomen horizontal, rather finely and almost densely punctate, 
ach puncture minutely setigerous; surface very finely or at times 
coarsely rugose, fourth and fifth segments generally less densely 
punctate and frequently more or less rufous. 


REVISION OF ELEODITNI—BLAISDELL. 895 


Legs moderate in stoutness and length. Femora rather densely 
punctate, each puncture with a very small appressed seta, the anterior 
mutic and more or less sinuate in apical fifth; tibie quite strongly 
muricate; anterior tibial spurs about equal and acute; anterior tarsi 
quite similar in the sexes, the first joint slightly thickened at tip be- 
neath, groove entire, the marginal and apico-marginal spinules rather 
prominent. 

Male—Somewhat narrow. Elytra with the sides not strongly 
arcuate, at times subparallel_ at the middle, and more or less arcuately 
and somewhat gradually obliquely declivous posteriorly; apex more 
or less obliquely narrowed. Abdomen slightly flattened at middle of 
first two segments, otherwise moderately convex. 

Female.—Somewhat robust, elytra more or less broadened, sides 
evenly arcuate, apex somewhat arcuately narrowed; dise arcuately 
and almost vertically declivous posteriorly, abdomen strongly convex. 

Measurements.—Males: Length, 20-25 mm.; width, 5.5-6.2 mm. 
Females: Length, 20-22.5 mm.; width, 8-8.5 mm. 

Genital characters, male.—Edeagophore somewhat oblong-fusiform. 

Basale oblong, sides more or less arcuate, scarcely arched and quite 
evenly convex, with a linear median groove in apical half. 

Apicale (Plate 1, fig. 3) triangular, surface more or less convex, 
generally depressed and more or less broadly membrano-chitinous at 
middle; sides feebly arcuate and more or less feebly sinuate near 
apex, the latter rather narrowly rounded and not recurved; base more 
or less acutely lobed at middle and sinuate laterally. 

Sternite transverse and semicircular in outline. Each lobe triangu- 
lar, with the external border quite evenly arcuate to the apex, the 
latter narrowly rounded; internal border slightly sinuous; surface 
feebly convex, rather densely punctate in apical half, setee moderately 
long and dense, most so at apex; membrane not setose across the 
bottom of the sinus, the latter more or less triangular. 

Female—Genital segment triangular to oblong-triangular in out- 
line, feebly setose, apex chitinous. 

Valvula (Plate 1, fig. 1).—Dorsal plate oblong, sides slightly con- 
verging apically, glabrous; surface more or less concave, feebly, 
sparsely, and irregularly punctate, punctures with fine reclinate sete, 
sides not reflexed; margins subparallel, the external nearly straight, 
or occasionally feebly arcuate to sinuous, and the internal more or 
less feebly sinuous or straight; apical margin more or less obliquely 
truncate, occasionally not well defined from the apical membrane 
internally; angle rounded but evident, occasionally with two or three 
sete. Apex well developed and moderately produced, flattened, more 
or less convex above, broadly and evenly rounded at tip, chitinous, 
with or without a few rather short sete; apical valvular membrane 
quite largely exposed within on apex and opposite the apical fourth 


396 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


of the dorsal plate, finely and densely setulose. Fossa largely visible 
from above and rather capacious, margins with a few scattered sete. 

Appendage moderate in size, mammilliform and penicillate, sete 
quite long, two or three extending beyond apex. 

Basal prominences slightly evident. 

Superior pudendal membrane longitudinally rugulose, rather 
broadly exposed and attaining the apical fourth of the dorsal plate. 

Ventrolateral surfaces (Plate 1, fig. 2) not swollen, surface flattened 
centrally to base and convex laterally. Apex concave beneath. Sub- 
marginal groove shallow and passing into a broad lateral and apical 
concavity; surface not noticeably setose; internal margins setulose at 
apex. Internal margins of the valves contiguous in basal half, the 
genital fissure in apical half, very narrow and nearly closed. 

Habitat—Oregon; Washington (Pullman, May, C. V. Piper) ; 
Northern California (Modoc County) ; Nevada (Verdi, April, abun- 
dant); Arizona (Williams, June, Barber and Schwarz; Humphrey’s 
Peak at the base, F. H. Snow): Colorado (mountains southwest of 
Montrose; Durango; South Park; San Luis Valley; Ouray; Veta 
Pass, 9.200 feet; Buena Vista; Colorado Springs; Florissant; Gar- 
land; Idaho Springs; Georgetown; near Long’s Peak; Fort Collins, 
Wickham’s List. South Park, elevation 8-10,000 feet, August, 
Golden, Dyar and Caudell; Chenney Gulch); New Mexico (Santa 
Fé, LeConte) ; Nebraska (Black Hills, LeConte; Sioux County, Coll. 
Univ. Nebraska) ; Idaho; Western Kansas; Dakota. 

Number of specimens studied, 350. 

Type in the LeConte collection. 

T ype-locality—Black Hills, Nebrask: 

Salient type-characters—Elongate, subopaque. Thorax subquad- 
rate, narrowed behind, sides broadly rounded, apex almost truncate, 
moderately convex, sharply and rather densely punctate. Elytra 
declivous behind and obliquely narrowed, irregularly punctate, pune- 
tures especially at the sides muricate and very briefly setiferous. 
Legs muricato-punctate; anterior femora obtusely sinuate (LeConte). 

Diagnostic characters.—Nigrina by its subopaque integuments and 
more or less muricate sculpturing differs from all the other species 
of the subgenus Ietablapylis. 

In the race perlonga the body surface is very bright and shiriing. 

In dissimilis the form is more ovate, usually less elongate and more 
depressed; the elytra are smoothly sculptured and the strie more 
evident and distant. 

Schwarzti more robust and the form is more like some Asid@, as 
convexa, the pronotum is less convex, more transverse, more dilated 
at the sides, so that the margin is distinct and prominent. 

Nevadensis is much more slender in every way, more or less aluta- 
ceous and opaque. 


P REVISION OF ELEODIINI—BLAISDELL. 397 


Nigrina is quite variable in form. There is before me a female 
that has the form of Awmeralis and was for years associated with that 
species, until I examined the genital segment, which revealed its true 
relationship. An examination of the tibial spurs and punctuation 
showed that it agreed with the present species. 

Another specimen, a male, has the form of a moderately large 
gigantea, the pronotum is strongly convex. ‘These two specimens are 
amphitypes. 

General observations—The middle lobe of the mentum is more or 
less triangular and comparatively small; the lateral lobes much ex- 
posed and larger; surface finely scabrous and punctate, not notice- 
ably setigerous. 

The prosternum is generally moderately protuberant ventrally with 
the cox, convex antero-posteriorly between the same and mucronate 
behind, the mucro is more or less subacute, the posterior edge may 
be vertically truncate. 

The mesosternum is arcuately declivous and more or less concave. 

The metasternum laterally between the cox is as long as the width 
of a mesotibia at apex. 

The abdominal process is quadrate and equal in width to the 
metasternal salient. 

In the male the post-coxal part of the first segment is equal in 
length to the second, twice as long as the fourth and a little longer 
than the third. 

In the female the post-coxal part of the first segment is a little 
longer than the third; the second twice as long as the fourth and a 
little longer than the post-coxal portion of the first. 

The profemora are moderately clavate and usually feebly com- 
pressed; the tibial grooves have flat floors, their margins more or 
less asperulate, the anterior somewhat arcuately laminate for a short 
distance internal to the apical sinuation; the margins converge to 
base. The mesofemora are not clavate, but somewhat parallel sided ; 
the grooves as on the profemora, but a little narrower and with the 
margins contiguous at basal fourth. 

The metafemora have the superior and inferior surface lines sub- 
parallel, grooves as on the mesofemora and with the margins evanes- 
cent at basal third. 

The tarsal grooves on the tibiz may be more or less indicated and 
asperately sculptured; the protibiz are more or less distinetly cari- 
nate externally in basal two-thirds; the articular cavities are closed. 

The tarsi are variable in length and stoutness. 

The relative lengths of the joints in each tarsus are apparently as 
in \Welaneleodes, where the test was fully applied. 


398 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Tarsal formula: 


> 
Pro. Meso. Meta. Metatibia. 
Male.— 2% 33 4 7 
Female.—2} 3t + 63 


ELEODES NIGRINA var. PERLONGA, new. 


Elongate, very bright and shining, sculpturing as in nigrina, but 
somewhat finer. Elytra obliquely narrowing in apical fourth, usu- 
ally distinctly pointed behind with apex narrowly rounded and sub- 
acute; disc somewhat flattened on the dorsum, arcuately and ob- 
liquely declivous behind. 

Otherwise as in nigrina. 

Measurements——Males: Length, 18.5-22 mm.; width, 6.5-7 mm. 

Habitat—Wyoming (June). 

Diagnostic characters—The present race differs mainly from 
nigrina in its very black and highly polished surface, as well as its 
very elongate form. Males are only known to me and I have seen 
eight examples all together. 


ELEODES DISSIMILIS, new species. 


Elongate, fusiform-ovate, or oblong-ovate to ovate, very black, mod- 
erately depressed, smooth, elytral striz rather distant. 

Head twice as wide as long, frons more or less plane, lateral im- 
pressions very feeble, frontal suture usually quite obsolete, distinctly 
but not very coarsely punctate, punctures dense laterally and on the 
epistoma. Antenne moderately long, somewhat stout, very feebly 
compressed, scarcely or very gradually widened in outer four joints, 
third joint about as long as the next two taken together, fourth 
evidently just the least longer than the fifth, the latter to the seventh 
inclusive subequal, eighth slightly shorter and subtriangular, ninth 
and tenth scarcely transversely oval, eleventh short-ovate, truncate 
at tip. 

Pronotum subquadrate, widest at or just a little in frout of the 
middle, about a sixth to a fifth wider than long, slightly depressed ; 
disc moderately and evenly convex, more or less arcuately declivous 
laterally, finely but distinctly, evenly and rather sparsely punctulate, 
punctules frequently somewhat obsolete in the central area. Apex 
slightly and evenly emarginate to subtruncate, more or less obsoletely 
margined; sides moderately and quite evenly arcuate, apparently 
more or less convergent behind the middle, where at times they may 
be somewhat straight or rarely feebly subsinuate, marginal bead fine; 
base feebly rounded and finely margined, scarcely wider than the 
apex; apical angles obtuse, feebly rounded or distinct, scarcely promi- 
nent anteriorly; basal angles obtuse and not in the least prominent. 


REVISION OF ELEODIINI—BLAISDELL. 399 


Propleurw distinctly convex, smooth, finely and sparsely or obso- 
letely punctulate, more or less rugulose. 

Elytra oval, about twice as long as wide, somewhat depressed, 
widest at or behind the middle; base subtruncate and distinctly 
adapted to the prothoracic base which it scarcely exceeds in width; 
humeri obtuse, angle distinct although small and not in the least 
prominent ; s#des evenly or more strongly arcuate behind the middle, 
more or less convergent in apical fourth, apex not broadly rounded; 
dise more or less moderately convex, rather evenly, quite strongly 
and somewhat narrowly rounded at the sides, more or less arcuately 
and obliquely declivous behind; surface punctate, punctures arranged 
in rather distant, not strongly marked and usually unimpressed 
series, the strial punctures are small and quite closely placed, the 
interstitial are serial and regular or irregular, fine and more distantly 
spaced, both series may be more or less irregular laterally and on 
apex, frequently of the same size, 

Epipleure comparatively rather wide, not noticeably dilated be- 
neath the humeri, narrowing gradually from base to apex, superior 
margin more or less visible at humeri when viewed from above; sur- 
face smooth, frequently slightly concave, dull in luster and quite 
impunctate, 

Sterna more or less feebly punctate and rugose, mesosternum 
slightly pubescent ; metasternum less so, hairs short. 

Parapleure not strongly punctate. 

Abdomen horizontal, smooth, sparsely and rather indistinctly pune- 
tulate, finely and inconspicuously piliferous, and more or less feebly 
rugulose. 

Legs moderate. in length, and in stoutness to somewhat slender. 
Anterior femora more or less sinuate beneath at apex, mutic; spurs 
of the protibix similar in the sexes, usually rather slender and acute, 
the anterior slightly longer than the posterior; protarsi similar in 
the sexes, the first joint not thickened beneath, groove distinct and 
entire. 

Male.—Fusiform-ovate to oblong-ovate, somewhat narrow. An- 
tenne reaching beyond the prothoracic base. Elytra rather gradu- 
ally and arcuately to almost obliquely and rather quickly declivous 
posteriorly. Abdomen very moderately convex, sometimes feebly 
impressed on first two segments. Anterior femora occasionally with 
the anterior margin of the tibial groove, feebly and broadly sub- 
dentate within the sinuation. 

Female.—Ovate, rather broad, antenne attaining the prothoracic 
base. Elytra rather broadly oval, arcuately but not vertically de- 
clivous posteriorly. Abdomen not very strongly convex. 

Measurements.—Males: Length, 14.5-19 mm.; width, 5.5-6.7 mm. 
Females; Length, 14.5-19 mm.; width, 6—-7.5 mm. 


. 


400 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Genital characters, male. 
fusiform, scarcely arched. 

Basale oblong, moderately convex with the sides feebly arcuate. 

Apicale subequilaterally triangular, moderately depressed, broadly 
impressed and membranous at middle; sides feebly arcuate and con- 
verging to apex, the latter not in the least attenuate and narrowly 
rounded; base subobtusely lobed at middle and feebly sinuate later- 
ally. 

Sternite rather short and slightly transverse, somewhat parabolic 
in outline. Each lobe short and triangular; external border straight 
or feebly sinuate in basal two-thirds, thence to apex feebly arcuate to 
subtruncate, apex obtuse; internal border oblique and feebly sinuate ; 
surface very feebly convex, coarsely and sparsely punctate in apical 
two-thirds, at apex rather densely so, sparsely setose, setee longer and 
denser on apex. Membrane not setose across the bottom of the sinus, 
the latter triangular and rather broad. 

Female.—Genital segment triangular, dorsal surface quite plane 
and feebly setose. 

Valvula (Plate 5, fig. 12).—Dorsal plate elongate oblong; surface 
plane or feebly concave, smooth and shining, with very few short 
setee scattered over the apical half; external border nearly straight, 
slightly converging apically and more or less explanate; apical mar- 
gin more or less defined from the apex and rounded with the angle; — 
internal border quite straight or feebly sinuous. Apex membrano- 
chitinous, feebly everted, slightly prolonged, tip rounded, convex 
above and feebly concave beneath, with two or three long sete near 
tip, valvular membrane within finely setose; fossa at base of the 
external surface. 

Basal prominences not evident. 

Appendage short mammilliform, smooth, with a tuft of some three 
or four quite long sete. 

Superior pudendal membrane longitudinally rugulose, attaining 
the apical sixth of the dorsal plate. 

Ventrolateral surfaces —Body triangular, ventral surface lines 
straight when viewed longitudinally, laterally very feebly convex 
in basal moiety, thence very slightly concave to apex: surface smooth 
and impunctate. Surface of apex finely setose, submarginal groove 
broad and shallow beneath the explanate external border of the dorsal 
plate. Internal margins of the valves contiguous in basal four-fifths, 
fissure in apical fifth and closed. Valves anteriorly produced in 
median line beneath. 

Habitat—Arizona (Fort Grant, July, Santa Rita Mountains, 
Chiricahua Mountains, Oracle, Hubbard and Schwarz; Williams, 
Barber and Schwarz, all in the U. S. National Museum collection. 


Edeagophore la oblong-sub- 


REVISION OF ELEODIINI—BLAISDELL. 401 


Oak Creek, September, E. G. Smith, collector, Warren Knaus) ; 
New Mexico (Highrolls, May). 

Number of specimens studied, 32. 

Type.—Cat. No. 12282, U.S.N.M.; co-types in my collection, 

Ty pe-locality.—Chiricahua Mountains (males), Santa Rita Moun- 
tains (female), Arizona. Collectors Hubbard and Schwarz. 

Salient type-characters.—Smooth; punctures simple; thorax gub- 
quadrate, widest just before the middle, somewhat narrowed behind; 
dise finely and rather sparsely punctulate; apex feebly and broadly 
emarginate; sides evenly and moderately arcuate, convergent behind 
the middle; apical angles obtuse, very slightly rounded at tip; basal 
angles obtuse, not rounded nor in the least prominent. Elytra widest 
at the middle; humeri obtuse with angle distinct but small; dise mod- 
erately convex, striato-punctate; strial punctures rather small and 
quite closely placed and not impressed; series rather distant, inter- 
stitial series of fine and more distantly spaced punctules, more or less 
irregular in arrangement; both series more irregular laterally and on 
apex. 

Male subfusiform oval. Elytra distinctly narrowed in apical 
fourth, disc gradually declivous behind. 

Female ovate. Elytra rather broadly oval, arcuately declivous at 
apex. 

Diagnostic characters.—This species is quite distinct from nigrina 
in its shining surface, smooth sculpturing and more depressed form ; 
the females are less elongate. 

Schwarzii is dull in lustre and more robust and not depressed, 
besides the sculpturing is subasperate; the sides of the pronotum is 
more dilated and as a whole transverse, with edge more strongly 
rounded and less convergent behind. 

Nevadensis is opaque; both sexes are typically subeylindrical; legs 
slender and the ventral surface distinctly but sparsely pubescent; the 
thorax is smaller. 

Dissimilis has heretofore been referred to some six species. There 
is nothing like it in the LeConte collection. 

A single male before me answers quite exactly to Colonel Casey's 
description of nitidus and it is possible that my name may have to 
be suppressed and Casey’s substituted. He refers n7f/dus to near 
longicollis. 

If dissimilis is nitidus no one who has compared my examples in 
the East has recognized this relationship, but it has invariably been 
associated with nigrina. The types are in the U. S. National Museum 
collection, and anyone can make comparisons. 

General observations—The middle lobe of the mentum is very 
small and triangular, the lateral lobes entirely exposed. more so than 

59780—Bull. 68—09——26 


402 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


in nigrina. This character has not been observed elsewhere in the 
genus Lleodes. The surface of the middle lobe is finely scabrous; the 
lateral lobes bear a few rather long sete, while the middle lobe is not 
noticeably setigerous. 

The prosternum is not strongly protuberant ventrally with the 
coxee, and distinetly setigerous. Between the cox it is more or less 
convex antero-posteriorly, sometimes horizontal. Generally with a 
small mucro behind, sometimes nearly vertically truncate with a 
mucronate angle. 

The mesosternum is arcuately declivous and more or less concave, 
with numerous small sete. 

The metasternum laterally between the coxe is as long as the width 
of a mesotibia at middle. 

The abdominal process is quadrate (male) or subquadrate (female) 
and about a fourth (male) of its width broader than the metasternal 
salient or subequal to it (female). 

The post-coxal part of the first segment is equal in length to the 
second ; the latter is twice as long as the fourth. 

In the male the third is about three-fourths longer than the fourth, 
in the female it is about two-thirds longer than the same segment. 

The profemora aré clavate, the mesofemora slightly swollen, and 
the metafemora somewhat narrowing toward base; the grooves are 
shallow and flat, the margins feebly subcariniform and sometimes 
feebly asperulate. The anterior margin of the profemoral grooves 
is not so distinctly sublaminate internal to the apical sinuation as in 
nigrina. On the meso- and metafemora the grooves vary as to their 
degree of convergence before evanescence. 

The tibie are without tarsal grooves; the external faces of the meso- 
and metatibie are somewhat flattened but not glabrous, and the 
articular cavities are closed. The protibie may or may not be 
carinate externally. 

The setee and spinules on the femora and tibie are usually black- 
ish, but frequently somewhat ferruginous on the tibie; on the tarsi 
they are nearly always ferruginous. 

The tarsi are somewhat slender, but frequently they appear mod- 
erately stout. 

Tarsal formula : 


Pro. Meso. Meta. Metatibia. 
Male.—24 34 . Be D4 
Female.—2 3 34 5 


ELEODES DISSIMILIS var. NEVADENSIS new. 


Cylindrico-fusiform to ovate, elongate, moderately depressed, 
opaque, more or less alutaceous and slender; thorax obsoletely punc- 
tulate; elytra striato-punctate; antenne, tibiee, and tarsi with the 


REVISION OF ELEODIINI—BLAISDELL. 403 


sete and spinules more or less strongly ferruginous; ventral surface 
distinctly pubescent. 

Head about twice as wide as long, frons frequently planate, or 
more or less convex, broadly and shallowly impressed laterally, 
frontal suture not evident, rather finely and not densely punctate, 
punctures denser at the periphery, each with a short reclining seta. 
Antenne long and moderately slender, outer four joints very feebly 
compressed and scarcely dilated, third joint about equal to the next 
two taken together, fourth to the seventh, inclusive, equal in length 
and almost twice as long as wide, eighth shorter and subtriangular, 
ninth and tenth suborbicular to feebly transversely oval, eleventh 
ovate. 

Pronotum subquadrate, widest at or slightly in advance of the 
middle, about one to two-sevenths wider than long: disc rather 
strongly convex and more or less declivousat the apical angles. 
quite obsoletely to evenly and not densely punctulate, the punctules 
slightly denser laterally: apex truncate in circular are to feebly 
emarginate, more or less finely to obsoletely margined; sides evenly 
and moderately arcuate in anterior two-thirds, more feebly so or more 
or less straight behind the middle, converging to base, finely mar- 
gined; base truncate or feebly rounded and about equal in width to 
the apex, finely margined; apical angles obtuse and not prominent: 
basal angles obtuse and not in the least rounded, rarely minutely 
prominent. 

Propleure finely, sparsely, and obsoletely punctulate, more or less 
feebly rugulose, most strongly on the acetabular convexities. 

Elytra elongate oval, smooth, widest usually at the middle: base 
truncato-emarginate and adapted more or less to the prothoracic base, 
which it scarcely exceeds in width: humeri obtuse, angle distinct 
although small and not prominent; sides evenly arcuate, apex not 
broadly rounded: disc slightly depressed and moderately convex on 
the dorsum, strongly and not very broadly rounded laterally, arcu- 
ately declivous behind ; surface striato-punctate, serial punctures small, 
closely placed in moderately distant rows, interstitial punctules more 
or less minute in a slightly irregular series, serial arrangement more 
or less evident laterally and on apex. 

Epipleure moderately wide, not dilated beneath the humeri and 
gradually narrowing to apex, superior margins feebly visible at 
humeri when viewed from above; surface smooth, more or less con- 
cave and obsoletely punctulate. 

Sterna more or less finely, densely, irregularly punctate and rugu- 
lose, sparsely clothed with erect and rather long ferruginous hairs. 

Parapleure more or less: obsoletely and irregularly punctate. 


404 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Abdomen finely punctate and rugulose, each puncture with a rather 
long, erect ferruginous hair; fourth and fifth segments frequently 
more or less rufous; horizontal. 

Legs slender and somewhat long. Femora finely and not very 
densely punctate, each puncture with a rather short reclinate seta; 
profemora feebly sinuate in both sexes. Anterior tibial spurs sub- 
equal and acute. Tarsi similar in the sexes; setee and spinules ferru- 
ginous throughout. 

Male.—Elongate, cylindrically subfusiform. Prothorax rather 
strongly convex. Elytra with sides not strongly arcuate, sometimes 
subparallel at middle, obliquely narrowed in apical fourth, arcuately, 
more or less obliquely and rather gradually declivous behind. 
Abdomen moderately convex and very feebly impressed along the 
middle on the first two segments. 

Female—Typically of the same form as the male, or more or less 
robust and ovate. Elytra with the sides moderately strongly arcuate, 
frequently broadened behind and narrowing more or less to base, 
rather suddenly arcuately declivous behind, slightly narrowed just 
before the apex (ovate form). Abdomen evenly and rather strongly 
convex. 

Measurements.—M ales: Length, 16.5-18 mm.; width 4-6 mm. Fe- 
males; Length, 17-17.5 mm.; width 7-6.5 mm. 

Genital characters, male-——Edeagophore rather small and fusiform. 

Basale evenly convex. 

Apicale quite equilaterally triangular; surface depressed and feebly 
membranous at middle; sides more or less feebly arcuate and shghtly 
sinuate before the apex, the latter not recurved, blunt and evenly 
rounded; base broadly lobed at middle and very feebly sinuate 
laterally. 

Sternite subparabolic in outline. Each lobe with the external 
border evenly arcuate to apex, the latter narrowly rounded; internal 
border more or less sinuate; surface rather densely and strongly 
punctate, setose, setee moderate in length. Membrane not setose at 
bottom of the sinus. 

Female.—Genital segment triangular, surface of the valves more or 
less inwardly declivous, feebly setose. 

Valvula (Plate 5, fig. 11).—Dorsal plate elongately triangular, nar- 
rowing from base to apex, smooth and shining, quite impuncetate, 
more or less feebly concave; external border not explanate, quite 
straight; apical margin narrow, not wider than the base of apex, 
angle not evident, rounded internally with the internal margin, the 
latter quite straight: apex quite narrow and slightly produced, 
convex above, slightly concave beneath, chitino-membranous, tip 
rounded and but slightly narrowed, smooth and shining, with few 


REVISION OF ELEODIINI—BLAISDELL. 405 


fine set on the internal surface; fossa small, located at base of the 
external surface. 

Appendage short mammilliform, smooth, with a pencil of some 
four short sete at tip. 

Basal prominences not in the least evident. 

Superior pudendal membrane rather long, quite attaining the 
apical margin of the dorsal plate, longitudinally rugulose. 

Ventrolateral surfaces —Body surface lines nearly straight when 
viewed longitudinally, slightly swollen in basal half, smooth, feebly 
setose apically; submarginal groove nearly obsolete, external margin 
of the dorsal plate being scarcely at all prominent laterally. Internal 
margins of the valves contiguous in basal two-thirds; fissure in 
apical third and closed. ; 

Habitat—Avizona (Tucson and Pinal mountains, Wickham; Pres- 
cott) ; Nevada (Pioche) ; California (Palm Springs, March). 

Number of specimens studied, 12. 

Type male in my collection; female (Cat. No. 12233) U.S. National 
Museum collection. 

Type-localities—Male, Tucson, Arizona; female, Palm Springs, 
California. 

Salient type-characters.—Both sexes elongate and subcylindrical. 
Antenne long. Head finely punctulate. Thorax small, a little wider 
than long; disc more or less obsoletely punctulate, rather strongly 
convex; apex feebly emarginate; sides moderately arcuate, narrowing 
posteriorly, marginal bead fine; base very feebly rounded; apical 
angles obtuse, distinct, not rounded nor prominent; basal angles 
obtuse, not rounded. Elytra at base more or less adapted to the pro- 
thoracic base, which it scarcely exceeds in width; humeri obtuse, 
angle distinct and small; dise more or less moderately convex, striato- 
punctate, strial punctures closely placed and rather small, strize mod- 
erately distant, interstitial punctules fine, slightly irregular and 
rather distantly spaced in a series, scarcely different laterally. 
Sterna and abdomen with distinct and sparse hairs: legs slender. 
Hairs and spinules ferruginous. 

Diagnostic characters.—The most slender and subcylindrical species 
of the subgenus. 

Easily recognized by its smooth, opaque, and alutaceous integu- 
ments, the antenne are long and the legs slender, the ventral surface 
is sparsely clothed with erect and rather long hairs. All hairs and 
spinules are ferruginous in color. 

The females are dimorphic. The type is of the same form as the 
male, while others are ovate and depressed like that sex in dissimilis. 

If it were not for this fact, I would consider nevadensis as very 
distinct; but as it is, there is difficulty at times in separating the 
females of the two races. 


406 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


The hairs of the ventral surface are easily lost, and specimens 
must be young or unworn. 

The hairs on the ventral surface of dissimilis ave short and dark. 
A male from Pioche, Nevada, represents a form with large pro- 
thorax, and the elytra are more oblong, with sides parallel at middle 
third. 

General observations—The mentum is as in dissimilis, but smaller. 

The prosternum in all of the examples at hand is not at all promi- 
nent ventrally with the coxe, feebly convex antero-posteriorly be- 
tween the coxe and distinctly mucronate behind, the mucro varying 
in size, but small and rounded. 

The mesosternum is small, quite vertically arcuate, and deeply 
excavated, so that the sides appear like, lobes. 

The metasternum laterally between the coxe is as long as the width 
of a mesotibia at apical fourth. 

The abdominal process is subquadrate and about a third of its 
width broader than the metasternal salient. 

The post-coxal part of the first segment is equal in iength to the 
second, the latter twice as long as the fourth (male). 

The third segment is about two-thirds (male) as long, or twice as 
long (female) as the fourth. 

The tarsi are long and slender. 

Tarsal formula: 


Pro. Meso. Meta. Metatibia. 
Male.—2 23 3 5 
Female.—24 3 34 54 


It is worthy of note that the female has unusually long tarsi as 
compared with the male; the types are quite similar in size. 


ELEODES SCHWARZII, new species. 


Suboblong-oval to suboblong-ovate, subopaque, rather robust. 
Thorax almost coarsely and densely punctate. Elytra submuricate 
laterally and on apex. 

Head twice as wide as long, feebly convex, moderately impressed 
laterally and somewhat along the frontal suture, the latter very 
slightly and vaguely indicated, densely and subscabrously punctate, 
punctures moderate. Antennw moderate in length and stoutness, 
outer four joints feebly compressed and slightly dilated, third joint 
about as long as the next two taken together, fourth very slightly 
longer than the fifth, the latter to the eighth, inclusive, subequal, 
the latter subtriangular, ninth about as long as wide, tenth slightly 
transversely oval, eleventh short-ovate. 

Pronotum somewhat transversely suboblong-oval, about a third 
wider than long, widest at about the middle; disc moderately and 
evenly convex, densely punctate, punctures almost coarse, granulate 


REVISION OF ELEODIINI—BLAISDELD. 407 


laterally; apew feebly and broadly emarginate, finely and more or less 
distinctly margined; sides rather strongly and quite evenly arcuate, 
feebly narrowed behind and slightly oblique, margin acute and promi- 
nent, marginal bead fine and thin, very narrowly reflexed; base very 
feebly rounded, finely and distinctly margined, about equal to the 
apex; apical angles obtuse, not prominent nor rounded; basal angles 
obtuse and not in the least prominent. 

Propleure rather finely, sparsely and evenly muricato-granulate, 
more or less rugulose. 

Elytra oval, less than twice as long as wide, widest at the middle: 
base transverse and not in the least margined; hwmeri rounded ; sides 
evenly arcuate, apex obtusely rounded but not broadly so; dise 
slightly convex on the dorsum, rather strongly and somewhat broadly 
rounded laterally, arcuately declivous behind; surface somewhat 
densely and scarcely subasperately punctate, the punctures about 
equal in size and in evident, rather closely placed series, laterally 
submuricate and irregular. 

E pipleure moderate in width, not dilated beneath the humeri, grad- 
ually narrowing from base to apex, superior margin not visible from 
above; surface smooth, dull, and obsoletely punctate. 

Sterna and paraplevrw more or less densely but not coarsely punc- 
tate. 

Abdomen more or less sparsely and subasperately punctate, irregu- 
larly regulose, surface more or less flattened at middle of the first two 
segments in both sexes. 

Legs moderate and quite densely sculptured. Anterior femora 
mutie and more or less sinuate beneath wm outer fifth; spurs of the 
protibiz similar in the sexes, the anterior slightly longer than the 
posterior, both quite slender and acute; anterior tarsi quite alike in 
the sexes, first joint moderately thickened and prominent beneath at 
apex, groove obliterated. 

Male——Scarcely robust, slightly narrow. Antenne reaching to the 
base of the prothorax. Elytra evenly arcuate and gradually decliv- 
ous behind. Abdomen noticeably oblique to the sterna, moderately 
convex and usually distinctly concave at middle on first three seg- 
ments. 

Female.—Moderately robust. Antenne attaining the posterior 
third of the prothorax. Elytra rather broadly oval, evenly and arcu- 
ately, but not vertically declivous posteriorly. Abdomen quite 
strongly convex and feebly flattened at middle toward base. 

Measurements.—Male: Length, 18 mm.; width, 7 mm. Females: 
Length, 19-20 mm.; width, 8.2-9 mm. 

Genital characters, male—Edeagophore elongate, oblong-ovate, 
sides subparallel and moderately arched. 


408 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Basale elongate oblong, moderately and evenly convex above, sides 
scarcely arcuate. ° 

Apicale triangular, slightly elongate, apex slightly recurved and 
flattened; surface moderately and evenly convex, with a narrow 
median groove from apex to base; sides feebly and broadly sinuate 
to near apex, thence briefly more convergent to tip, the latter subacute 
and narrowly rounded; base obtusely lobed at middle, laterally feebly 
sinuate. 

Sternite subparabolic in outline. Each lobe triangular and mod- 
erately distant from each other; ‘external border broadly sinuate in 
basal half, thence arcuate to apex, the latter subacute; internal mar- 
gin nearly straight; surface feebly convex, strongly and rather 
densely punctate in apical two-thirds, setose, setee long, shorter at the 
internal margin. Membrane not setose across the base of the sinus, 
the latter rather broad and deep. 

Female.—Genital segment (Plate 5, fig. 10) subquadrate, dorsal 
surface quite plane, glabrous and slightly setose. 

Valvula.—Dorsal plate oblong, smooth and shining; surface feebly 
and irregularly concave, coarsely and very sparsely punctate, each 
puncture with a rather long seta; external border feebly and broadly 
arcuate, more or less explanate apically; apical margin truncate and 
transverse, angle rounded; internal border more or less feebly sinu- 
ate. Apex small and subacute, apparently membrano-chitinous, very 
feebly and inconspicuously setose. 

Appendage small and mammilliform, with a pencil of short sete. 

Basal prominences not evident. 

Superior pudendal membrane not usually rugulose, quite broadly 
exposed and reaching about to the apical third of the dorsal plate. 

Ventrolateral surfaces—Body subequilaterally triangular; ventral 
surface lines nearly straight when viewed longitudinally, laterally at 
base slightly prominent, thence inwardly oblique and then broadly 
sinuate to apex, scarcely transversely concave before the apex; sur- 
face smooth, sparsely setose, setae short. Submarginal groove broad 
and shallow beneath the moderately expanded external border of the 
dorsal plate. Inferior surface of apex finely setose. Internal margins 
of the valves contiguous in basal two-thirds; fissure about closed. 

Habitat.—W ashington (Pullman, May, C. V. Piper, U. S. National 
Museum collection). 

Number of specimens studied, 3. 

Sexitypes in the U. S. National Museum collection (Cat. No. 12234). 

Type-locality.— Pullman, Washington. 

Diagnostic characters —Quite distinct in its more robust form, 
more transverse pronotum, the sides of which are more strongly 
rounded and dilated. 


REVISION OF ELEODIINI—BLAISDELL. 409 


The dull luster and sculpturing is nearly as in nigrina, although 
in the latter it is rather more coarse. In nigrina the pronotal dise 
is usually rather feebly defined from the sides by the comparatively 
feeble margins, which are often a mere line. 

In dissimilis the integuments are more shining and smooth, the 
pronotum rather quadrate and altogether it is a much more depressed 
species, the vertical diameter of schiwarzii being about a third greater. 

Nevadensis is a slender species, but the females tend toward 
schwarzii in their broader form, and the surface luster is opaque and 
alutaceous. 

General observations—The mentum is comparatively large and 
subtriangular rather than parabolic; surface is rather coarsely and 
distinctly punctate, somewhat feebly and broadly foveate laterally, 
setose, but inconspicuously so. 

The prosternum is not prominent ventrally with the coxe. In the 
single male before me it is arcuately rounded antero-posteriorly and 
not in the least mucronate; in one female, rounded between the coxve 
and distinctly mucronate behind, and in another example very 
slightly mucronate. 

The mesosternum is feebly arcuate, but obliquely declivous and 
very slightly and broadly concave. 

The metasternum laterally between the coxe is as long as the 
width of a mesotibia at the middle. 

The abdominal process is subquadrate and about a fifth (male) to 
a fourth (female) of its width broader than the metasternal salient. 

The process is equal in length to the post-coxal part of the first 
segment, also to the third; the second is about two-thirds longer than 
the fourth. These proportions appear quite similar in the two sexes. 

The femora are quite like those of nigrina, the grooves on the meso- 
and metafemora are more evanescent and the margins weaker. 

The tibizw are quite like those of nigrina; the protibie in the male 
are quite strongly constricted at base: this is also observed in per- 
longa. 

The tarsi are quite stout and somewhat long. 


Tarsal formula: 
Pro. Meso. Meta. Metatibia. 
Male.—2} 34 34 6 
Female.—24 3 4 6 


Subgenus STENELEODES, new. 


As in other subgenera of the genus //eodes, the present one is 
founded upon the female genital characters. which are very dis- 
tinctive. ; 

Otherwise the subgenus would be difficult to characterize or account 
of the paucity of intrinsic superficial characters, although the specific 
constituents possess a more or less distinctive habitus. 


410 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


The salient subgeneric qualities are as follows: Anterior femora 
mutic, integuments smooth, the form elongate "and more or less 
attenuate. 

Other characters of less diagnostic value may be mentioned: The 
humeri are feebly developed, except in innocens, where they are 
distinct; the first Joint of the anterior tarsi in the male is slightly 
thickened at tip beneath and bears a small transverse tuft of yellowish 
or brownish modified spinules, which interrupts the groove. 

The elytra are never caudate; the anterior tibial spurs are similar 
in the sexes and usually subequal, although subject to a slight varia- 
tion in their relative lengths, the anterior often appear slightly longer 
than the posterior and both may be longer, slightly larger in the 
female, but the males often exhibit this variation among themselves. 

The internal spurs of the meso- and metatibiz are always notice- 
ably longer than the external. | 

The abdomen is horizontal in the males of gigantea and gentilis, 
slightly oblique in estriata and most so in innocens; always hori- 
zontal in the females. 

Subgeneric genital characters, male——Apical lobe of the edea- 
gophore triangular, dorsal surface broadly depressed and concave, 
central area more or less membranous; apex blunt and not pro- 
duced; base broadly rounded at the middle third. 

Female.—Genital segment subparabolic and not setose, external 
lobe of the apex obsolete, the internal chitinous and more or less 
strongly developed, produced, convex above and excurved. 

Appendage minute and quite rudimentary. 

Dorsal plate of the valves elongate oval to subfalcate. 

Superior pudendal membrane broadly exposed by the distant dorsal 
plates. 

In the form of the apices of the genital segment and the minute 
appendages the present subgenus appears closely related to Désco- 
genta. 

Distribution.—Gigantea occurs on the peninsula of Lower Cali- 
fornia and in California as far north as the vicinity of San Fran- 
cisco Bay in the north central part of the State. 

Gentilis undoubtedly occurs in southern and central California; 
no definite locality has been recorded except San Diego. Specimens 
collected at San Diego by Colonel Casey and others received from 
Lower California by Doctor Horn, and therefore recorded from 
definite localities, were not gentilis at all, but either oméssa or its 
race pygmea. All specimens of gentilis that I have seen had simply 
i “Cal.” label, and these were carefully compared with a LeContian 
type by Mr. Blanchard and myself and the identification is correct. 
The species probably occurs in Lower California. 

Estriata is only known from the San Franciscan peninsula and 
adjoining region. 


REVISION OF ELEODIINI—BLAISDELL. 411 


Longicollis has a wide range of distribution. In Mexico it occurs 
in the States of Coahuila, Chihuahua, and Sonora. 

In the United States it is abundant in Texas, Arizona, New 
Mexico, Kansas, and Colorado. It probably extends into Nebraska, 
Wyoming, and Utah. A specimen in the National Museum collec- 
tion bears a label “ Oregon.” I believe this is erroneous; at any 
rate it remains to be verified. Longicollis occurs in Colorado atean 
elevation of 7,000 feet (Wickham). /nnocens thus far is only known 
to occur in Lower California. I have no knowledge of it occurring 
on the mainland in Mexico. 

Genealogy.—\lt would appear after a careful survey of the large 
series before me representative of the subgenus Steneleodes that the 
species have descended from a common ancestral stock along two 
distinct and divergent lines. Of course this only applies to those 
species included in our fauna. Similar study of the Mexican species 
would be the only way to definitely settle this question. 

The Mexican forms related to innocens I 
know nothing about. Longicollis and gigan- Ciguntna 
tea are closely related, and I believe the - Gantt 
former had its birth in Mexico, diffusion 
taking place to the northward, a western 
modification resulting in the latter. 

Unfortunately I have not seen any marked 
races of Jongicollis; there are undoubtedly 
some to be found in Mexico. Gigantea has 
apparently thrown off two divergent forms— 
gentilis, the older, and estriata, the more 
recent. I might say that ‘anocens is the more jyq 7 Gewvanoctear pta- 
heteromorphic; Jongicollis, gentilis, and — Gram or THR sUBGENUS 
gigantea as the more homomorphic. E'striata 9“ **°*°?"™ 
is slightly intermediate. 

Innocens has the abdomen oblique in the male; /ongicollis, gentilis, 
and gigantea have the abdomen horizontal in both sexes; estriata has 
the abdomen slightly, but undoubtedly, oblique. This may be an 
atavistic reversion to the primitive form, which may have had an 
oblique abdomen. 

Our species and their races may be separated as follows: 


Imocens — 


Elytra estriate; surface luster shining and bright. 

Pronotum feebly convex antero-posteriorly, normally strongly convex from 
side to side; sides rather evenly arcuate when viewed from above. 
mene On cane Oommrago’ Waiver. a. ee ____longicollis. 

Pronotum rather strongly convex antero-posteriorly ; sides strongly arcuate 
anteriorly. West of the Colorado River and Sierra Nevada Moun- 
per ap CEA SE RAE hs ree EB ee ee ee ee ee ee gigantea, 

Pronotum moderately and evenly convex, subquadrate. 


412 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Surface very smooth and shining, punctures very fine; legs slender; 
elytra widest at middle (male) or widest® behind the middle 


(female). Se ee ee oe var. estriata. 

Surface less shining, punctuation coarser; legs stouter; elytra widest 

at the middle (male) ; female unknown____----------_-_ var. gentilis. 

Elytra more or less: striate; surface luster dull____.___-.~-------=_-3 innocens, 


The above is the best table that I can devise at the present time. 
More material is necessary to elucidate the difficulties, that are very 
evident. 


ELEODES GIGANTEA Mannerheim. 


Eleodes gigantea MANNERHEIM, Bull. Soc. Nat. Moscow, XVI, 1848, p. 267; 
Mag. Zool., XIII, 1843, pl. 127—LeContr, Proc. Acad. Nat. Sci. 
Phila., 1858, p. 181—Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 312. 

yar. gentilis LeContTE, Proc. Acad. Nat. Sci. Phila., 1858, p. 187.—Horn, 
Trans. Amer. Phil. Soc., XIV, 1870, p. 311. 

var. estriata Casey, Ann, N. Y. Acad. Sci., V, Nov. 1890, p. 398. 


Strongly elongate, convex, smooth, and moderately shining. 

Head about twice as wide as long, moderately convex, often more 
prominently so in the median line of the frons as far as the frontal 
suture, not coarsely, somewhat sparsely and irregularly punctate, 
the punctures becoming denser at the sides; frons generally impressed 
within the supra-antennal convexities, the impressions frequently 
extending along the frontal suture; antenna rather stout, not reach- 
ing to the base of the prothorax; outer four joints very slightly com- 
pressed, gradually and very feebly dilated, third joint slightly longer 
than the next two taken together, fourth just perceptibly longer than 
the fifth, joints 5-8, inclusive, subequal in length, the seventh and 
eighth subtriangular, ninth suborbicular, tenth transversely oval, 
eleventh subovate. 

Pronotum widest just in advance of the middle and about one-fifth 
wider than long; disc strongly convex, arcuately declivous laterally, 
finely, evenly, and rather sparsely punctulate, punctures becoming 
slightly denser laterally; apex truncate or very feebly emarginate in 
circular are, finely to obsoletely margined; sides evenly and rather 
strongly arcuate in the anterior two-thirds, thence nearly straight and 
converging to the base, at times evenly and moderately arcuate 
throughout, finely margined; base very feebly arcuate, more strongly 
margined, about one-fourth widet than the apex; apical angles small, 
subacute, and rarely slightly everted ; basal angles obtuse, not rounded 
nor prominent. 

Propleure opaque, obsoletely or very sparsely punctulate and rugu- 
lose. 

Elytra elongate oval, usually widest at the middle; dase scarcely 
emarginate and about as wide as the contiguous prothoracic base; 
humeri obtuse; sides evenly arcuate; apex feebly produced and nar- 
rowly rounded; disc convex; suture frequently slightly impressed ; 


REVISION OF ELEODIINI—BLAISDELL. 413 


dorsum sometimes depressed, arcuately declivous posteriorly ; surface 
finely, irregularly, and evenly punctate, rarely with closely placed 
series of slightly larger punctures, the intervals with a more or less 
irregular series; the punctures are slightly denser at the sides and on 
the apex, never asperate. 

Epipleure moderately wide beneath the humeri, evenly and gradu- 
ally narrowing to the apex; surface finely and sparsely or obsoletely 
punctate, more or less finely rugulose. 

Sterna rather strongly and sparsely punctate and rugose; the pro- 
sternum anteriorly and the metasternum between the coxe laterally, 
frequently obsoletely punctate. 

Parapleure strongly punctate anteriorly, sparsely so posteriorty. 

Abdomen horizontal, finely and sparsely punctate, the punctures 
becoming denser at the periphery of the fifth segment; surface more 
or less rugulose, at times strongly so. 

Legs moderately long. Femora not usually densely punctate, the 
anterior mutic in the sexes. Anterior tibial spurs subequal. Pro- 
tarsi slightly dissimilar in the sexes. 

Male.—Very elongate and attenuate. Antenne attaining the pos- 
terior fifth of the prothorax. Elytra gradually narrowed and very 
gradually arcuately declivous posteriorly. Abdomen moderately 
convex and very feebly, broadly impressed on first three segments. 
Posterior tibiz more or less arcuate. Anterior tarsi with the first 
joint slightly thickened at apex beneath; apico-marginal tufts of 
spinules somewhat attenuate, brownish or yellowish, those of the first 
joint more or less transversely confluent and slightly modified; groove 
subentire. 

Female.——Elongate and rather broad. Antenne scarcely attaining 
the posterior fifth of the prothorax. Elytra slightly depressed on the 
dorsum, not gradually attenuate and less gradually or more strongly 
arcuately declivous posteriorly. Abdomen strongly convex. Pos- 
terior tibiw quite stout and scarcely arcuate. Apico-marginal tufts 
of spinules of the tarsi rather stout, those on the first joint of the 
protarsi not transversely confluent; groove entire. 

' Measurements.—Males: Length, 31-35 mm.; width, 9.5-10 mm, 
Females: Length, 28-35 mm.; width, 10.5-13 mm. 

Genital characters, male.—Edeagophore oblong-ovate and some- 
what fusiform, moderately arched. 

Basale elongate oblong, longitudinally convex, and evenly so from 
side to side; apex with a rounded emargination. 

Apicale (Plate 1, fig. 6) triangular. Surface broardly and strongly 
depressed, very broadly membrano-chitinous and with a linear median 
groove at middle two-fourths; sides nearly straight, feebly sinuate 
at apex, the latter narrowly rounded, gradually and feebly recurved ; 
base with a rounded lobe at middle third and sinuate laterally. 


414 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Sternite subhemispherical in outline. Each lobe quite triangular; 
external border evenly arcuate, apex broadly rounded, with the in- 
ternal margin more or less arcuate; surface densely punctate and 
setose in apical half, setee long and dense. Membrane not setose 
across the bottom of the sinus, the latter moderately wide. 

Female.—Genital segment (Plate 1, fig. 4) subparabolic in outline, 
strongly chitinized, not setose. Superior pudendal membrane 
broadly exposed, dorsal plates widely separated. 

Valvula.—)Dorsal plate subfaleate, with surface more or less con- 
cave, and rather coarsely, sparsely punctate; margins slightly con- 
verging to the apical margin, which is more or less rounded. Internal 
surface of the valve inwardly declivous and more or less exposed, 
finely and densely punctate, slightly excavated beneath the apex, 
the latter chitinous, semi-oval in outline, flattened, projecting up- 
ward and backward, moderately divergent, tip broadly and evenly 
rounded, with the surface feebly convex and quite strongly punctate. 

Appendage minute, scarcely projecting beyond the margins of the 
minute fossa. 

Basal protuberances feeble. 

Superior pudendal membrane attaining the apical fourth of the 
segment, longitudinally rugulose. 

Ventrolateral surfaces conjointly ogival when viewed from below, 
straight when viewed longitudinally and not inflated, sparsely punc- 
tate. Apex concave beneath, the concavity extending forward and 
laterally, so that the dorsal plate appears expanded and the sub- 
marginal groove broad. Internal margins of the valves contiguous, 
the genital fissure closed. - 

Habitat—California (San Diego, Orange, Los Angeles, Kern, and 
Santa Cruz counties). Lower California (San Pedro Martir). 

Number of specimens studied, 51. 

Location of type not known. 

T ype-locality —* California ” (Mannerheim). 

Salient type-characters.—Elongate, shining. Thorax, convex and 
rounded; femora, mutic; elytra, finely granulato-punctate (Manner- 
heim). 

Diagnostic characters.—Readily recognized by the elongate and at- 
tenuated form of the males; the females, while elongate, are also more 
robust, at times decidedly so. The most common form of sculpturing 
observed is that in which the elytra are smooth, polished, minutely 
and rather sparsely punctate, the punctures being diffusely placed. 

I have many specimens with more or less distinctly indicated and 
closely placed series of punctures on the elytra, and they were not 
from any one region. - Mannerheim’s type had the elytra finely granu- 
lato-punctate. This form of sculpturing is lesscommon, The thorax 
is finely and sparsely punctulate. 


REVISION OF ELEODIINI—BLAISDELL. 415 


From longicollis (see p. 427) it is separated by feeble characters. 

In gigantea the thorax is usually much more strongly arcuate at 
the sides, the disc less strongly arcuately declivous and inflexed, the 
pronotum appearing broader, as compared to dongicollis where the 
side margins are more strongly declivous and inflexed, the margins 
appearing less strongly arcuate and the pronotum longer. 

In gigantea the thorax has a much stronger antero-posterior eon- 
vexity. 

The mutic profemora of both sexes distinguishes gigantea from all 
of the members of the subgenus //eodes, except sponsa, caudifera, 
and Jongipilosa; these species are roughly sculptured, or caudate, and 
therefore not easily mistaken for gigantea. 

The genital characters serve to separate it from all other subgenera 
if there be any doubt. 

The form found about San Francisco Bay and named by Colonel 
Casey as estriata is related to gentilis (see p. 423) ; the typcial form is 
rare, but found there. 

Some of the smaller individuals might be more difficult to separate 
from the larger specimens of nigrina. The latter species has dull 
integuments and the sculpturing is coarser, especially that of the 
prothorax. 

I have one specimen of nigrina that was collected at Verdi, Nevada, 
which simulates gigantea quite perfectly; it measures 24 mm. in 
length, which is larger than some of the smaller examples of gigantea. 

General observations.—The mentum is triangulo-parabolic in out- 
line, slightly wider at base than long; the apex is usually rounded, 
sometimes apparently truncate on account of the tip being slightly 
and briefly recurved; rarely the apical margin is feebly impressed 
at middle so as to appear emarginate; the surface is quite coarsely 
punctate, the punctures are not usually crowded, each bearing a 
rather long hair, giving it a bearded appearance. 

The prosternum is strongly arcuate between the cox, both being 
decidedly protuberant ventrally: the ante-coxal surface more or less 
sinuate and oblique, continuing the surface line of the intercoxal 
part; behind it bears a small conical mucre. These characters are 
quite constant in the series before me. 

The mesosternum is quite vertical and moderately arcuate, more or 
less concave. 

The abdominal and metasternal salients are about equal in width. 
The abdominal process is slightly longer than wide, that is, slightly 
oblong. 

The post-coxal portion of the first segment is equal in length to the 
second, the latter is about, or a little more than twice as long as the 
fourth; the third is a little less than twice as long as the fourth, 


416 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


The metasternum laterally between the cox is about as long as the 
width of a metatibia at middle. 

The profemora are very feebly swollen in external half; this condi- 
tion is very gradually attained from the base; the surface lines are 
very feebly arcuate when viewed lengthwise; the tibial grooves are 
moderate in width, with margins rather coarse and slightly arcuate, 
becoming evanescent a little distance from the base but not contigu- 
ous at point of evanescence; floors feebly concave, opaque, and more 
or less minutely reticulate. 

Meso- and metafemora almost subequal in width throughout when 
taken individually, each having the corresponding sides subparallel. 
The grooves of the mesofemora are a little narrower than the pro- 
femoral, the margins are coarse and subgranulate, evanescent at in- 
ternal third; floors feebly concave, smooth and opaque, more or less 
rugulose, sometimes transversely so. | 

Metafemoral grooves a little narrower than the mesofemoral and 
very feebly concave; margins coarse, more or less granulate, the 
posterior frequently finely serrulate, both evanescent a little internal 
to the middle. 

The tibia are usually quite stout and usually more or less feebly 
arcuate. The protibie are scarcely compressed and more or less 
carinate externally ; the tarsal grooves are more or less developed and 
asperate; the articular cavities are more or less closed. 

The mesotibix are somewhat circular in section, usually not at all 
carinate externally; grooves more or less developed, rather narrow, 
opaque and limited by the general muricate sculpturing. 

The metatibie are quite cylindrical in section, otherwise like the 
mesotibie. The articular cavities are closed in each. 

The tarsi are stout and moderate in length. 

The protarsi are about a fourth of their length shorter than a 
mesotarsus. Joints 24 subequal in length and apparently decreasing 
in width in the order named, a little wider than long and together 
about equal in length to the fifth; first joint a little longer than wide. 

The mesotarsi are about equal to a metatarsus in length. Joints 
2-4, inclusive, subequal in length and very slightly decreasing in size 
in the order named, and together subequal in length to the fifth; the 
first about twice as long as wide. 

The metatarsi are about half as long as their respective metatibia. 
Joints 2 and 3 slightly decreasing in length, width, and size in the 
order named, each distinctly a little longer than wide, together 
scarcely as long as the fourth and quite as long as the first. 


ELEODES GIGANTEA var. GENTILIS LeConte. 


Elongate fusiform, more or less shining and smooth. 
Head about twice as wide as long, feebly convex, lateral impres- 
sions very feeble, frontal suture evident and usually marked by a line 


REVISION OF ELEODIINI—BLAISDELL. 417 


of punctures, very feebly impressed; punctures somewhat coarse, 
sparsely and irregularly placed, more closely so at the periphery. 
Antenne moderately slender and moderate in length, scarcely attain- 
ing the prothoracic base, feebly compressed and very slightly dilated 
in outer four joints, the third joint equal in length to the next two 
taken together, fourth apparently very slightly longer than the fifth, 
5-8, inclusive, subequal in length, eighth feebly triangular and about 
as wide as long, ninth and tenth orbicular or feebly transversely oval, 
eleventh short-ovate. 

Pronotum subquadrate, widest at about the middle, about one- 
seventh wider than long; disc moderately convex, more arcuately and 
strongly so at the sides, finely and rather sparsely punctulate, pune- 
tures scarcely at all denser laterally; apex subtruncate in circular 
are, and finely beaded; s/des rather evenly but not strongly arcuate 
when viewed from above, slightly narrowed posteriorly and finely 
beaded; base feebly rounded and less finely beaded, about one-sixth 
wider than the apex; apical angles not in the least prominent 
anteriorly, scarcely at all rounded and not acute; basal angles obtuse 
and not rounded. 

Propleure smooth and more or less sparsely and obsoletely punc- 
tulate, rugulose. 

Elytra elongate oval, widest at the middle; base very feebly 
emarginate, scarcely wider than the contiguous prothoracic base; 
humeri obtuse, not prominent nor rounded; s#des evenly arcuate, apex 
rather narrowly rounded and not produced; disc convex and quite 
evenly rounded at the sides, arcuately declivous and rather obliquely 
attenuate posteriorly; surface finely, sparsely, and irregularly punc- 
tulate, sometimes scarcely serially arranged, the punctures slightly 
denser at the sides and on apex, never asperate, scarcely subgranulate 
at times. 

Epipleure moderately narrow beneath the humeri, very gradually 
and evenly narrowing to apex; surface finely, sparsely, and obsoletely 
punctate, sometimes feebly rugulose. 

Sterna not coarsely rugose and more or less punctate. 

Parapleure rather sparsely but distinctly and not coarsely punc- 
tate. 

Abdomen horizontal, sparsely punctulate and more or less rugu- 
lose, smooth, and shining. 

Legs moderate in stoutness and length, not densely sculptured, 
most so on the tibige; anterior femora mutic in the sexes; tibial spurs 
subequal; tarsi as in gigantea. 

Male.—Flongate and quite fusiform. Elytra gradually narrowed 
in apical third, evenly, gradually arcuately declivous posteriorly ; 
abdomen moderately convex, more or less broadly and not strongly 

59780—Bull. 683—09——27 


418 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


impressed at middle on the first three segments; posterior tibiae more 
or less feebly arcuate. 

Measurements —Males: Length, 20-27 mm.; width, 7-10 mm. 
Females: Unknown. 

Genital characters, male-—Edeagophore comparatively large, ob- 
long-ovate, very elongate and slightly arched. 

Basale elongate oblong, evenly convex from side to side, moder- 
ately strongly convex antero-posteriorly, with a moderate longitudi- 
nal impression at middle in apical moiety; sides subparallel; surface 
glabrous and shining, sparsely punctate apically at the sides; apex 
with a rounded emargination at middle. 

Apicale triangular. Surface convex laterally, with a depressed 
oval chitino-membranous area at the middle, thence to apex broadly 
and not deeply grooved; apex recurved and quite broadly rounded; 
sides quite straight, although feebly but broadly sinuate at apex; 
base with a well-marked, rounded lobe at middle and quite strongly 
sinuate laterally. 

Sternite subparabolic in outline and slightly transverse. Each 
lobe with the external border quite evenly and broadly arcuate from 
base to apex, the latter narrowly rounded; internal border short and 
arcuate; surface quite convex, glabrous in basal half, thence to apex 
strongly punctate, the punctures increasing in density, each with a 
rather long seta, the latter dense in apical third. Sinus broad and 
more or less oblong-oval, the inner and contiguous surface of the 
lobes inwardly and gradually declivous. Membrane not setose and 
irregularly rugose. 

Habitat.—California. The exact locality is not indicated on the 
specimens at hand. LeConte’s cotype before me bears a green 
“Calif.” label, but in his description of the species he gives San 
Diego, California. 

All the specimens that I have seen and which were referred to 
gentilis by Doctor Horn have thus far turned out to be something 
else, and I am not sure that he had typical specimens in any of these 
instances. 

Gentilis, no doubt, occurs in Central and Lower California, as well 
as in the southern part of the State. 

Lower California specimens* referred to gentilis by Horn were 
insularis or some variation of omissa. Colonel Casey failed com- 
pletely to recognize gentélis, and considered the race described in this 
paper as pygmea as that species, saying that it is abundant at San 
Diego.” I have a specimen which he identified as gentilis. It is 
simply a pygmea with a very quadrate prothorax. 

Pygmea is abundant at San Diego, but gentilis seems to be very 
rare, for IT have not seen any recently collected specimens. During 


@See Proc. California Acad. Sci., 2nd ser., IV, Pt. 1, pp. 306 and 349. 
d> Ann. N. Y. Acad. Sci., V, Novr., 1S90, p. 395. 


REVISION OF ELEODIINI—BLAISDELL. 419 


many years’ residence in San Diego County and after collecting over 
the entire western half of the county I had not a single specimen. 

Number of specimens studied, 3; one a LeContian cotype. 

Type in the LeConte collection in Cambridge, Massachusetts. 

Type-locality.—San Diego, California. 

Salient type-characters.—Elongate. Thorax convex with the sides 
rounded and slightly narrowed behind, finely and sparsely punctuate. 
Elytra elongate oval, declivous and obliquely attenuate behind, finely, 
sparsely, and scarcely seriately punctulate (LeConte). 

Diagnostic characters.—Closely allied to gigantea but very much 
smaller, with thorax less rounded on the sides and less narrowed 
toward base. 

In gentilis the elytral punctulation is a little coarser than in gigan- 
tea. 

One specimen before me measures 27 mm. and is as large as an or- 
dinary gigantea. 

There is something strange about the manner in which gen¢i/is has 
been misunderstood, and it has given me more trouble to study it out 
than any other species: it was with a feeling of deep satisfaction 
when, through the kindness of Mr. Samuel Henshaw, I was permitted 
to receive one of LeConte’s cotypes through transmission from the 
museum in Cambridge, Massachusetts. Two of the specimens already 
in the material at hand had been referred to gigantea. Mr. Blanchard 


‘truly defined gentilis when he wrote me that it looked to him like a 


small fusiform gigantea. 

A specimen in Mr. Ralph Hopping’s collection was wrongly identi- 
fied when received by him and bears the number 7337, and was evi- 
dently received from Reinecke, for this name is on the label. This 
shows how the earlier collectors obtained these specimens, while all 
recently collected material yields no examples of this race. 

Estriata is a modified gentilis. It is more glabrous, the elytra of 
the male is more inflated and with a tendency to become widest behind 
the middle. 

The male in its genital characters shows some approach to innocens 
in the oval chitino-membranous area of the apicale of the edeago- 
phore, while the same characters in estriata are intermediate be- 
tween gentilis and gigantea. But these characters are no doubt 
subject to considerable individual variation. 

Gentilis therefore appears to be nothing more than a race of 
gigantea, LeConte’s types being simply an extreme nanoid form, 
while Hopping’s specimen has the normal size of gigantea. The 
females probably do not differ from the general form of the female 
gigantea, the males being dimorphic. 

General observations—The mentum is trapezoido-parabolic in out- 
line and comparatively small, setose, the sete being moderately long 


420 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


and somewhat transversely arranged on the antefior half so as to 
give a distinctly bearded appearance. Otherwise as in gigantea. 

The prosternum is moderately prominent ventrally with the coxe, 
convex antero-posteriorly and grooved; there is a small mucro 
behind. 

The mesosternum is arcuately declivous and more or less vertical, 
concave to a varying degree. 

The metasternum laterally between the cox is about as long as 
the width of a mesotibia at middle. 

Comparative measurements gave the same relative proportions 
between the abdominal segments, metasternal and abdominal salients 
of the two extreme specimens of gentilis before me. LeContian 
cotype, male, 20 mm.; Hopping’s large example, male, 27 mm. 

The abdominal salient is about a third of its: width broader than 
the metasternal process; the abdominal salient is subquadrate, a little 
longer than wide; the post-coxal portion of the first segment is equal 
to the third in length, and the second is twice as long as the fourth. 

The profemora are feebly swollen in outer half, but very gradually 
so; the grooves are moderate in width with margins feebly sub- 
cariniform, feebly arcuate, converging to become contiguous and 
evanescent at basal fourth; the floors of the grooves are smooth and 
scarcely concave. 

The mesofemora are very feebly and just noticeably widened in 
middle third, narrowing slightly and gradually each way; the 
grooves are a little narrower, margins scarcely subcariniform and 
more or less granulate, scarcely at all arcuate, converging slightly 
to become evanescent at basal third without becoming contiguous; 
floors of grooves nearly plane. 

Metafemora are not swollen, but with the opposed surfaces sub- 
parallel; grooves narrower and otherwise as those of the mesofemora. 

The tibie are very slightly arcuate in the three males at hand. 

The protibie are obsoletely subcarinate externally, tarsal grooves 
more or less obsolete; the articular cavities are closed. 

The mesotibie are obsoletely subcarinate externally at basal third, 
otherwise as in the protibie. 

The metatibie are more or less rounded, not carinate and obsoletely 
grooved. 

The tarsi vary in stoutness according to the size of the example. 

The protarsi are about a fifth of their length shorter than a meso- 
tarsus. Joints 2-4, inclusive, are subequal in size, apparently very 
feebly wider than long and together about equal to the fifth; joint 
one distinctly longer than wide. 

The mesotarsi are about equal in length to a metatarsus. Joints 
2-4, inclusive, are subequal in size, very feebly longer than wide, 


REVISION OF ELEODIINI—BLAISDELL. 421 


together about equal to the first; the fifth scarcely as long as the three 
preceding joints taken together. 

The metatarsi are a little longer than half the length of their 
respective metatibia. Joins 2 and 3 subequal in size, distinctly longer 
than wide, and together equal in length to the first or fourth taken 
individually. 


ELEODES GIGANTEA var. ESTRIATA Casey. 


Elongate, ovato-fusiform, moderately robust, strongly convex, 
smooth and polished. 

Head rather variable in size, nieteasiely convex, impressions obso- 
lete, rather coarsely punctate, the punctures sparse toward the middle, 
denser and setose laterally. 

Antenne somewhat variable in stoutness, short, reaching to about 
the prothoracic base, slightly compressed and scarcely dilated in the 
outer five joints; third joint scarcely longer than the next two to- 
gether; fourth to the seventh, inclusive, subequal in length; fourth 
sometimes slightly longer; eighth triangular; ninth orbicular; tenth 
feebly transverse; eleventh subovate and more or less obliquely trun- 
cate at tip. 

Pronotum subquadrate, widest at about the middle, slightly wider 
than long; disc moderately convex, very finely and sparsely punctate 
‘throughout; @pea truncate in circular arc, more or less obsoletely 
beaded; s¢des broadly arcuate anteriorly, more or less straight and 
convergent in basal half, at times somewhat arcuate throughout, very 
finely beaded; base subtruncate, rather finely beaded, and a little 
wider than the apex; apical angles obtuse, not rounded nor anteriorly 
prominent; basal angles obtuse and not prominent. 

Propleure finely punctulate anteriorly and more or less obsoletely 
rugulose. 

Elytra elongate oval, widest at or behind the middle; base subtrun- 
eate and about equal in width to the contiguous prothoracic base ; 
humeri not rounded nor prominent; sides arcuate, apex subacute and 
not at all produced; disc moderately convex, more strongly rounded 
laterally and arcuately declivous posteriorly; surface finely or rather 
coarsely punctate, punctures sparsely placed and not asperate, not 
denser laterally and arranged without trace of order throughout, at 
times with evident closely placed series. 

Epipleure moderate in width, gradually narrowing from base to 
apex, superior margin quite straight when viewed longitudinally; 
surface smooth, sparsely and obsoletely punctulate. 

Sterna quite strongly punctate and rugose throughout. 

Parapleure moderately closely punctate. 

Abdomen shining, sparsely and irregularly punctulate, rather obso- 
letely and reticulately rugulose. 


422 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Legs moderate in length and variable in stoutne§s. Femora rather 
sparsely punctate, the anterior mutic in the sexes; anterior tibial 
spurs almost equal. Tarsi slightly dissimilar in the sexes. 

Male.—Rather narrow, not distinctly ovate, more fusiform. Elytra 
quite evenly arcuately declivous and somewhat gradually narrowed 
posteriorly. Abdomen feebly oblique, moderately convex, and more 
or less broadly and feebly impressed at middle of first two segments. 
Legs somewhat slender, especially the tibize. Anterior tarsi with a 
small transverse tuft of modified spinules at tip of first joint beneath, 
the tuft closing the plantar groove. 

Female.—Rather robust. Elytra widest behind the middle, areu- 
ately and somewhat obliquely declivous and somewhat rapidly 
narrowing posteriorly. Abdomen horizontal, rather strongly and 
evenly convex. Legs rather stout, the tibiae moderately so. First 
joint of the protarsi not modified at tip beneath, plantar groove wide. 

Measurements —Male: Length, 20 mm.; width, 7.2 mm. Females: 
Length, 22.5-26 mm; width, 9-10 mm. 

Genital characters, male—Edeagophore as in gigantea. Sternite 
‘ather short and transversely parabolic in outline. Each lobe rather 
small and subtriangular; external border subangulate at middle, 
thence to base feebly arcuate, to apex obliquely truncate, apex nar- 
rowly rounded; inner border feebly arcuate; surface densely and_ 
strongly punctate in apical moiety. each puncture with a long black 
seta. Membrane not setose, sinus broadly oval. 

Female.—Genital segment subparabolic in outline, feebly elongate 
and finely setose. Superior pudendal membrane broadly exposed, 
and the dorsal plates widely separated. 

Valvula—Dorsal plate subfalcate, with surface more or less con- 
cave, facing upwards and outwards, coarsely punctate, each puncture 
with a short, rather stout semirecumbent seta; borders gradually 
converging to the apical border, which is more or less rounded ; inter- 
nal border quite sharp and broadly sinuate; internal surface of the 
valve much exposed and inwardly declivous, densely and rather 
strongly punctate in apical third beneath the apex, each puncture 
with a short, curved, and semirecumbent seta. Apex chitinous, semi- 
oval in outline, flattened, projecting upward and backward, mod- 
erately divergent, tip broadly rounded; surface feebly convex and 
more or less punctate. 

Appendage minute, scarcely projecting beyond the fossa, which is 
situated in an excavation between the external surface of apical base 
and the apical margin of the dorsal plate; short, semioval in form, 
with a few short set at tip. 

Basal prominences not noticeable. 

Ventrolateral surfaces as in gigantea. 


<— 


REVISION OF ELEODIINI—BLAISDELL. 423 


Habitat—California (Presidio sand hills, San Francisco, June; 
San Mateo County, at Holy Cross Cemetery; Alameda County). 

Number of specimens studied, 8. 

Type in Colonel Casey’s collection. 

Type-locality.—San Francisco, Cal. 

Salient type-characters.—Head transverse and moderate in size; 
antenne rather short and robust, third joint very slightly longer than 
the next two together, fourth less than twice as long as wide. 

Prothorax just visibly wider than long, apex truncate. and equal 
in width to the base which is subtruncate; sides broadly arcuate 
anteriorly, convergent and nearly straight in basal half, basal angles 
obtuse and not at all prominent; dise moderately convex, very finely 
and sparsely punctate. 

Elytra widest behind the middle, humeri not rounded nor promi- 
nent; apex acute but not greatly prolonged; dise finely but distinetly 
and very sparsely punctate, the punctures not asperate and without 
trace of order throughout (Casey). 

Diagnostic characters.—A local race of gigantea, from which it 
differs in its “ smaller size, less attenuate form, wider epipleure, less 
convex pronotum, less arcuate sides of the prothorax, and shorter 
elytra” (Casey). 

From longicollis it is recognized by having the “ prothorax more 
anteriorly dilated, much shorter and rather more robust antenne, 
and coarser punctuation ” (Casey). 

It is undoubtedly more closely allied to genfi/is than to any other 
form belonging to the present subgenus. It is also very variable as 
indicated by Casey’s two specimens and the small series before me. 


‘The male, as directly compared with the LeContian co-type of 


gentilis, has the same form, but is more polished, has finer punctu- 
ation, more slender legs, especially the tibiz, and is altogether a more 
graceful insect. 

Unfortunately the female of gentilis has not been recognized. Two 
females of estriata now before me are the homomorphic companions 
of the male above compared with genfélis, and are more robust with 
the elytra widest behind the middle and the tibiz are distinctly stouter. 
I judge that Casey’s smaller specimen with longer and more slender 
legs is a male, and the larger example with a more robust habitus, 
is undoubtedly a female. A specimen which I took in the Holy 
Cross Cemetery in San Mateo county, is the most robust specimen 
that I have seen and it is a female, it has exactly the same measure- 
ments as Casey’s large specimen and agrees with his description, the 
antenne are distinctly stout and shorter than in the other specimens 
before me. 

The individuals taken about the sand hills on the peninsula are 
very graceful insects and at first sight apear quite distinct from 


494 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


gigantea; specimens taken more inland gradually approach the 
typical form of the last named species. £’stviata is not common. 

General observations.—The mentum is trapezoidal to trapezoido- 
parabolic, more or less feebly convex, rather coarsely punctate, each 
puncture with a rather long hair. 

The prosternum as in gigantea, except that the small conical mucro 
is usually obsolete. 

Mesosternum as in gigantea. 

The abdominal intercoxal salient is quite quadrate and about a 
fifth of its width broader than the metasternal process. 

The post-coxal part of the first abdominal segment is about equal 
to the second in length; the second segment is about a fourth longer 
than the third, the latter being about a half longer than the fourth. 

The metasterum laterally between the coxe is about equal in length 
to the width of a mesotibia at apical fourth. 

The profemora are scarcely at all swollen, and the surface lines 
are evenly and rather feebly arcuate, not distinctly subclavate, al- 
though shghtly narrowed at base; tibial grooves strongly defined by 
the rather coarse cariniform margins, which are feebly arcuate and 
convergent, becoming contiguous at the femoral base and _ there 
shghtly evanescent. The protibie are obsoletely carinate externally 
and oval in section, the tarsal grooves are obsolescent. 

The mesofemora are somewhat variable as to point of greatest 
width; surface lines subparallel, noticeably convergent in basal 
fourth; tibial grooves distinctly concave, margins not cariniform; 
evanescent at basal third before becoming contiguous. The mese- 
tibiz are oval to cylindrical in transverse section; tarsal grooves not 
impressed and recognized as a smooth surface between the sparse 
asperities. 

The metafemora have the surface lines subparallel, slightly nar- 
rowed in basal fifth; tibial grooves as on the mesofemora and evanes- 
cent at or basad to the middle. Metatibiz variable, but with general 
characters as in the mesotibie. 

The articular cavities are quite closed on all the tibie. 

The tarsi are variable as to stoutness and moderate in length. 

The protarsi are about a fifth of their length shorter than a meso- 
tarsus. Joints 2-4, inclusive, short and subequal in size and together 
about equal to the length of the fifth; first joint one-half longer than 
the second. 

The mesotarsi are just a little longer than a metatarsus. Joints 
2-4 are subequal and together just a little longer than the fifth; the 
first is subequal to the second and third taken together. A metatarsus 
is a little more than half as long as its metatibia. Joints two and 
three are subequal in size, a little longer than wide, and together 
equal to the length of the fourth or first. 


_ 


REVISION OF ELEODITNI—BLAISDELL. 425 


ELEODES LONGICOLLIS LeConte. 


Bleodes longicollis LeConrr, Ann. Lyc. Nat. Hist. N. Y., V., 1851, p. 134; 
Proc. Acad. Nat. Sci. Phila., 1858, p. 181.—Horn, Trans. Amer. Phil. 
Soc., XIV, 1870, p. 311—CHAmpion, Biol. Cent.-Amer., 1V, Pt. 1, 1884. 
p. 84. 

Bleodes haydenii LeConrr, Proc. Acad. Nat. Sci. Phila., 1858, p. 186. 

Elongate to elongate-fusiform, black, the elytra often more or less 
reddish, smooth and shining. 

Head twice as wide as long, more or less moderately convex, finely 
and quite evenly punctate, the punctures becoming slightly denser at 
the sides. Antenne moderately stout, about reaching to the base of 
the prothorax; outer five joints very feebly compressed and scarcely 
dilated ; third joint about as long as the next two taken together; 4-7, 
inclusive, subequal and nearly evlindrical; eighth triangular; ninth 
and tenth suborbicular; eleventh ovate. 

Pronotum widest at the middle and about one-seventh wider than 
long; dise quite evenly convex, the convexity strongest transversely, 
slight antero-posteriorly, evenly and rather sparsely punctulate, punc- 
tures not denser laterally; apex feebly emarginate or truncate in cir- 
cular are, very finely margined; sides quite evenly arcuate through- 
out when viewed from above, slightly converging to apex, finely 
margined ; base more or less arcuate, feebly and very broadly sinuate 
ut middle, rather finely margined, about one-fourth wider than the 
apex and equal to the length; apical angles not prominent anteriorly, 
obtuse but not rounded; basal angles obtuse and more or less feebly 
rounded. 

Propleure smooth, obsoletely punctate and sometimes slightly 
rugulose. 

Elytra elongate to broadly oval, widest at the middle; dase trun- 
cate and feebly bisinuate, usually equal in width to the contiguous 
prothoracie base; humeri obtuse and not prominent; sides evenly 
arcuate, apex feebly produced and rounded; disc convex, frequently 
broadly depressed, suture also occasionally impressed, arcuately 
declivous posteriorly ; surface finely, irregularly and evenly punctate, 
frequently with closely placed series of punctures, the intervals with 
a more or less irregular series, punctures rarely denser at the sides 
and apex, never asperate. 

Epipleure moderately wide at the humeri, gradually narrowing to 
apex; surface sparsely and rather evenly punctate. 

Sterna and parapleure more or less irregularly punctate and 
rugose. 

Abdomen horizontal; surface finely, sparsely or obsoletely punctate, 
punctures denser at the periphery of the fifth segment, more or less 
rugulose. 


426 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Legs moderate in length. Femora not densel¥ punctate, the an- 
terior mutic in both sexes: protibial spurs subequal; tarsi dissimilar 
in the sexes, 

Male.—Elongate. Elytra twice to a little more than twice as long 
as wide, arcuately and not suddenly declivous posteriorly. Abdomen 
moderately convex and broadly impressed on first three segments. 
Posterior tibize more or less arcuate. First joint of the protarsi 
somewhat produced at tip beneath, bearing a tuft of golden pubes- 
cence, which obliterates the plantar groove at that point. 

Female.—Robust, often fusiform. Elytra broadly oval, often one- 
half longer than wide, very suddenly arcuately or vertically declivous 
posteriorly. Abdomen rather strongly and evenly convex. Posterior 
tibiz not arcuate. Anterior tarsi unmodified, 

Measurements —M ales: Length, 27-35 mm.; width, 8.5-9.5 mm. 
Females: Length, 25-30.5 mm.; width, 10-12 mm. 

Genital characters, male—KKdeagophore with the apex of the 
apicale moderately wide and evenly rounded, slightly recurved in 
apical fourth and punctulate above; general surface feebly convex 
although broadly depressed. Otherwise as in gigantea. 

Female.—Genital segment with the general characters as in 
gigantea, 

Valvula (Plate 1, fig. 7).—Dorsal plate subfalcate, surface oblique— 
facing upwards and outwards, quite densely and rather coarsely 
punctate in apical half, each puncture with a very small decurved, 
subrecumbent seta; internal margin not strongly defined; the internal 
or pudendal surface of the valve very densely punctate and setose as 
above, scarcely excavated below the apex. 

Appendage and fossa minute, feebly visible from above. Other- 
wise as in gigantea. 

Habitat—Arizona (Kearn’s District, Navajo Indian Reservation, 
April, A. W. Barber; Peach Springs, July; Santa Rita Mountains, 
June, Hubbard and Schwarz; Phoenix, collection E. C. Van Dyke; 
Fort Grant, July, Hubbard and Schwarz; Tucson, March; Sulphur 
Springs Valley, May, Hubbard and Schwarz: Oracle, July; Galliuro 
Mountains, May, Hubbard and Schwarz; Fort Yuma, Prescott, June, 
Barber and Schwarz); New Mexico (Las Vegas, August, collection 
H. S. Barber; Coolidge, Luna, Hubbard and Schwarz; Santa Fé, 
collection C. V. Riley) ; Colorado (Colorado Springs, June, Hubbard 
and Schwarz; elevation 6,000-7,000 feet, Wickham; La Junta, 
Southwestern (Strecker) ; Holly, Fort Collins, San Luis Valley, and 
Trinidad, Wickham’s list); Texas (Marfia, July, elevation 4,600— 
4,800 feet, H. F. Wickham; Alpine, July, elevation 4,400—6,000 feet) ; 
Kansas (Western, Hubbard and Schwarz) ; Oregon ? (one specimen 
bearing this state label in the collection of the U. S. National 
Museum); Mexico (Northern Sonora, Morrison; San Luis Potosi; 


> 


REVISION OF ELEODIINI—BLAISDELL. 427 


Hacienda de Bleados; Parras; San Pedro in Coahuila, Doctor 
Palmer). 

Number of specimens studied, 86, 

Type is in the LeConte collection. 

Type-locality.—* Flumen Gila” (LeConte). 

Salient type-characters.—Elongate, black. Thorax obsoletely punc- 
tulate. Elytra finely punctulate, punctures subseriate (LeConte). 

Diagnostic characters.—Closely related to gigantea, from which 
it differs by the frequent elongate fusiform or subcylindrical shape. 
The females frequently have the elytra more or less flattened on the 
dorsum. 

The prothorax is less arcuate at the sides, chiefly because the dise 


‘is quite strongly and transversely convex, with sides more or less 


strongly and arcuately deflexed and in many instances more or less 
feebly inflexed as well, so that the margin is completely invisible 
when the prothorax is viewed vertically from above. 

The pronotum is only apparently longer than wide, for by direct 
measurements and in tracings I do not find much difference between 
longicollis and gigantea. 

There is usually only a very slight antero-posterior convexity and 
this is a good differential character in separating the species from 
gigantea, but it can not be relied upon entirely. 

Some specimens from the Colorado Desert Region and Orange 
County, California, are as nearly intermediate between Jongicollis 
and gigantea as any forms can be, and these mesotypes approach 
estriata as well. 

I consider the two species as modifications of a single and rather 
recent ancestral form, one ramus of this dichotomous divergence 
extending westward and northward, the other eastward and north- 
ward. 

The sculpturing of /ongicollis is the same as in gigantea—surface 
smooth, polished, finely and sparsely punctate, but more coarsely 
sculptured specimens are frequently found, and some are even sub- 
striate. ; 

The specimens described by LeConte as haydenii is a variation 
along this line; the differential characters given are as follows: 
“Thorax is more rounded on the sides, more distinctly punctulate, 
the rows of punctures of the elytra are much more evident, and the 
curvature from the back to the sides is abrupt, while in /ongicollis 
it is regular and uniform; the elytra are also more elongate near the 
apex, so as to produce at the inflexed margin a very slight concavity.” 
It is only a synonym. 

Specimens collected by Doctor Palmer in Mexico had the elytra 
distinetly but finely punctato-striate; other forms were taken but all 
referable to the present species. 


428 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


General observations.—The mentum appears to Be more trapezoido- 
parabolic, otherwise as in gigantea. 

The prosternum is also about the same as in that species, probably 
not as prominent ventrally, and the ante-coxal portion more gradually 
and evenly continuing the surface line; frequently more strongly 
mucronate behind, the mucro may be conical and more or less de- 
flexed, or vertically truncate behind with the angle mucroid and de- 
flexed. In the series before me the prosternum is not even in one 
instance horizontal between the cox, but always more or less convex. 

The mesosternum is variable, at times quite flat to vertically arcu- 
ate, broadly, and more or less strongly concave. 


The abdominal process is wider than long and about a fourth of 


its own breadth wider than the metasternal salient. 

Phe post-coxal portion of the first abdominal segment is equal in 
length to that of the second; the latter is about a fifth longer than 
the third and almost twice as long as the fourth. 

The metasternum laterally between the coxze is about as long as 
the width of a mesotibia at apex. 

The profemora are subclavate and not noticeably swollen, greatest 
dorso-ventral diameter is at the junction of the middle and apical 
thirds, the surface lines are feebly arcuate; the tibial grooves are 
well developed, their margins converging and becoming contiguous 
some distance from the base. 

In the series at hand all of the tibiz are more or less arcuate. The 
protibiz are slightly compressed, obsoletely carinate externally, and 
the tarsal grooves are not evident. 

The mesofemora narrow but slightly toward base and not at all 
swollen; the grooves are rather short, their rather coarse margins 
converging and becoming contiguous near the middle, the floors are 
slightly concave. } 

The mesotibiz are quite cylindrical in section; their tarsal grooves 
are more or less evident as glabrous, opaque grooves limited by 
asperities. 

The metafemora are almost subequal in width throughout, the 
grooves are rather short, the margins converging to become almost 
evanescent before uniting at the distal side of the middle. 

The metatibiz are subcylindrical in section, slightly flattened ex- 
ternally and asperate. 

The articular cavities are quite closed. 

The tarsi are variable in thickness, usually moderately long and 
somewhat stout. In one specimen collected at Tucson, Arizona, the 
tarsi are distinctly slender. 

The protarsi are about a fourth of their length shorter than a 
mesotarsus. Joints 2-4, inclusive, subequal in length and very 


ated Ae A 


Pee > eee) ee 


eee = ph Per i 


REVISION OF ELEODIINI—BLAISDELL. 429 


slightly wider than long and together quite equal to the fifth; the 
first joint is short, about a half longer than the second. 

A mesotarsus is about a tenth of its length shorter than a meta- 
tarsus. Joints 2-4 subequal in length and scarcely longer than wide, 
together apparently just a little longer than the fifth; first joint 
almost as long as the second and third taken together. 

The metatarsi are half as long as their respective tibie. Joints 
2 and 3 are about subequal in length and distinctly longer than 
wide, together about equal in length to the fourth or first. 


ELEODES INNOCENS LeConte. 


Bleodes innocens LeConte, Smithson. Miscell. Coll., No. 167, 1866, p. 114.— 
Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 312. 

Elongate, suboblong to ovate, subopaque to feebly shining, moder- 
ately convex. Elytra striato-punctate. 

Head nearly twice as wide as long, feebly convex, more or less 
feebly impressed laterally and along the frontal suture, finely, rather 
densely and more or less irregularly punctate, antenne stout, reaching 
to the prothoracic base, outer four joints scarcely or very feebly com- 
pressed and not noticeably dilated; third joint shorter than the next 
two taken together: fourth to the seventh, inclusive, subequal and 
subeylindrical; eighth just noticeably shorter; ninth and tenth sub- 
orbicular; eleventh oval. 

Pronotum quadrate, widest at or a little in advance of the middle, 
very little wider than long; disc very moderately and quite evenly 
convex, at times noticeably declivous at the apical angles, very finely 
to obsoletely punctulate; apex truncate to feebly emarginate later- 
ally within the angles, obsoletely margined; sides more or less feebly 
and evenly arcuate from base to apex, often rather straight behind 
the middle, very finely margined; base truncate to feebly arcuate 
and finely margined, little wider than the apex and about equal to 
the length; apical angles obtuse, scarcely rounded and more or less 
prominent, not dentiform; basal angles obtuse, not rounded nor 
prominent. 

Propleure obsoletely punctulate and more or less rugulose, feebly 
convex and very feebly defined from the pronotal dise by the fine 
marginal bead. 

Elytra oval, smooth, about twice as long as wide, widest at or a 
little behind the middle; base feebly emarginate or truncate, fre- 
quently slightly wider than the contiguous prothoracic base; Auwmers 
obtuse, not rounded; sides evenly arcuate, apex not produced and 
rather narrowly rounded; disc slightly flattened on the dorsum and 
quite strongly convex laterally, arcuately, and vertically declivous 
posteriorly; surface striato-punctate, the strial series moderate in 


430 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


size and closely placed, usually more or less ereéded, intervals more 
or less feebly convex, finely, sparsely, and obsoletely punctulate; 
striz more or less impressed, the punctures slightly denser and ir- 
regular on the apical declivity. 

Epipleure smooth and usually impunctate, moderately wide, with 
margins parallel in middle two-fourths, thence gradually converging 
to near the apex. 

Sterna more or less punctate and rugulose. 

Parapleure rather coarsely punctate. 

Abdomen more or less obsoletely punctulate on the first three seg- 
ments, fourth and fifth more or less punctate; rather strongly rugu- 
lose on the first segments, the others obsoletely so. 

Legs moderate in length and stoutness. Profemora more or less 
feebly sinuate in both sexes; tibial spurs rather short, the anterior 
similar. Tarsi slightly dissimilar in the sexes. 

Male.—Elongate, somewhat narrow and suboblong. Elytra usually 
widest at about the middle. Abdomen oblique, feebly convex, 
strongly and broadly impressed on the first three segments. Anterior 
tarsi with the first two joints rather densely clothed at tips beneath, 
with yellowish pubescence, that more or less obliterates the plantar 
groove. 

Female-——Somewhat robust and more or less elongately ovate. 
Elytra usually widest behind the middle. Abdomen horizontal, 
rather evenly but not strongly convex, scarcely longitudinally convex. 
Tarsi simple. 

Measurements.—M ales: Length, 17-17.5 mm.; width, 6-6.8 mm. 
Females: Length, 19-19.5 mm.; width, 7.5-8 mm. 

Genital characters, male.——Edeagophore elongate, oval-fusiform 
and not strongly arched. 

Basale oblong oval, sides feebly arcuate and evenly convex from 
side to side. 

Apicale (Plate 4, fig. 20).—Subequilaterally triangular, quite 
strongly depressed; surface with a rather broad fusiform membra- 
nous depression at middle three-fifths, laterally more or less convex, 
with a few scattered and rather coarse punctures; the general surface 
is flattened and somewhat declivous at the terminal fifth; sides rather 
strongly arcuate at middle third, thence to the base rather straight, 
with small accessory lobes visible laterally at basal third, toward 
apex rather deeply sinuate; apex somewhat produced and narrowly 
rounded at tip; base arcuately lobed at middle and sinuate laterally. 

Sternite somewhat transversely semicircular in outline. Each lobe 
triangular with the external border evenly and obliquely arcuate 
from base to apex; internal border more or less straight and con- 
verging to the mid-line of the base; apex narrowly rounded; surface 


beat hd 


REVISION OF ELEODIINI—RBLAISDELL. 431 


diagonally convex, basal half smooth and shining, impunctate, the 
apical moiety feebly and inwardly declivous, quite densely and 
coarsely punctate, setose, sete long and dense at apex. Membrane 
not setose, sinus triangular. 

Female.—Genital segment (Plate 4, fig. 21) subparabolie in out- 
line—exclusive of the apices, not setose and strongly chitinized. 
Superior pudendal membrane broadly exposed. 

Valvula—Dorsal plate suboval to oval-oblong; surface more or less 
concave, with sides very slightly reflexed, shining and very sparsely 
punctate; external border rather evenly arcuate; internal margin 
more or less arcuate, apical margin rounded with angle obsolete; 
there are a few coarse punctures about the basal margin. Apex 
strongly produced, excurved, divergent and slightly decurved, convex 
above, broadly rounded at tip and chitinous, more or less excavated 
externally at base by the fossa, which has its margins very finely and 
inconspicuously setose. 

Appendage punctiform. The valvular membrane is exposed on 
the internal surface of the valve, and is more or less chitinous and 
impunctate. : 

Superior pudendal membrane is triangularly exposed, longitudi- 
nally and reticulately rugulose. 

Ventrolateral surfaces convex and not inflated, more or less im- 
pressed laterally, sparsely and irregularly punctate, glabrous; sub- 
marginal groove well developed beneath the external border of the 
dorsal plate. Apex more or less concave beneath. Internal margins 
of the valves contiguous to near the apex, genital fissure small, nar- 
row, and subapical. 

Habitat—Lower California (Cape St. Lucas, LeConte and Horn; 
El Taste, Gustav Beyer; Sierra El Chinche, collection of Charles 
Fuchs). 

Number of specimens studied, 8. 

Type in the LeConte collection. 

Type-locality.—Cape St. Lucas. 

Salient type-characters.—Elongate, somewhat shining; thorax 
quadrate, apex emarginate, apical angles acute and not acuminate, 
sides broadly rounded, also the base, basal angles obtuse; elytra 
elongate oval, strongly declivous behind, apex not prolonged, strize 
strongly punctate, slightly impressed, interstices with a few sparsely 
placed punctures. Anterior femora mutic (LeConte). 

Diagnostic characters.—lnnocens is the only species of the sub- 
genus with striate elytra at present known to our fauna; it is much 
smaller than gigantea and duller in luster. Horn writes that the 
anterior femora are feebly sinuate, but I can not see that this character 
is any more marked here than in the other members of the group. 
By the nearly mutic femora it is recognized from the members of the 


432 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


subgenus /leodes, except sponsa, caudifera, and longipilosa which 
have the profemora more or less obtusely armed or sinuate, and 
besides have a rougher sculpturing, with differently formed elytral 
apices, which render the diagnosis easy; in innocens the elytra are 
quite suddenly and arcuately declivous and never in the least pro- 
duced. 

Care must be taken not to confound specimens of the peninsular 
form of insularis with the present species. This mistake has already 
been observed in collections. 

There is at hand a female znsularis, collected at Sierra San Lazaro, 
Lower California, which has a subopaque luster and the elytra striato- 
punctate, very similar to the sculpturing observed in innocens; the 
apical angles are more acute and anteriorly prominent than in the 
latter species. I had to resort to the examination of the genital seg- 
ment to feel satisfied as to its true relationship. 

In innocens the antenne are proportionally stouter than in most 
species. . 

In énsularis (peninsular form) the profemora are more distinctly 
sinuate or obtusely dentate. 

General observations—The mentum is variable, triangular, 
triangulo-parabolic, sometimes triangulo-trapezoidal in outline; the 
surface may be feebly convex to foveate laterally with a median 
convex ridge, rather finely and obsoletely to rather coarsely punctate, 
each puncture with a short seta. Very variable considering the small 
series before me. 

The prosternum moderately convex antero-posteriorly in the 
median line, rather more gibbous and moderately wide between the 
cox, more or less grooved, scarcely widened behind and irregularly 
sculptured; more or less produced posteriorly, the mucro scabrous 
and more or less appressed to the body. 

Mesosternum almost flat to feebly oblique, broadly and moderately 
concave. 

The abdominal intercoxal process is subquadrate and about a third 
of its own width broader than the mesosternal salient; it is also equal 
in length to the second abdominal segment. The post-coxal part of 
the first segment is equal in length to that of the third, the latter 
just a little longer than the fourth. 

.The metasternum laterally between the coxe is about equal in 
length to the width of a mesofemur at base. 

The profemora are gradually and moderately clavate externally; 
the tibial grooves are well developed, their margins well defined by 
finely cariniform edges, which gradually converge to become contig- 
uous at the femoral base; their floors are quite flat, scarcely concave. 


a 


a 


REVISION OF ELEODIINI—BLAISDELL. 433 


The mesofemora are gradually and slightly thickened externally ; 
the grooves are structurally similar to those of the profemora, but 
the edges become contiguous a short distance from the femotal base. 

The metafemora are but slightly narrowed at base, and therefore 
have the opposed surfaces subparallel; the grooves are as on the other 
femora, but- have the edges evanescent near the basal fourth before 
becoming contiguous. , 

The protibie are obsoletely carinate externally and the tarsal 
grooves more or less obsolete. 

The mesotibiz are subeylindrical and very slightly increasing in 
diameter from base to apex; the grooves are faintly indicated and 
the surface asperate. 

The metatibiw are subcylindrical, narrowing slightly toward base; 
the grooves are feeble and exist as narrow linear impressions between 
the marginal asperities. 

The articular cavities of all the tibiz are quite closed. 

The tarsi are moderate in length and rather stout. 

The protarsi are about two-sevenths of their length shorter than a 
mesotarsus. Joints 2, 3, and 4 subequal in length and just slightly 
wider than long; the first is about one-half longer than the second; 
the fifth about equal in length to the third and fourth taker together. 

The mesotarsi are a_ little shorter than a metatarsus. Joints 2, 3, 
and 4 subequal in length and slightly decreasing in width in the order 
named, together subequal to the length of the fifth; the first about 
one-half longer than wide and a little shorter than the second and 
third taken together. 

A metatarsus is at least half as long as its tibia. Joints 2 and 3 
are about equal in length and breadth, together quite equal to the 
length of the first or fourth. 

Nore.—I sent to Mr. Blanchard an example of the peninsular form, 
which I refer to énsularis for comparison with énnocens, with the fol- 
lowing result: “It is surely a smooth innocens// The types are 
1 male, 1 female, Arizona, having the elytra subsuleate, anterior 
femora of the male angulate at outer three-fourths. In both sexes 
the same femora markedly flattened and strongly margined beneath.” 

I had labeled the specimen Z. insularis Linell, and Mr. Blanchard 
still further writes: “ I have been looking at the descriptions of in- 
nocens and insularis. Tt seems that the former was described from 
Cape St. Lucas, but only specimens now in the collection are from 
Arizona. The male does not have the large dense brush of golden 
hairs on the first joint of the front tarsi as is said of énsularis.” 

Doctors LeConte and Horn both give Cape St. Lucas as the type- 
locality for ‘nnocens. The specimens which I consider as true repre- 
sentatives of the species are quite like the peninsular form of insw- 

59780—Bull. 68—09——28 


434 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


laris in general habitus, but more elongate and more opaque, and 
these specimens are from Cape St. Lucas and El"Taste, Lower Cali- 
fornia. *I do not believe true ‘nnocens occurs in Arizona. Doctor 
Horn referred the peninsular form of insularis to gentilis and quadri- 
collis. This form of insularis occurs in the same region with énno- 
cens; both were collected at El Taste by Gustav Beyer. If my con- 
clusions should prove incorrect I should be very much surprised. 


Subgenus DISCOGENIA LeConte. 


Mentum transverse, trilobed, inflexed lateral lobes rudimentary, 
invisible without dissection. Thorax transverse. Elytra with the 
humeri rounded, rarely obtuse. 

Profemora armed or sinuate in the male; feebly armed or mutic 
in the female. . 

The internal spurs of the meso- and metatibie are slightly longer 
than the external. 

Subgeneric genital characters, male-—The characters, although not 
distinctive of the present subgenus, may be stated as follows: A picale 
of the edeagophore triangular, depressed, with a more or less widened 
median membranous groove. 

Female.—Genital segment triangular in outline, glabrous, not 
setose, and fully chitinized; valvular apices strongly produced, diver- 
gent and excurved. 

Superior pudendal membrane reaching at least to the middle of 
the dorsal plate. Internal ventral margins of the valves contiguous. 
Genital fissure closed and subapical. Appendages minutely puncti- 
form. 

The subgenus as at present defined contains three species that are 
not strictly homomorphic, and may therefore be divided into two 
groups as follows: 

Thorax widest at the middle, sides evenly arcuate from base to apex; first 
joint of the protarsi distinctly thickened at apex beneath, more strongly so 
in the female. ii 2 cee bk ss ee a ee Group A. 

Thorax widest just in advance of the middle, sides slightly convergent behind 
and briefly sinuate just in front of the basal angles; protarsi with the first 
one or two joints slightly thickened at apex beneath, more strongly so in 
[Cee ab: a ea ee eee Group B. 
Group A contains marginata and scabricula; group B the somewhat 

remarkable planipennis. 

The taxonomical arrangement of the subgenera constituting the 
genus /Hleodes is that in the order of morphological sequence, pri- 
marily based upon the structural modification of the valves of the 
genital segment in the female. 

The subgenera most closely related to Discogenia are Metablapylis 
and WSteneleodes. The approach to the closely related genus 


REVISION OF ELEODIINI—BLAISDELL. 435 


Embaphion is gradual and complete, apparently constituting a 
perfect evolutionary series. 

In Metablapylis with nigrina as the central form the valvular 
apices are short, rounded at tip, convex above and somewhat concave 
beneath, membrano-chitinous, scarcely divergent or produced. 

In Steneleodes with gigantea as typical the apex of each valve is 
broad, flattened, rounded at tip, convex above and concave beneath, 
strongly chitinized, and distinctly divergent and moderately produced. 

Taking marginata as typical of Discogenia, the valvular apex is 
moderately broad, slightly flattened, rounded at tip, convex above 
and concave beneath, strongly chitinized, divergent and excurved, 
strongly produced. 

In Embaphion the general characters are similar to the latter, but 
the strongly produced and excurved apex is more laminate and more 
strongly defined from the dorsal plate of the valve. 

The other segmental characters are distinctive in each subgenus 
and render the characters just enumerated as valuable differential 
criteria. 

An examination and analysis of the plates which form an im- 
portant part of this paper, will elucidate what has been stated above 
and also demonstrate that the species preserve a wonderful com- 
munity of subgeneric genital structure. 

The species constituting Group A may be differentiated as follows: 
Smaller species, scarcely robust, more or less shining, subasperately sculptured ; 

elytra distinctly gradually narrowed posteriorly, apex quite acute. Males 

LSI Con | w 0:2 Ba I ea Saree a a ae marginata, 
Larger species, rather robust, more or less opaque, asperately sculptured ; elytra 

moderately narrowed posteriorly, apex obtuse. Both sexes more or less 

ae ape Re Bg SE a ae sa ee a ee A scabricula, 

The single species constituting Group B can be recognized from the 
group characters; the most noticeable characteristic of planipennis 
is the depressed elytra. 

The following synopsis is probably the most practical one that 
ean be formulated from the specific genital characters for any sub- 
generic section of the genus /Jeodes, and is constructed from the 
female characters alone: 

Appendages rudimentary and without setx. 

External margin of the dorsal plate continuous with that of the apex, not 
angulate, not excavated dorsally at base of apex. Appendages puncti- 
ee ep es Ste RE We EES SUP See Ca!) ees Seen ee marginata, 

External margin interrupted, angulate; apical margin of the dorsal plate de- 
fined by a deep excavation at base of apex; appendages larger__scabricula. 

Appendages mammilliform, with pencil of set at tip- __.._planipennis. 

The males apparently have no constant specific differences; at least 

I have not been able to differentiate any. 


436 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Relationships.—There is no doubt but that marginata and scab- 


: ie 
vicula are closely related, and have undoubtedly diverged from a. 


common ancestral form within recent times. The more obscure rela- 
tionships of planipennis have been referred to elsewhere. 

Distribution —Discogenia is a subgenus peculiar to California and 
extends eastward into the Sierra Mavis Mountains as far as Lake 
Tahoe (elevation 6,280 feet), possibly into western Nevada; in 
Alpine County to an altitude of 7,000 feet (Blood’s meadow). 

I do not consider planipennis as a normal constituent of the pres- 
ent subgenus; if it should prove to be, then it must be considered as 


an eastward modification. In the latter case the first statement will — 


have to be altered. When the obscure question of relationships shall 
have undergone greater solution, then we will be able to speak more 
intelligently upon this problem. 

Thus far I have only seen specimens of marginata from the mari- 
time regions of central and northern California. The nucleus of 
distribution seems to be about the Bay of San Francisco. 

Scabricula is both montane and submaritime, evidently confined 
chiefly to central California, and undoubtedly extending more or 
less north and south of this central region. 

Planipennis has only come to me from Arizona, New Mexico, and 
Colorado. 

ELEODES MARGINATA Eschscholtz. 


Eleodes marginata Escuscuoirz, Zool. Atlas, III, 1833, p. 10. 
HEIM, Bull. Soc. Nat. Moscow, XVI, 1848, p. 268.—LEConrTE, Proc. Acad. 
Nat. Sci. Phila., 1858, p. 182. 

Eleodes fischeri MANNERHEIM, Revue Zool., III, 1840, p. 157; Bull. Soe. 

Nat. Moscow, XVI, 1848, p. 269. 

Discogenia marginata LeContre, Smithson. Miscell. Coll., No. 167, 1866, 

p. 117.—Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 320. 


Black, oval, and more or less elongate. 

Head twice as wide as long, equal to one-half the width of the 
prothorax, plane to feebly convex, shining, more or less impressed 
laterally near the frontal suture, sparsely punctate, confluently so 
at the sides. Antenne reaching to the posterior fourth of the pro- 
thorax, outer joints gradually and feebly dilated; the third about 
three times as long as the second; fourth twice as long as the same; 
fifth, ninth, and tenth subequal; sixth, seventh, and eleventh equal in 
length and slightly longer than the fifth. 

Pronotum feebly shining, widest at the middle, about two-fifths 
wider than long; dis¢ evenly and not strongly convex, rather coarsely, 
irregularly, at times confluently punctate, laterally not broadly and 
very feebly impressed, and there scabrous from minute granules; 
apex moderately and evenly emarginate, finely margined; sides 


. REVISION OF ELEODIINI—BLAISDELL. 437 


evenly and broadly rounded, narrowly and acutely margined, margin 
reflexed; base more or less feebly arcuate, about one-eighth of its- 
width greater than the apex; apical angles nearly obtuse; basal 
angles obtuse and more or less minutely prominent. 

Propleure smooth, shining, sparsely and irregularly granulate; 


- more or less rugulose over the acetabular convexities. 


Elytra oval, shining, widest at the middle; Awmeri obsolete, 
rounded; base not margined, somewhat declivous, and scarcely wider 
than the contiguous prothoracic base; seutellum rather large and 
triangular; sides evenly arcuate, rather gradually converging at 
apex, the latter gradually and somewhat produced, rather narrowly 
rounded; disc quite evenly and moderately convex, more strongly and 
quite evenly rounded laterally, arcuately and more or less obliquely 
declivous posteriorly; surface rather coarsely and somewhat densely 
punctate on the dorsum, becoming gradually more coarsely and sub- 
asperately muricato-granulate laterally, and without any sign of 
order. 

Epipleure moderate in width at the humeri, gradually narrowing 
to apex; surface feebly concave beneath the humeri and subasperately 
punctured. 

Sterna more or less strongly rugulose and punctate; punctures 
setigerous. 

Parapleure irregularly and rather strongly punctate, interstices 
glabrous and shining. 

Abdomen rather coarsely, thickly punctate and rugulose on seg- 
ments one and two, less so on the third, very sparsely so on the fourth 
and fifth. 

Legs moderate in length and thickness. Anterior tibial spurs simi- 
lar in the sexes the anterior spur longer than the posterior, both 
acute and rather stout. Protarsi nearly similar in the sexes, both 
with the first joint distinctly thickened at tip beneath. 

Male.—Elongate oval and more or less pointed posteriorly. Elytra 
more or less slightly wider than the thorax, evenly, obliquely, and 
arcuately declivous posteriorly, with the apex acute and more or less 
feebly produced. Abdomen moderately oblique and convex, feebly 
and broadly flattened on first two segments. Anterior femora armed 
with an obtuse tooth. Protarsi with the first joint feebly and trans- 
versely produced at apex beneath, plantar groove obliterated apically, 
space between the apico-marginal tufts of spinules simulating it. 

Female-—Somewhat robust. Elytra rather broadly oval and wider 
than the prothorax, slightly pointed, arcuately and slightly obliquely 
declivous posteriorly. Abdomen horizontal, quite strongly and evenly 
convex. Profemora mutic, scarcely sinuate. Protarsi with the first 
joint distinctly and transversely produced at tip beneath, plantar 
groove quite obsolete. 


438 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Measurements.—Males: Length, 18.5-19 mm. ; width, 6.5-T mm. 
Females: Length, 16-17 mm.; width, 6.5-7 mm. 

Genital characters, male-——KEdeagophore of the usual oblong-ovate 
form. 

Basale variable, oblong, more or less feebly convex, with sides very 
slightly arcuate. 

Apicale triangular, sides nearly straight, shghtly swollen toward 
base; apex more or less acute; base with middle lobe rather small 
and rounded, sinuate laterally; dorsal median groove moderately 
wide apically and linear toward base. 

Clave stouter than usual, shorter and not so attenuate toward apex. 

Sternite (Plate 2, fig. 11) with each lobe rather long, narrowing 
somewhat toward apex, the latter rounded; external border evenly 
arcuate, sometimes slightly angulate at middle, thence oblique and 
straight from angle to apex; internal border arcuate. Strongly 
chitinized, surface feebly convex and thickly set with long stiff sete 
in apical half. Sete do not extend across membrane at bottom of 
the sinus. 

Female (Plate 2, figs. 12 and 13) genital segment triangular, not 
setose. 

Valvula.—Dorsal plate nearly horizontal, feebly concave or convex, 
impunctate; external border nearly straight and directly continuous 
with that of the apex, becoming slightly more prominent at the 
transitional point on account of the external margin of apex becom- 
ing slightly sinuate; internal margin slightly reflexed, feebly arcuate 
and directly continuous with the internal apical margin which is 
strongly arcuate on account of the excurvature; apical margin not 
in the least defined from apex, the dorsal surface of both being di- 
rectly continuous, the transitional area forming a concave surface. 

Apex produced, not strongly chitinized at base externally, ex- 
curved, dorsal surface more or less convex from side to side and 
directed upward and slightly outward; ventral surface concave, 
directed downward and inward; internal surface more or less mem- 
branous and set with very short chitinous sete ; tip narrowly rounded; 
fossa obsolete with surface not in the least excavated at its usual site. 

Appendage minute and punctiform, without sete and located on 
the external margin, projecting but slightly beyond the margins of 
its minute cavity. ? 

Superior pudendal membrane attaining the middle of the dorsal 
plate. 

Ventrolateral surfaces not inflated, not strongly convex and gradu- 
ally tapering from base to apex; submarginal groove scarcely visible. 
Internal margins of the valves contiguous. 


REVISION OF ELEODITNI—BLAISDELL. 439 


Genital fissure closed and very narrow, finely setose within. In- 
ferior pudendal membrane not visible when the valves are in addue- 
tion. 

Habitat.—California (San Francisco all the year around; Alameda 
County; Trinity County, Dr. E. C. Van Dyke). 

Number of specimens studied, 60. 

Type probably in the Eschscholtz collection. 

Ty pe-locality.—Near San Francisco, California. 

Salient type-characters—Thorax subopaque, slightly transverse, 
sides rounded, surface densely and more or less confluently punctate, 
scabrous and feebly impressed laterally along the margins. Elytra 
densely subscabrous with reclinate muricate granules, declivous and 
rather strongly acute behind. Anterior femora armed in the male; 
mutic in the female. 

Diagnostic characters.—Although usually smaller than scabricula, 
it is at times quite difficult to differentiate; this is best done by direct 
comparison: It is less strongly sculptured, thorax not so broad, 
antennz more slender, shorter, decidedly so in the female, last joint 
of the labial palpi is narrower and less triangular, legs are more 
slender. The males have the elytra narrower, more obliquely de- 
clivous and acute at apex, and the profemora armed with an acute 
tooth. 

The female has the elytra less broad, less suddenly, more obliquely 
declivous posteriorly and more produced. The profemora are mutic. 

The female genital characters will always decide the diagnosis. 
In the male the femoral teeth are more acute in marginata than in 
scabricula. I believe that the genital characters are quite constant, 
for in the large series examined no mesotypical characters were 
observed. 

Marginata is rather abundant when its habits are known. About 
San Francisco it is found around the roots of prostrate shrubs. 

Mannerheim’s fischeri was considered by LeConte as a large and 
well-developed form of marginata. The inaccessibility of the type 
precludes any efforts to determine its true relations. It might be a 
scabricula; if so Mannerheim’s name would have priority over that 
of LeConte’s. 

General observations —The mentum (Plate 6, fig. 20) has the 
middle lobe large and somewhat variable. Usually quite evenly 
rounded anteriorly, sometimes subtruncate, and again the sides may 
be somewhat straight; surface feebly convex and at times more or 
less foveate laterally. Lateral lobes rudimentary. 

The anterior margin of the prosternum is more or less strongly 
deflexed. The prosternum and mesosternum the same as in scabricuda. 

The metasternum laterally between the cox is short, and equal in 
length to the width of a mesotibia at middle. 


440) BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


The abdominal process is subquadrate and slightly transverse, 
its width is equal to the length of the third segment; the post-coxal 
part of the first segment is subequal in length to the third, the latter 
is one-third of its length shorter than the second; the fourth is a 
iittle more than one-half as long as the second. 

The abdominal intercoxal salient is one-fifth of its width greater 
than that of the metasternal process. 

The tibial grooves on all of the femora are quite strongly mar- 
gined, distinctly concave and glabrous; the margins are subasperate 
and converge gradually to the femoral bases. 

The anterior margins of the profemoral grooves are quite dis- 
tinctly carinate and not very strongly dentately laminate at about 
the outer sixth, thence more or less obsolete to apex. 

The external borders of-the protibize are more or less subearinate, 
those of the meso- and metatibie are rounded, muricately sculp- 
tured, and not in the least carinate. 

The protibie are slightly compressed, the others subcylindrical. 
The tarsal grooves are more or less indicated on all of the tibia, but 
most marked on the anterior. These grooves are all asperately 
sculptured. On the protibiz the more or less developed carinz pass 
to the outer margin of the tarsal grooves, while the internal margins 
are more or less serrulate. 

The anterior surfaces of the protibize are feebly subasperately 
sculptured, and the posterior surfaces are strongly asperate. 

Tarsi somewhat slender and moderate in length. 

The protarsi are about one-third of their length shorter than a 
mesotarsus. Joints 1 to 4, inclusive, are small, short, and slightly 
transverse; the first shghtly the thickest and longest, the second, 
third, and fourth subequal in length. 

A mesotarsus is equal in length to a metatarsus. Joint one slightly 
shorter than the third and fourth taken together; the second, third, 
and fourth subequal, short, and scarcely longer than wide; second 
slightly the longest. 

A metatarsus is about one-third of its length shorter than its meta- 
tibia. Joint one equal to the next two taken together; second and 
‘third subequal in length. 


ELEODES SCABRICULA LeConte. 


Eleodes scabricula LECONTE, Proc. Acad. Nat. Sci. Phila., 1858, p. 187. 
Discogenia scabricula LeContrr, Smithson. Miscell. Coll., No. 167, 1866, p. 
117.—Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 320. 


Black, oblong-oval. Head nearly one-half as wide as the pro- 
thorax, plane to shghtly convex, opaque, rather coarsely and irregu- 
larly punctate, punctures confluent laterally. Antenne not quite 


REVISION OF ELEODITINI—BLAISDELL. 441 


attaining the prothoracic base, outer four joints feebly compressed, 
slightly and gradually dilated, third joint scarcely as long as the 
next two taken together; fourth slightly longer than the fifth; the 
latter, sixth, and seventh subequal; eighth about as wide as long and 
subtriangular; ninth suborbicular in outline; tenth feebly trans- 
versely oval; eleventh ovate. 

Pronotum opaque, widest at the middle and about one-half wider 
than long; disc evenly and moderately convex, rather coarsely, 
densely, and more or less confluently punctate, the punctures coales- 
cing more or less transversely; sides broadly and slightly impressed 
throughout along the margins, and there finely scabrous from minute 
granules; apex moderately, broadly, and evenly. emarginate, finely 
margined ; sides evenly and broadly rounded, with the narrow margin 
reflexed; base very feebly arcuate, finely margined, and about one- 
fourth wider than the apex; apical angles obtuse and more or less 
feebly rounded; basal angles obtuse and not in the least prominent. 

Propleure smooth, more or less opaque, very sparsely and irregu- 
larly granulate, more or less rugulose and concave beneath the ap- 
parently explanate pronotal margins. 

Elytra oval, somewhat shining, widest at the middle; base not 
margined, slightly declivous, slightly wider than the contiguous pro- 
thoracic base; humeri very obtuse, scarcely evident; sides evenly 
arcuate, slightly convergent posteriorly, with apex rather broadly 
rounded; disc moderately convex and more strongly rounded later- 
ally, arcuately declivous posteriorly; swrface coarsely and somewhat 
densely punctate on the dorsum at center, becoming more and more 
scabrous laterally and about apex, the sculpturing consisting of 
densely placed reclinate and more or less muricate granules that are 
confused throughout. Frequently the disc is more or less flattened, 
and more strongly, less broadly, and somewhat more suddenly 
rounded laterally. 

Epipleure moderate in width beneath the humeri, and gradually 
narrowing to apex; surface rather thickly and submuricately punc- 
tate, the interstices glabrous and shining. 

‘Sterna scabrous from irregularly placed setigerous punctures and 
ruge. 

Parapleure coarsely punctate, frequently slightly muricate. 

Abdomen more or less moderately, and rather densely muricately 
punctate, less so on the fourth and fifth segments, coarser and most 
scabrous on the first. 

Legs moderate in length and thickness. Anterior femora more or 
less armed; anterior tibial spurs similar in the sexes, the anterior 
longer than the posterior, both acute. Anterior tarsi with the first 
joint more or less thickened at tip beneath and slightly dissimilar in 
the sexes. 


442 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Male——Elongate oval. Elytra scarcely wider than the prothorax, 
moderately convex, arcuately and obliquely decli¥ous posteriorly and 
noticeably attenuate. Abdomen moderately oblique, not strongly 
convex, more or less broadly and obsoletely impressed on first two 
segments. Anterior femora armed with an obtuse tooth or simply 
sinuate. Anterior tarsi with the first joint slightly thickened at tip 
beneath, with small acute tufts of modified spinules at the apico- 
marginal angles; the tufts do not completely obliterate the plantar 
groove. 

Female——Robust. Elytra broadly oval and noticeably wider than 
the prothorax, convex, less attenuate at apex, arcuately and quite 
vertically declivous posteriorly. Abdomen rather strongly convex. 
Profemora slightly sinuate and with a small obtuse tooth, or entirely 
mutic. Protarsi with the basal joint distinctly thickened at tip be- 
neath, and bearing a small tuft of modified spinules, plantar groove 


obliterated. 

Measurements.—Males: Length, 16.5-23.5 mm.; width, 7.2-10 mm. 
Females: Length, 20-23 mm.; width, 9-11 mm. 

Genital characters, male-—Edeagophore elongately oblong-ovate 
and more or less arched. 

Basale oblong-oval, moderately convex and variable; surface longi- 
tudinally grooved at middle in apical half. 

Apicale rather elongate, triangular, sides nearly straight, apex not 
produced ; base moderately and distinctly lobed at the middle, sinuate 
laterally; dorsal surface moderately convex, with a moderately wide 
median groove extending from apex nearly to the base. 

Sternite with lobes rather large, each distinctly longer than wide, 
rounded at apex, with the external margin nearly straight in basal 
half, thence arcuate; surface densely punctate and setose apically, 
setee moderately long and dense at tip. Membrane not setose across 
the bottom of the sinus. 

Female.—Genital segment subquadrate in outline exclusive of the 
apices, well chitinized, apex and dorsal plate continuous internally, 
not setose. 

Valwula (Plate 4, fig. 1).—Dorsal plate horizontal and oblong; 
surface varying from feebly concave to slightly convex, occasionally 
with a few Sian punctures; external margin arcuate in apical 
moiety and more or less sinuate to the base in basal half, where the 
submarginal groove is visible from above, making somewhat of a 
sigmoid curve upon the basal prominence of the lateral plate; in- 
ternal margin slightly or not at all inflexed and feebly sinuous; 
apical margin feebly defined from the dorsal surface of apex inter- 
nally, but distinct externally with the angle rectangular or narrowly 
rounded, 


—_—— 


REVISION OF ELEODIINI—BLAISDELL. 443 


Apex produced and completely chitinized, divergent and feebly 
decurved or recurved; dorsal surface convex, directed upwards and 
inwards; ventral surface concave and directed downwards and out- 
wards; externally and caudad to the apical margin of the dorsal 
plate, deeply excavated forming a fossa, at margin of which is the 
minute subpunctiform appendage, that scarcely protrudes beyond 
the margins of its articular cavity, and frequently it bears a couple 
of short sete at tip. 

Superior pudendal membrane reaches to the middle of the dorsal 
plates, the latter being contiguous apically, but divergent basally. 
Internal valvular membrane set with few fine, short sete. 

Ventrolateral surfaces evenly convex, with surface lines straight 
when viewed longitudinally, rather deeply excavated beneath the 
apical half of the external margin of the dorsal plate, the execava- 
tion extending along the ventral surface of the apex. 

Submarginal groove fine, sigmoid towards base. Internal margins 
of the valves contiguous; genital fissure a narrow cleft in apical 
third, the margins finely setose, sete soft and scattered. Inferior 
pudendal membrane not visible. 

Habitat—California (Lake Tahoe, Eldorado County, July, ele- 
vation 6,280 feet; Big Trees, Calaveras County, August, elevation 
4,702 feet; Blood’s Meadow, Alpine County, July, elevation 7,000 
feet; Tuolumne County, D. W. Coquillet and E. C. Van Dyke; 
Colony Mills, Tulare County, elevation 5,415 feet, Charles Fuchs 
and Ralph Hopping; Fresno, Alameda, Marin, and Sacramento 
counties). 

Number of specimens studied, 50. 

Type (female) in the LeConte collection. 

Type-locality.— Sacramento, California; collector, Mr. J. Wittick. 

Salient type-characters.—Thorax opaque, transverse, sides rounded, 
densely and confluently scabro-punctate, on each side broadly im- 
pressed. Elytra densely scabrous with reclinate granules, declivous 
and acute behind. Anterior femora with an obtuse tooth (LeConte). 

Diagnostic characters —Scabricula has to be carefully separated 
from marginata, which it resembles more than any other species, and 
this can best be done by comparison: It is more roughly sculptured, 
usually larger, last joint of the labial palpi triangular, antenne 
stouter and longer, thorax broader and the legs stouter, there is also 
less difference between the sexes. 

The males have the elytra narrower and obliquely declivous pos- 
teriorly and produced apically; the females have broader elytra, 
which are more suddenly declivous posteriorly and scarcely produced 
apically, the anterior femora are feebly armed. 

This species is undoubtedly characteristic of the western foothills 
of the Sierra Nevada Mountains and where the more typical form is 


444 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


found. Its presence about Sacramento and San Francisco Bay can 
undoubtedly be explained by carriage by the rivers that drain the 
western slopes of the mountains. Its area of distribution no doubt 
naturally overlaps that of marginata, the latter being a more or less 
maritime form. Specimens of scabricula indigenous to the valley 
regions are smaller and with greater difficulty separated from mar- 


ginata. 
The female genital characters are distinctive, those of the mrale are 
not. a» 


A male example recently loaned to me by Mr. A. Koebele, presents 
varietal characteristics as follows: Form as usual, distinctly depressed 
on the elytra and pronotum, the latter distinctly sinuate before the 
basal angles, where the sides are parallel for a short distance; elytra 
densely, asperately muricato-tuberculate, base transverse and the 
humeri rather prominent and obtusely rounded. sides subparallel 
nearly as in producta. The protibie are suddenly and distinetly con- 
stricted at base as in E'mbaphion; this character is also observed in 
planipennis. 

This form may be known as scabricula forma deplanata. 

Habitat.—Napa County, California. 

This specimen surely forms a connecting link betwen scabricula 
and planipennis. On the other hand it may be an aberration, and it 
remains for other examples to settle this question. 

General observations —The mentum has the middle lobe large and 
variable, usually rounded anteriorly from the middle; frequently 
subtrapezoidal and subtruncate anteriorly, even slightly emarginate 
at apex; surface may be feebly convex and simply punctate or more 
or less foveate laterally. The lateral lobes are very rudimentary. 

The anterior margin of the prosternum is usually more or less 
stoutly deflexed ; between the coxe usually strongly arcuate and more 
or less vertically declivous posteriorly and not in the least mucronate. 
The surface may gradually slope downward and backward from the 
anterior margin to between the cox, with which it is quite prominent 
ventrally, but less suddenly so than in planipennis. The prosternum 
is frequently strongly grooved between the coxe. 

In a specimen from Plumas County there is a small but well marked 
mucro present. In some specimens from Tulare County (Colony 
Mills) the posterior margin is feebly compressed and quite: vertical 
with a feeble attempt at a mucronate angle. The mesosternum is 
arcuately declivous and more or less concave. 

The metasternum laterally between the coxe is very short, about 
equal in length to the width of a mesotibia at apex. 

The abdominal intercoxal process is subquadrate and_ slightly 
transverse; its width is equal to the length of the third abdominal 
segment; the post-coxal portion of the first segment equal to the 


REVISION OF ELEODIINI—BLAISDELL. 445 


length of the second, the third about equal to the fourth, the latter 
one-third of its length shorter than the second. 

— *The abdominal salient is one-fifth of its own width greater than 
_ thatof the metacoxal process. 

4 The tibial grooves of the femora are distinctly concave, with well 
defined asperate or subserrulate margins. The margins of the ante- 
rior and middle grooves converge gradually to the femoral bases; 
those of the metafemora become evanescent at about the middle, 
thence to base the femoral surfaces are simply more or less flattened. 

The grooves of the profemora have their anterior margins more or 
less dentately laminate at about the outer fifth. 

The external surfaces of the protibizw are subcarinate; those of the 
meso- and metatibiz are rounded and scabrously sculptured, not in 
the least carinate. The protibie are slightly compressed; the others 
are subeylindrical in transverse section. ‘The tarsal grooves are rudi- 
mentary on all of the tibiz, but most marked on the protibixw; all are 
roughly sculptured. On the protibiz the more or less evident carina 
passes to the external margin of the groove, while the internal margin 
is quite distinctly serrulate. 

The anterior surfaces of the protibiz are much less roughly seulp- 
tured than the posterior surfaces. 

The articular cavities of the protibix are partly closed, those of the 
mesotibie almost closed, and those of the metatibie entirely so. 

Tarsi apparently variable in stoutness and independent of sex. 

The protarsi about one-third of their length shorter than a meso- 
tarsus. Joints 1 to 4, inclusive, are short and subequal in length; the 
first is apparently slightly longer and slightly produced beneath. 

The mesotarsi are about one-fourth of their length shorter than a 
metatarsus. Joints 2 to 4, inclusive, are subequal in length and 
slightly longer than wide; the first is slightly longer and stouter. 

The metatarsi are about two-fifths of their length longer than a 
protarsus and scarcely two-thirds as long as the metatibia. Joint 
one is distinctly shorter than the next two taken together; the second 
and third joints are subequal in length, and together about equal to 
the fourth. 

ELEODES PLANIPENNIS LeConte. 


Eleodes planipennis LeConte, Smithson, Miscell. Coll. No. 167, April, 1866, 
p. 116.—Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 316. 

Elongate oval to ovate, more or less shining, elytra strongly de- 
pressed. 

Head \ess than twice as-wide as long, quite flat between the more or 
less prominent sides of the frons, finely and sparsely punctate, punc- 
tures denser laterally and on the epistoma. Anfenne rather long, 
reaching a little beyond the prothoracic base, moderate in stoutness, 


446 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


scarcely compressed or incrassate, third joint slightly shorter than the 
next two taken together, fourth slightly longer than the fifth, the 
latter, sixth, and seventh subequal, eighth just a trifle shorter, ninth 
and tenth suborbicular, eleventh short-ovate. 

Pronotum cordately and transversely oval, widest in front of the 
middle, where it is scarcely one-half wider than long; disc moderately 
and quite evenly convex, finely and rather sparsely punctate, narrowly 
impressed along the sides and feebly granulate; apex rather deeply 
and evenly emarginate, more or less obsoletely margined, and one- 
eighth wider than the base; sides evenly and strongly arcuate in 
anterior three-fourths, thence to base obliquely and more or less 
sinuate, margin feebly but distinctly reflexed to near the base, the 
latter quite truncate and one-sixth to one-fourth shorter than the 
length; apical angles quite acute, not rounded, and slightly promi- 
nent; basal angles quite rectangular, not prominent nor rounded. 

Propleure smooth and impunctate or sparsely and feebly muricato- 
punctate and rugulose. 

Elytra oval, widest at the middle, less than twice as long as wide; 
base not emarginate, usually slightly wider than the contiguous pro- 
thoracic base; Auwmeri rounded; sides evenly arcuate, apex feebly 
produced and narrowly rounded; disc quite flat, sometimes very feebly 
convex on the dorsum, laterally rather narrowly rounded and quite 
suddenly inflexed, especially near the humeri, inflexed sides quite 
plane and oblique, evenly and more or less gradually arcuately 
declivous posteriorly, feebly sinuate at apex; surface finely and 
scarcely densely muricately punctate on the sides and apex, some- 
times quite densely so, punctures quite simple on dorsum along the 
suture. 

Epipleure smooth, obsoletely and very sparsely punctate; moderate 
in width beneath the humeri, thence gradually narrowing to apex. 

Sterna sometimes smooth, obsoletely punctate and rugose, at other 
times more coarsely sculptured. 

Parapleure more or less indistinctly punctate. 

Abdomen smooth, more or less dull in luster, very minutely, 
sparsely and more or less indistinctly punctate, more or less rugulose 
and horizontal. 

Legs moderate in length and stoutness, rather densely sculptured. 
Profemora more or less sinuate in outer fifth. Protibie quite dis- 
tinctly and rather suddenly constricted at base, the constriction 
opposed to the femoral sinuation; spurs similar in the sexes, slender 
and acute, the anterior slightly longer than the posterior. Protarsi 
nearly similar in the sexes. 

Male.—Body somewhat narrow. Elytra rather obliquely to gradu- 
ally declivous posteriorly. Abdomen very moderately convex and 
very feebly flattened at middle of the first two segments. Protarsi 


REVISION OF ELEODIINI—BLAISDELL. 447 


with the first two joints feebly but distinctly thickened at tips beneath, 
each bearing a small tuft of spinules at the apico-marginal angles, the 
second joint is less thickened than the first, in each instance obliter- 
ating the plantar groove. 

Female.—Body quite broadly oval. Elytra rather evenly arcuately 
declivous posteriorly. Abdomen rather strongly convex from side 
to side. Protarsi with the first joint slightly thickened at tip beneath, 
bearing a small tuft of spinules. 

Measurements.—Males: Length, 16-20 mm.; width, 6.8-8 mm. 
Females: Length, 16.2-19.5 mm.; width, 7.5-9 mm. 

Genital characters, male.—Edeagophore rather small, narrow, 
oblong-ovate, slightly arched. 

Basale oblong; surface not strongly convex, flattened more or less 
on the dorsum, sides nearly parallel. 

Apicale triangular, about one-half longer than wide, apical fourth 
more or less decurved; surface flattened, with a broad median mem- 
branous groove in basal half; sides nearly straight to tip, the latter 
broad and narrowly rounded; base rather narrowly lobed at middle 
and sinuate laterally. 

Sternite slightly transversely quadrate in outline. Each lobe rather 
long, with external border straight or slightly sinuate in basal half, 
thence evenly arcuate to apex; internal border symmetrically curved 
with the external border, both borders meeting in a rather narrowly 
rounded apex; surface very feebly convex, smooth, impunctate, and 
glabrous in basal moiety, apically coarsely and sparsely punctate, 
not densely setose, sete quite long. Sinus triangular, slightly closed 
at bottom by membrane which is not setose. 

Female.—Genital segment (Plate 5, fig. 6) triangular in outline, 
about one-half longer than wide, strongly chitinized, glabrous, valves 
contiguous. 

Valvula.—Dorsal plate elongately triangular, impunctate; surface 
directed slightly outward as well as upward; external border quite 
straight, slightly convergent to apex; apical border oblique, angle 
rounded, internally continuous with the chitinous apex, the latter 
produced, moderately excurved, convex from side to side and concave 
beneath, thin and sublaminate at tip, where it is rather broad and 
rounded, internal surface moderately and finely setose, dorsally 
without sete. 

Appendage small and conical, bearing three or four long sete at 
tip; fossa an excavation at base of the external surface of apex 
and contiguous to the apical border of the dorsal plate. 

Superior pudendal membrane attaining the apical third of the 
dorsal plate and more or less longitudinally rugulose, 

Basal prominences not developed. 


448 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Ventrolateral surfaces.—Body triangular, jyith surface lines 
straight, evenly convex from side to side; surface smooth and im- 
punctate; submarginal groove distinct and moderate beneath the . 
narrowly expanded external margin of the dorsal plate, terminating 
at the fossa. Apex membranous beneath its chitinous and expanded 
dorsal surface, finely setose; ventral plate distinctly defined apically. 
Internal margins of the valves contiguous, fissure short and apical. 

Habitat—Arizona (Williams, May, Barber and Schwarz; South- 
ern, coll, E. C. Van Dyke; Chiricahua Mountains, June, Hub- 
bard and Schwarz; at base of Humphreys Peak, August, elevation 
9,500 feet, F. H. Snow); New Mexico (F. H. Snow; Hubbard and 
Schwarz; Coolidge; Magdalena, August, F. H. Snow, Warren 
Knaus) ; Colorado (Manitou, Packard). 

Number of specimens studied, 25. 

Type (female) in the LeConte collection. 

T'ype-locality—New Mexico. 

Salient type-characters—HHead and thorax sparsely and _ finely 
punctate, subdepressed, apex broadly emarginate, base truncate, 
sides strongly rounded, posteriorly briefly sinuate, basal angles rec- 
tangular, small and not prominent. Elytra obliquely declivous 
behind and sinuately attenuate at apex, humeri obtuse, sides suddenly 
inflexed, along the suture finely punctate, at the sides and apex 
densely sculptured with small slightly elevated granules, anterior 
femora subsinuate (LeConte). 

Diagnostic characters—Quickly recognized from marginata and 
scabricula by the flattened elytral disc. The prothoracic apex is gen- 
erally slightly wider than the base, the sides are evenly arcuate 
anteriorly and converge somewhat posteriorly, becoming slightly 
sinuate in front of the basal angles, which are rectangular; dise 
widest in front of the middle. 

In some specimens the elytra are very feebly flattened—in fact, 
quite evenly but not strongly convex—and the smaller of such ex- 
amples greatly resemble the elongate form of snowii found in Ari- 
zona. The genital characters must here be relied upon for their 
separation. 

Planipennis has heretofore been considered a Blapylis. It is the 
nearest approach to E’mbaphion that we possess; in fact, it could 
with as much propriety form a section in the latter genus as the one 
in which it is now retained, and where it also must form a section 
intermediate between Discogenia and Embaphion. In genital char- 
acters it is an Hmbaphion, in facies an Eleodes. 

The constriction of the anterior tibizw at base is not peculiar to the 
genus Embaphion. It is observed to a greater or less degree in the 
different species of Blapylis; it is absent in the genus Hleodes, and 


REVISION OF ELEODIINI—BLAISDELL. 449 


appears to be a structural modification with a function whose analogy 
is found in the femoral teeth of that subgenus. 

It might have been a wise course to have created a new genus for 
its reception, but I am opposed to contributing unnecessarily to an 
increase in the complexity of our terminology when relationships can 
be understood without it. 

General observations —Mentum variable, the middle lobe is rather 
large, usually rounded anteriorly; many examples have the anterior 
margin truncate and often distinctly emarginate, with sides arcuate 
or obliquely straight; surface may be feebly and evenly convex, to 
strongly foveate laterally, with a well-marked meridian ridge. 

The prosternum is quite evenly rounded antero-posteriorly between 
the cox and very feebly widened behind the equator of the acetab- 
ula; it is quite prominent ventrally with the coxw; posteriorly it is 
vertically and scarcely arcuately declivous. In the series at hand 
there is not the least sign of a mucro. 

The mesosternum is arcuately declivous and more or less concave. 

The abdominal intercoxal process is comparatively small, sub- 
quadrate, and equal in width to the length of the post-coxal portion 
of the same segment, the latter about equal to the length of the third; 
the second is one-third of its length longer than the third; the fourth 
is equal to half the length of the second. 

The abdominal process is one-fifth of its width greater than that 
of the metasternal salient. 

The metasternum laterally between the coxe is short and about 
equal in length to the width of a mesotibia at middle. 

The femora have their tibial grooves well defined and glabrous, 
their margins are rather thin and inwardly converging quite to the 
femoral bases. Those of the profemora have the anterior margins 
feebly laminate at the usual site of the femoral tooth. 

The external surfaces of the tibiz are more or less carinate, most 
marked on the protibiz, which are also more or less compressed. 

The tarsal grooves are more or less indicated on all of the tibia, 
frequently extending two-thirds of their lengths, most strongly de- 
veloped on the protibia; frequently the meso- and metatibiw are 
simply flattened externally. 

Tarsi variable in stoutness, noticeably so in the larger specimens 
and somewhat slender in the smaller ones. 

The protarsi are about one-fifth of their length shorter than a 
mesotarsus. Joint one distinctly longer than the second, the suc- 
ceeding three about equal in length, just the least longer than wide; 
the fifth about as long as the preceding three taken together. 

The mesotarsi are about one-tenth of their length shorter than a 
metatarsus. First joint about one and a half times longer than the 
second, the latter to the fourth inclusive subequal and distinctly 


59780—Bull. 68—09——-29 


450 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


longer than wide; fifth slightly longer than the pgeceding two taken 
together. 

The metatarsi are about three-sevenths of their length shorter than 
a metatibia. First and fourth joints subequal, each about equal in 
length to the second and third taken together; the latter two are sub- 
equal in length. 


UNRECOGNIZED SPECIES. 
ELEODES VICINA LeConte. 


Eleodes vicina LEContrE, Ann. Lyc. Nat. Hist., New York, V, 1851, p. 
133.—Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 308.—CAsgEy, Ann. 
New York Acad. Sci., V, Noy. 1890, p. 595. 


ELEODES NITIDA Casey. 
Bleodes nitida Casey, Ann. N. Y. Acad. Sci., VI, Nov. 1891, p. 58. 
ELEODES REFLEXICOLLIS Mannerheim. 
Eleodes reflericollis MANNERHEIM, Bull. Soc. Moscow, XVI, 1843, p. 270. 
Genus EMBAPHION Say. 


Embaphion Say, Journ. Acad. Nat. Hist. Phila., III, 1824, p. 254.— 
LACORDAIRE, Genera des Coléopt., V. 1859, p. 152.—LeContr, Classif. 
Coléopt. North Amer. (Smithson. Miscell. Coll.), 1861, p. 226—Horn, 
Trans. Amer. Phil. Soc. Phila., XIV, 1870, p. 320—LeConTr and 
Horn, Smithson. Miscell. Coll. No. 507, 1883, p. 374. 


Thorax and elytra always acutely and sometimes broadly mar- 
gined, margin more or less reflexed. 

The epipleure always narrow, rarely defined from the inflexed 
sides of the elytra, except at apex, where they are always well defined, 
not suddenly widened at base and never attaining the humeral angles 
of the elytra. 

The above characters are distinctive of the genus and are not 
observed elsewhere in the Eleodiini. 

The constriction of the protibiz at base is not peculiar to Emba- 
phion, for it is also observed in Blapylis, and to a less degree in 
Discogenia. ; 

The mentum presents no distinctive characters; it is as variable here 
as elsewhere. The middle lobe is somewhat larger than in the true 
Eleodes, slightly smaller than in Discogenia; although the lateral 
lobes are not so strongly evident as in Eleodes, they are larger than 
in the subgenus Discogenia. 

The abdominal and metasternal processes are moderate and quite 
equal in width in the present genus, and this is also occasionally 
observed in Lleodes (fusiformis and opaca). 

The profemora are always mutic; the tibial spurs are quite similar 
in the sexes, The anterior spurs of the protibie are never noticeably 


— 


re 


REVISION OF ELEODIINI—BLAISDELL. 451 


thickened, although slightly longer and larger than the posterior. 
The internal spurs of the meso- and metatibia are always slightly 
longer than the external. The tarsi are similar in the sexes and 
simple. 

The abdomen is always on the same horizontal plane as the sterna; 
its segments are on the same plane in muricatum, contusum, con- 
tractum and planum; the third segment slightly oblique in glabrum 
and elongatum, and distinctly so in depressum. 

The genital characters are quite characteristic and immediately 
follow planipennis of the subgenus Discogenia in the evolutionary 
series; they are very different from those observed in the remaining 
genera of the tribe, and only phyllogenetically related to Blaps. 

The generic genital characters are as follows: 

Male.—A picale of the edeagophore triangular; dorsal surface more 
or less strongly convex, and without median groove; sternite bilobed; 
lobes united by a membranous interval. 

Female.—Genital segment triangular, or without the apices trian- 
gulo-trapezoidal, somewhat depressed ; dorsal surface quite plane, not 
setose. Apices of the valves produced, chitinous, more or less everted, 
convex above and concave beneath. Va/ves contiguous beneath; in- 
ferior pudendal membrane not visible. Genital fissure narrow and 
subapical. 

Superior pudendal membrane reaching to about the middle of the 
dorsal plates. Appendages short mammilliform, with a pencil of 
rather long hairs at their tips. 

Although the genital characters present slight specific differentia- 
tion, they can not at present be tabulated so as to be of any practical 
use. 

The species should be arranged in the following morphological 
sequence: depressum, elongatum, glabrum, contractum, planum, con- 
tusum, and muricatum. This is also an evolutionary series indicating 
genealogical relationship with Discogenia. 

Such an arrangement will present to the notice of the student that 
there is a gradual increase of the foliaceous margin from depressum 
to muricatum; as a consequence of this the apical emargination of 
the prothorax will increase from a simple broad shallow emargina- 
tion of the apex, so obvious in LJeodes, to the deep notch with almost 
parallel sides, about equaling in depth the length of the head, as seen 
in muricatum. 

In the narrowly margined species the prothorax is much broader 
than long (depressum), and with an increase in the width of the 
margin the pronotum proper becomes narrower, even longer than 
wide, 

The epipleural line follows the same rule; in the very narrowly 
margined species it is quite distinct, and indicated in the whole 


452 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


course either by a faint groove or a distinctly glevated line; it be- 
comes almost entirely absent in the broadly margined species. In the 
middle of the series the marginal line of the elytra does not attain 
the apex (glabrum, contractum, and planum) ; at the extremes of the 
series it does. 

In those species with feeble elytral margin the humeral angles are 
more distinct and even prominent (depressum), and less so as the 
margins increase. Note again what has already been said relative to 
the obliquity of the third abdominal segment (see p. 451 and below). 

Genealogy.—The scheme herewith presented illustrates what has 
already been said above relative to the apparent progressive differ- 
entiation of the species. 

The two species which form the extremes of the whole series are the 
most distinct, not only from each other, but from the other members 
of the same dichotic stem of the ancestral trunk. 

Muricatum las the broadest foliaceous margin; the basal angles 
of the pronotal margin project 
backward over the humeral an- 
gles of the elytra; the anterior 
tibiz are not carinate externally. 

Depressum has the narrowest 
margin; elytral base more dis- 

ae tant from the prothorax, and 
3rd. Segment Horizontal “3rd Segment Obique  (uite densely sculptured. 

In each hemi-series the mar- 
ginal line decreases —that is, 
strongest in muricatum, weakest 
at apex in contractum, strongest 
in depressum, weakest at apex in glabrum. 

It can be assumed that the acquisition of the obliquity of the third 
segment might determine the lines of divergence from the ancestral 
trunk. 

A careful consideration of the diagram in connection with the 
characters enumerated under the specific headings should further 
elucidate the interesting relationships. 

Distribution—Thus far I have studied material from the follow- 
ing States and Territories: California, Arizona, Nevada, New Mex- 
ico, Texas, Colorado, Kansas, Nebraska, Wyoming, and South 
Dakota. 

It will be observed that Hmbaphion has a large area of distribu- 
tion—approximately two-thirds of the region west of the Mississippi 
River, not entering the region west of the Sierra Nevada Mountains, 
nor the extreme northwestern and northern States. 

Depressum from eastern central California. 

Elongatum from eastern northern California and western Nevada, 


lic. 8.—GENEALOGICAL DIAGRAM OF THE 
GENUS EMBALPHION. 


REVISION OF ELEODITNI—BLAISDELL. 453 


Glabrum from Arizona and New Mexico. 

Contractum from New Mexico. 

Planum from Kansas and Wyoming. 

Contusum from Arizona, New Mexico, Colorado, Kansas, and 
Wyoming. 

Laminatum from Texas. 

Muricatum from Texas, Colorado, Kansas, Nebraska, and South 
Dakota. 

The data here given are suggestive and I shall refrain from making 
any further comments upon specific distribution until more collecting 
shall have been done, and it is to be hoped that future collectors will 
be careful in giving full accounts of locality and elevation. 


ANALYTICAL KEY TO SPECIES OF GENUS EMBAPHION. 


Hind angles of prothorax subacute and overlapping the humeral angles of the 
Pinay PrOnGly Mareined_. = a muricatum. 
Hind angles obtuse not overlapping the humeral angles; body acutely but not 
broadly margined. 
Thorax sparsely and not coarsely punctured, distinctly margined; not dis- 
tant from the elytra; legs black. 
Elytral margin attaining apex. 
Broadly oval; males distinctly caudate____._.__________- contusum. 
Elongate oval; elytra gradually narrowed__---------~- elongatum. 
Elytral margin not attaining the apex. 
Thorax with a moderate foliaceous margin; elytral dis¢ more or 


less concave, smooth and obsoletely sculptured___--~~~- glabrum. 
Thorax with margin acute, not foliaceous; elytral disc flat or 
feebly convex. |. sfy 
Acute margin ‘not attaining the apex _-__----- contractum. 
Acute mareir “attaining ‘the: apex... 2-4 e planum. 
Thorax very coarsely punctured, slightly distant from elytra, the latter 
broadly oval, subrugose; legs ferruginous brown _--~--~-~-~--- depressum., 


EMBAPHION DEPRESSUM LeConte. 


Bleodes depressa LeConte, Ain, Lyc. Nat. Hist. N. Y., V. 1851, p. 136. 
Embaphion depressum LeConte, Smith, Miscell. Collections, No. 140, 1866, 
p. 60,—Horn, Trans. Amer. Phil. Soc, Phila., XIV, 1870, p. 322. 

Rather robust, ovate, black, thin margin of the body almost totally 
wanting; thorax and elytra slightly distant. 

Antenne moderately long and somewhat stout, basal joints moder- 
ately pubescent, last three joints slightly thickened. 

Prothorax about one-third wider than long, slightly narrowed be- 
hind; dise moderately convex, very coarsely punctate; aper rather 
feebly emarginate; sides moderately rounded, with margin feebly re- 
flexed ; base with the middle slightly prominent; basal angles obtuse. 

Elytra broadly oval, acutely margined, the marginal line reaching 
the apex; base emarginate; humeral angles prominent ; sides strongly 


454 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


rounded, gradually narrowing to apex; disc scargely convex, acutely 
inflexed at the sides, sculptured with rows of moderately coarse 
punctures, bearing short hairs, and more or less distinetly rugose. 

Epipleure with superior marginal line distinctly elevated. 

Abdomen with the third segment placed more or less obliquely to 
the second. Legs ferruginous brown. 

Male—Third abdominal segment distinctly obliquely placed to 
the second. Anterior tibize suddenly and briefly sinuate at base. 

Female——Third abdominal segment slightly oblique to the second. 
Anterior tibiz gradually narrowing at base. 

Measurements.—Length, .88—44 inch. 

Habitat—California (Vallecito, LeConte; Maricopa Desert, 
Horn). 

Unknown to me in nature. Extremes of this species should always 
be carefully compared with those of /. planipennis. 


EMBAPHION ELONGATUM Horn. 
Embaphion elongatum Horn, Trans. Amer. Phil. Soc. Phila., XTV, 1870, p. 
321. 

Elongate, oblong-oval, nearly three times longer than wide, mar- 
gins of body not foliaceous, surface dull, acute elytral margin reach- 
ing to the apex; usually finely sculptured. 

Head small, scarcely twice as wide as long, quite flat, impressed 
laterally within the supra-antennal convexities, the latter quite prom- 
inent, impression extending more or less feebly along the frontal 
suture; surface finely, sparsely, irregularly and muricately punctured, 
obsoletely so at times. Antenne long, reaching beyond the pro- 
thoracic base, outer four joints feebly compressed, scarcely widened, 
third joint equal to the next two combined, fourth to the seventh, 
inclusive, subequal; eighth slightly compressed and subtriangular; 
ninth circular, and at times slightly widened; tenth suborbicular; 
eleventh oval to ovate. 

Pronotum about one-fourth wider than long, evidently narrowed 
behind; margin acute, moderately explanate, more or less reflexed ; 
disc feebly to moderately convex, finely and sparsely asperately 
punctate, granular laterally on the margins; apea more or less 
moderately emarginate—the emargination less than one-third to one- 
fourth wider than deep, obsoletely margined; sides evenly and 
moderately arcuate from apex to base; base proper truncate to feebly 
arcuate and about equal to the apex proper, obsoletely margined ; 
apical angles narrowly rounded, subacute, attaining the eyes; basal 
angles obtusely rounded and projecting slightly beyond the middle 
of the base. 

Propleure more or less concave beneath the margin at point of 
reflexion, surface smooth, opaque and very sparsely, finely punctulate. 


—— aay. 


REVISION OF ELEODIINI—BLAISDELL. 455 


Elytra elongate oval, fully twice as long as wide; base subtruncate, 
feebly sinuate laterally ; humeral angles more or less distinet ; margin 
acute, not foliaceous nor suddenly formed, evenly and gradually 
reflexed and attaining the apex; sides feebly arcuate, gradually nar- 
rowing to apex and attenuately converging, apex acute and not pro- 
duced ; dise more or less concave, suture frequently raised posteriorly : 
here the dise is more or less biconcave, gradually, evenly arcuately 
declivous behind, acutely inflexed laterally, the inflexed sides moder- 
ately oblique and plane; surface sculptured with fine, irregularly, 
sparsely, and evenly placed, subasperate punctures, the inflexed sides 
similarly sculptured; each puncture bears a very minute decurved 
seta. : 

E pipleure narrow, scarcely at all differentiated from the inflexed 
sides of the elytra and similarly punctured; distinctly defined at apex 
and feebly, minutely dilated beneath the elytral apices. 

Sterna opaque and finely, irregularly sculptured. 

Parapleure opaque and obsoletely punctate. 

Abdomen opaque, finely, rather evenly and not densely punctulate ; 
first two segments horizontal, third slightly oblique, last two much 
less convex. 

Legs long and rather slender; posterior femora usually as long as 
the first four abdominal segments; profemora mutic; tibial spurs 
quite similar and the protarsi simple. 

Male——FElongate and rather narrow. Abdomen quite strongly 
convex, median surface of the first three segments more or less broadly 
and feebly impressed. Anterior tibiz suddenly and briefly con- 
stricted at base. 

Measurements.—M ales: Length, 15-20 mm.; width, 6-8 mm. 

Genital characters, male.—Edeagophore elongately subfusiform, six 
to eight times longer than wide, more or less arched. 

Basale elongate, about five times longer than wide; sides sub- 
parallel to slightly arcuate; surface transversely and moderately 
convex, distinctly membranous centrally at the distal extremity. 

Apicale rather slender, elongately triangular ; dorsal surface moder- 
ately convex, with a median linear groove at middle two-thirds, very 
sparsely and finely punctulate: sides briefly arcuate at base and 
middle, with intervening edge and side of apex feebly sinuate; apex 
subacute and slightly produced; base distinctly bisinuate with a 
median subacute lobe. 

Sternite subparabolic in outline, circularly concave at middle like 
a watch-glass between the bases of the lobes. Each lobe triangular 
and distinctly longer than wide; external border evenly arcuate, 
apex subobtusely rounded, and the internal border straight or feebly 
sinuate; surface more or less convex, concave internally to sinus, 
basal third smooth and almost impunctate, apical two-thirds densely 


456 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


punctured and clothed with quite long sete. Sinus triangular, two- 
thirds closed by membrane, the latter setose. 

Habitat—Nevada (Horn; Hubbard and Schwarz); California 
(Lassen County, Charles Fuchs). 

Number of specimens studied, 2 (males). 

Type in the Horn collection; collected by Mr. William M. Gabb. 

Type-locality—_Western Nevada. 

Salient type-characters.—The most elongate of our species. Pro- 
thoracic margin acute, moderately explanate and reflexed. Elytra 
with margin simply acute and extending to apex. Epipleuree indis- 
tinctly defined except at apex. 

Diagnostic characters.—The elongate form, with elytra gradually 
and arcuately narrowing to apex give a characteristic facies. 

Separated from contusum by its narrower form, besides the 
body has a greater dorso-ventral thickness, and the males are not 
‘audate. In planuwm the elytral margins do not reach the apex, in 
clongatum they do. From depressum by its elongate oval elytra, 
obsoletely defined epipleuree and by having the basal joints of the 
antenne less pubescent. 

General observations —The mentum is rather small, middle lobe 
variable, sides converging anteriorly, straight or more or less arcuate, 
with apex rounded or subacute; surface very slightly convex and more 
or less feebiy foveate laterally. 

The prosternum is arcuately prominent ventrally with the coxe, 
feebly compressed and sometimes subvertically truncate posteriorly. 

Mesosternum arcuately declivous and rather deeply concave. 

The abdominal intercoxal salient is small and quite quadrate, one- 
half of its width shorter than the post-coxal portion of the same seg- 
ment, the latter subequal in length to the second; third segment a 
little shorter than the second; the fourth is one-half the length of the 
second. 

The abdominal and metacoxal salients are quite equal in width. 

The metasternum laterally between the coxe is short and equal in 
length to the width of a mesotibia at middle. 

The tibial grooves of the femora are well developed, more or less 
concave externally, becoming flat internally, glabrous and impune- 
tate; margins distinct, more or less acute and gradually converging 
to the femoral base. 

The protibie are distinctly compressed, and carinate externally; 
‘arina quite entire, surface longitudinally grooved behind the same, 
groove asperately sculptured. 

Meso- and metatibize subcylindrical in section, flattened or grooved 
externally, grooves opaque, glabrous and more or less punctate. 

The tarsi are moderate in length and stoutness. 


REVISION OF ELEODITNI—BLAISDELL. 457 


Protarsi about one-fifth of their length shorter than a mesotarsus. 
First four joints short, subequal in size and length, the first not 
noticeably longer, each just a little wider than long; the fifth equal 
to about the preceding three in length. 

Mesotarsi about one-fourth of their length shorter than a meta- 
tarsus. The first joint just the least shorter than the next two com- 
bined; joints two to four inclusive about equal in length and just a 
little longer than wide; the fifth about equal to the preceding two 
combined. 

The metatarsi are slightly more than one-half as long as a meta- 
tibia. The first joint is just a little shorter than the next two com- 
bined; the second and third subequal and distinctly longer than wide, 
both together equal to the fourth. 


EMBAPHION GLABRUM, new species. 


Suboblong oval, twice as long as wide, nigro-piceous, surface more 
or less shining, quite smooth, obsoletely sculptured, margins of the 
pronotum moderately reflexed. 

Head about twice as wide as long, plane or feebly convex, slightly 
impressed laterally, sometimes along the frontal suture, sides not 
strongly prominent, obsoletely punctulate. Antenna rather slender 
and equal in length in the sexes, reaching slightly beyond the pro- 
thoracic base, very slightly compressed in outer four joints, which are 
also feebly dilated, third joint equal to the‘next two taken together, 
fourth scarcely longer than the fifth, sixth and seventh very slightly 
shorter and subequal, eighth subtriangular, ninth and tenth sub- 
orbicular, eleventh ovate. 

Pronotum proper slightly wider than long and about as wide as 
one elytron, margins thin and moderately foliaceous, reflexed and 
concave; disc moderately convex, smooth and obsoletely punctulate, 
laterally obsoletely granulate and rugulose; apex moderately deeply 
emarginate, the emargination about four times wider than deep, 
obsoletely margined; sides evenly rounded from base to apex, not 
noticeably widening at base; base truncate and about equal to the 
apex proper, finely margined; apical angles moderately rounded, 
reaching to the eyes; basal angles rounded and slightly more promi- 
nent posteriorly than the base. 

Propleure smooth and impunctate. 

Elytra oval, width equal to about two-thirds of the length, margins 
acute, slightly reflexed and not quite attaining the apex; base sub- 
truncate; humeri proper obtuse and rounded, not anteriorly promi- 
nent; sides quite evenly but not strongly arcuate, apex simply sub- 
acute; disc smooth, more or less transversely concave or feebly bi- 
concave, sometimes plane between the reflexed margins, frequently 


458 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


slightly and briefly arcuately declivous at base, and arcuately de- 
clivous posteriorly as usual; inflewed sides not strongly oblique, 
quite plane at the middle, feebly concave beneath the humeri and 
distinctly so where it passes into the apex; surface smooth and obso- 
letely punctate, inflexed sides smooth and impunctate. 

Epipleure smooth, impunctate and obsoletely defined, except at 
apex; surface on the same plane as the inflexed sides of elytra. 

Sterna and parapleure smooth and more or less obsoletely seulp- 
tured. 

Abdomen obsoletely sculptured and smooth; third segment more 
or less oblique to the fourth and fitth segments which have a feeble 
transverse convexity. 

Legs moderate in length and thickness. Anterior femora mutic, 
tibial spurs quite similar and the tarsi are alike in the sexes, and 
simple. 

Male-—Elytral dise rather obliquely arcuately declivous pos- 
teriorly, apex acute and just noticeably produced. Abdomen moder- 
ately convex and broadly impressed on first two segments. Protibiee 
briefly and moderately constricted at base. 

Female.—Elytral dise arcuately and more or less vertically decliv- 
ous behind, apex simply subacute, abdomen rather strongly convex. 
Protibiz gradually and feebly narrowed at base. 

Measurements—Male: Length, 17 mm.; width, 8.2mm. Females: 
Length, 15-17 mm.; width, 7.5-8.5 mm. 

Genital characters, Male-—Edeagophore elongate subfusiform, six 
to eight times longer than wide and more or less arched. 

PBasale elongate and about five times longer than wide, sides more 
or less feebly arcuate, surface moderately convex. 

Apicale elongately triangular; dorsal surface moderately convex, 
without median groove (in the few specimens examined), impunctate ; 
sides broadly-and feebly sinuate; apex subacute and rather slender; 
base bisinuate with a median triangular lobe. 

Sternite as in elongatum. 

Female.—Genital segment triangular in outline, rather elongate, 
surface quite plane and not setose. 

Valvula (Plate 5, fig. 14).—Dorsal plate oblong-triangular; surface 
very slightly concave, smooth, obsoletely and sparsely punctate; ex- 
ternal border more or less sinuate; internal border very feebly areu- 
ate; apical margin short and more or less feebly rounded or truneate, 
directly continuous with the dorsal surface of the apex in the internal 
third, angle rounded, not setose. Apex chitinous, produced and 
arcuately everted, convex above and concave beneath, tip rounded, 
excavated externally at base for the appendage, the latter rather 
short mammilliform, with a few moderately long sete at tip. Basal 
prominences obsolete. 


REVISION OF ELEODITNI-—-BLAISDELL. 459 


Superior pudendal membrane attaining the middle of the dorsal 
plate, finely and longitudinally rugulose. 

Ventrolateral surfaces.—Body somewhat depressed, surface lines 
straight when viewed longitudinally, moderately convex transversely, 
smooth and shining; submarginal groove distinct beneath the not ex- 
planate external margin of the dorsal plate. Internal margins of the 
valves contiguous in basal four-fifths. Fissure very narrowly fusi- 
form in apical fifth, membrane not visible. 

Habitat—Arizona (Winslow, Barber and Schwarz; Holbrook, 
H. F. Wickhiam) ; New Mexico (Coolidge, Wickham). 

Number of specimens studied, 7. 

Sexitypes (Cat. No. 12235) in the U. 8. National Museum collection. 

Type-locality.—W inslow, Arizona. 

Salient type-characters.—Surface smooth, more or less shining and 
obsoletely sculptured. Pronotum with the lateral margins thin and 
moderately foliaceous, reflexed and concave. Elytral margins acute, 
slightly reflexed and not quite attaining the apex; dise more or less 
slightly concave. 

Diagnostic characters.—Conspicuous on account of its smooth in- 
teguments and entire absence of the short sete that are so noticeable 
in all of the other species. The body has a much greater dorso- 
ventral thickness than any of the species before me. The sexes are 
also less differentiated than in any other. It differs from contusum, 
elongatum, and depressum in not having the marginal lines of the 
elytra attain the apex. From planum by its moderately foliaceous 
prothoracic margins, more or less concave elytral dise and in the 
smooth and obsolete sculpturing. 

Glabrum appears more robust than any other form on account of 
the unusual dorso-ventral thickness. 

General observations—Mentum with the middle lobe slightly 
smaller than usual, subtruncate anteriorly and feebly straightened 
laterally, the sides oblique and converging to apex, the outline is 
more semicircular than otherwise; surface smooth and somewhat 
punctate laterally. 

The prosternum is quite prominent ventrally with the cox, 
arcuate antero-posteriorly and rather vertical posteriorly, not in the 
least mucronate. 

Mesosternum arcuate and more or less vertically declivous, longi- 
tudinally concave. 

The abdominal process is slightly transverse, in width scarcely as 
long as the third abdominal segment; the post-coxal portion of the 
first about equal to the’ second, the latter is twice as long as the 
fourth: the third is about one-half of its length shorter than the 
second segment. 

The abdominal and metasternal salients are equal in width. 


460 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


The metasternum laterally between the coxe Js short, in length 
about equal to the width of a mesotibia at middle. 

The tibial grooves of the femora are entire, glabrous, flat to more 
or less concave, their margins moderate to feeble and acute, converg- 
ing to the femoral base in each, where they are scarcely contiguous. 

The external surfaces of the tibie are as in planwm. 

The tarsi moderate as usual. 

The protarsi are about one-fourth of their length shorter than a 
mesotarsus. First four joints small and subequal, the first distinctly 
longer than wide, second, third, and fourth as wide as long: the fifth 
about equal in length to the preceding three combined. 

The mesotarsi are about one-fifth of their length shorter than a 
metatarsus. The first joint is about one and a half times longer than 
the second; joints two to four inclusive are subequal in length and a 
little longer than wide; the fifth about equal in length to the pre- 
ceding three taken together. 

The metatarsi are about two-thirds as long as their respective meta- 
tibia. The first joint is about equal to the second and third combined ; 
the fourth equal to the first; the second and third are subequal in 
length and about one and a half times longer than wide. 


EMBAPHION CONTRACTUM, new species. 


Oblong-oval, piceous black, feebly shining to subopaque, sides of 
the body not foliaceous, acute elytral margin not attaining the apex; 
pronotum constricted in basal eighth; each puncture with a very 
short, semierect seta. 

/Tead moderately large, more or less plane to feebly convex between 
the eyes, rather broadly and distinctly impressed within the supra- 
antennal convexities, frontal suture more or less feebly evident; 
surface sparsely, irregularly, and somewhat finely punctate, punctures 
slightly denser at the periphery. 

Antennw long and moderately slender, outer four joints feebly 
compressed and searcely dilated; third joint equal to the next two 
taken together; fourth scarcely longer than the fifth; the latter to 
the seventh, inclusive, subequal in length: eighth subtriangular and 
longer than wide; ninth and tenth very feebly transversely oval; 
eleventh pointed ovate and scarcely longer than wide. 

Pronotum with the total width about one-half greater than the 
length, margin acute and not foliaceous; disc proper about as wide 
as long, moderately, broadly, and evenly convex, sparsely, irregularly, 
and not strongly punctate, with a more or less feeble transverse anti- 
basal impression, laterally rather broadly impressed or feebly con- 
cave within the slightly reflexed acute margins, surface here more 
opaque and obsoletely sculptured; apew broadly, evenly, and moder- 


REVISION OF ELEODIINI—BLAISDELL. 461 


ately emarginate, evidently not beaded; s/des evenly and moderately 
arcuate to the basal seventh or eighth, there quite abruptly con- 
stricted with sides parallel or slightly convergent to ase, the latter 
slightly arcuate and rather coarsely but not strongly beaded, equal 
in width to the apex; apical angles very narrowly rounded; basal 
angles subrectangular. 

Propleure smooth, opaque, and obsoletely sculptured. 

Elytra oblong-oval, widest at the middle, a third of the length 
longer than wide; Jase feebly emarginate between the rather broadly 
rounded and slightly more anteriorly prominent Awmeri, sides evenly 
and moderately arcuate, more strongly convergent in apical fourth 
where they are feebly and broadly sinuate, the apex appearing some- 
what produced and subparabolically rounded; disc flat to feebly con- 
vex, limited laterally from the obliquely inflexed sides by an acute 
margin, that does not attain the apex, arcuately declivous pos- 
teriorly in apical third; surface irregularly and not densely subas- 
perately punctate, at times there is a distinct tendency to a serial 
arrangement, the punctures are slightly denser and a little more 
muricate at the periphery, the acute margin is subserrulate; inflexed 
sides more or less obsoletely punctate. 

E pipleure narrow and not dilated at the base, gradually narrow- 
ing to apex; surface smooth and impunctate; superior margin not 
defined in basal half, thence to apex well developed. 

Sterna quite smooth and more or less obsoletely sculptured. 

Parapleurw rather coarsely and more or less obscurely punctate. 

Abdomen smooth, very sparsely and finely punctulate, more or less 
obsoletely rugulose; punctules more distinct and denser on the fifth 
segment. 

Legs slender and moderate in length, femora mutic, anterior tibial 
spurs subequal and the tarsi similar in the sexes. 

Male—Unknown to me, the secondary sexual characters are prob- 
ably as in planum. 

Female—Moderately broad. Abdomen not very strongly but 
evenly convex. Anterior tibiz quite gradually and slightly sinuate at 
base. 

Measurements—Males: Not studied. Females: Length, 18-21.5 
mm; width, 9-10 mm. 

Genital characters, male—Not studied. 

Female.—Genital segment elongate and triangular in outline, dor- 
sal surface more or less transversely ogival in section, impunctate and 
not setose. 

-Valvula (Plate 5, fig. 20).—Dorsal plate oblong-triangular, about 
four times longer than wide; surface feebly convex basally and 
concave apically: external border more or less sinuous; internal bor- 
der nearly straight and somewhat beaded; apical margin arcuate in 


462 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


external two-thirds, becoming continuous internally with the inner 
margin of the fossa. Apex produced and not everted, sides sub- 
parallel, externally continuing the line of the dorsal plate, tip rather 
broadly rounded, dorsal surface concave; internally the valvular 
membrane is finely setose and semi-chitinous caudad to the pudendal 
membrane. 

Appendage short and nearly hemispherical, projecting very little 
beyond the fossa, setose at tip, sete long forming a loose pencil. 
Fossa rather broad and not sharply defined. 

Superior pudendal membrane reaching to the apical fourth of the 
dorsal plate and longitudinally rugulose. 

Basal prominences scarcely at all prominent. 

Ventrolateral surfaces transversely convex, surface lines straight 
when viewed longitudinally, concave apically before the apices. Sub- 
marginal groove distinct beneath the feebly explanate external margin 
of the dorsal plate. Internal margins of the valves contiguous in 
basal two-thirds; genital fissure very narrow in apical third. Inferior 
membrane not visible. 

Habitat—New Mexico (Cloudcroft, elevation 9,000 feet). 

Number of specimens studied, 7 (females). 

Types in my own collection. 

Ty pe-locality —Cloudcroft, New Mexico; collector, Warren Knaus. 

Diagnostic characters.—The seven specimens studied are homomor- 
phic and distinct from their congeners. The shortening of the ex- 
panded pronotal margin is quite remarkable, and the basal constric- 
tion causes the prothorax to appear more distant from the elytra than 
in depressum. | 

In fact, the form of the prothorax is an approach to that observed 
in ELleodes planipennis. Contractum appears as a step toward the 
bridging over of the gap between Lleodes planipennis and Em- 
hbaphion. It is also significant that both should inhabit the same 
geographical region. Some specimens of planipennis exhibit a ten- 
dency to a thinning and reflexion of the pronotal margins, but never 
to an acuteness of the elytral margins. When such examples are 
found planipennis must be referred to Lmbaphion. 

In depressum the acute elytral margin attains the apex, in contrac- 
tum it does not. 

In planum the acute and expanded pronotal margins reach to and 
slightly beyond the base, while in contractum they are abbreviated 
posteriorly and not extending beyond the basal seventh or eighth, the 
true basal angles being exposed. 

Contractum is without doubt an extreme and perpetuated modifi- 
cation of planum. In the series of contractum examined the abdo- 
minal segments are not oblique. 


. REVISION OF ELEODILINI—BLAISDELL. 463 


Tam indebted to Mr. Warren Knaus for this interesting addition to 
our list. 

General observations.—The mentum is rather small, feebly trans- 
verse, more or less arcuate anteriorly, although at times apparently 
subtruncate, sometimes feebly impressed at apex so as to appear some- 
what emarginate; surface feebly convex, glabrous, and obsoletely 
punctate at center, narrowly impressed peripherally within the mar- 
gin, there opaque and distinctly punctate. 

Prosternum less suddenly protuberant ventrally than in planwm, 
otherwise the same. 

The mesosternum is quite vertical and arcuately declivous, more or 
less concave as usual. 

The metasternum laterally between the coxw is as long as the width 
of a mesotibia at apex. 

The metasternal and abdominal salients are quite equal in width. 
The abdominal process is a little longer than wide. The post-coxal 
portion of the first abdominal segment is about equal to the third in 
length; the second is about twice as long as the fourth segment. 

The tibial grooves of the femora are well developed, more or less 
concave, but not strongly so. The margins are subcariniform exter- 
nally, becoming more or less evanescent internally. On the profemora 
the grooves have the margins slightly arcuate and about attaining 
the femoral base; those of the meso- and metafemora have the mar- 
gins quite straight, scarcely attaining the base on the former and 
evanescent at basal fifth on the latter. 

The protibie are feebly compressed and distinctly carinate ex- 
ternally; tarsal grooves more or less obsolete, articular cavities 
closed. On the meso- and metatibiew the grooves are more or less de- 
veloped and narrow on apical two-thirds, slightly glabrous, margins 
asperate, and the articular cavities are closed. 

The tarsi are the same as in planum. 

The protarsi are about two to three-fifths of their length shorter 
than a mesotarsus, joints two to four subequal in length and scarcely 
as long as wide; the first joint is a little longer than wide: the fifth 
is just the least longer than the three preceding joints taken together. 

The mesotarsi are about two-fifths of their length shorter than a 
metatarsus. Joints two to four, inclusive, are subequal in length and 
slightly longer than wide; the first joint is about one-half longer than 
wide; the fifth is slightly shorter than the preceding three joints taken 
together. 

The metatarsi are distinctly more than a half longer than their 
respective tibia. Joints two and three are subequal in length and 
distinctly more than one-half longer than wide, and together as long 
as the fourth; the first is scarcely as long as the next two taken 
together. 


464 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


EMBAPHION PLANUM Horn. 
. 


Embaphion planum Horn, Trans. Amer. Phil. Soc. Phila., XIV, 1870, p. 321. 

Oblong-oval, about twice as long as wide; margins of body not 

foliaceous, acute elytral margins not attaining the apex; surface more 
or less dull. 

Head moderate, about twice as wide as long, quite plane between 
the eyes, feebly impressed laterally ; sides of the frons slightly promi- 
nent, finely, rather sparsely punctate, each puncture with a minute 
curved seta, frontal sttture at times distinct, sometimes slightly 
impressed. Antenne quite long; outer four joints scarcely com- 
pressed or widened; third joint about equal to the next two taken 
together; fourth to the seventh, inclusive, subequal; eighth triangu- 
lar; ninth and tenth suborbicular; eleventh ovate. 

Pronotum one-sixth to one-half wider than long and slightly nar- 
rowed behind; margin acute, not foliaceous nor suddenly formed; 
disc feebly convex, finely and rather sparsely punctate, punctures 
becoming slightly denser laterally, each with a small curved seta; 
apex evenly but not deeply emarginate, the emargination four or five 
times wider than deep, usually very finely margined; sides evenly 
and moderately arcuate from base to apex; margin slightly explanate, 
widening a little posteriorly, feebly reflexed; ase distinctly shorter 
than apex, transverse, and obsoletely margined ; apical angles obtusely 
rounded, attaining the eyes; basal angles obtuse, rounded, and pro- 
jecting slightly beyond the middle of the base. 

Propleure smooth, impunctate, and obsoletely rugulose, defined 
from the inferior surface of the pronotal margin by a shallow groove. 

Elytra elongate oval, base at middle slightly prolonged and feebly 
sinuate laterally ; humeral angles distinct and rounded ; margi acute, 
feebly reflexed and not attaining the apex; sides evenly arcuate, grad- 
ually converging apically; apex acute and not caudate; disc flattened, 
at times feebly convex, never concave, evenly and rather gradually 
arcuately declivous posteriorly ; acutely inflexed laterally, the inflexed 
sides feebly convex; surface sculptured with subasperate punctures, 
that may be serially arranged or irregular and somewhat denser 
laterally ; the inflexed sides are irregularly and rather sparsely punc- 
tate. Each puncture bears a short curved seta. 

Epipleure narrow and not dilated at base, gradually narrowing to 
apex: superior margin distinctly defined; surface glabrous and im- 
punctate. 

Sterna more or less feebly punctate and rugulose, feebly shining. 

Parapleure quite glabrous, feebly and obsoletely punctate. 

Abdomen horizontal, more or less shining, sparsely or not very 
densely punctulate, obsoletely rugulose. 

Legs moderate and rather slender, anterior femora mutic, tibial 
spurs similar and the protarsi simple in the sexes. 


REVISION OF ELEODIINI—BLAISDELL. 465 


Male—Somewhat narrow. Antenne reaching considerably be- 
yond the prothoracic base. Abdomen moderately convex, first three 
segments rather broadly impressed along the middle. Protibize 
suddenly and briefly sinuate at base. 

Female——More robust. Antenne reaching slightly beyond base 
of the prothorax. Abdomen rather strongly convex.  Protibixe 
gradually narrowing at base. 

Measurements.—M ales: Length, 14-19.5 mm.; width, 6-9 mm. Fe- 
males: Length, 13.2-17 mm.; width, 7-8.5 mm. 

Genital characters, male—Edeagophore very slender, elongate, and 
moderately arched. 

Basale about five times longer than wide, rather strongly convex 
from side to side, the latter feebly arcuate. 

Apicale small, triangular, slightly elongate ; surface rather strongly 
convex and without groove; sides feebly and broadly sinuate; apex 
gradually attenuated and acute; base with a rather short, subacute 
lobe at middle and feebly sinuate laterally. 

Sternite transversely triangulo-parabolic in outline. Each lobe 
triangular, with external border more or less arcuate to apex, the 
latter subacute and rather gradually narrowed; internal margin 
nearly straight and oblique; surface more strongly convex along the 
external border, thence sloping inward, rather densely punctate in 
apical two-thirds, setose, setee long and rather dense, especially in 
apical third. Sinus more or less semicircular, membrane more or 
less setose across the same. 

Female—Genital segment (Plate 5, fig. 9) triangular, surface quite 
plane, impunctate and not setose. 

Valvula—Dorsal plate elongately subtriangular, moderately nar- 
rowed from base to apex; surface plane and glabrous; external 
border more or less feebly sinuate; apical margin truncate at middle, 
internally continuous with surface of the apex, angle evenly rounded; 
internal margin very feebly arcuate. Apex produced and chitinous, 
more or less everted, convex above and concave beneath, tip rounded. 
Fossa at base of the external surface of apex and beneath apical mar- 
gin of dorsal plate. 

Appendage short mammilliform, with a pencil of four or five 
rather long sete at tip. 

Basal prominences moderately developed. 

Superior pudendal membrane attaining the apical fourth of the 
dorsal plate, and longitudinally rugulose. 

Ventrolateral surfaces—Body slightly protuberant. laterally in 
basal half, feebly concave laterally before the apices; surface lines 
straight. Submarginal groove shallow beneath the feebly expanded 
external border of the dorsal plate. Internal margins of the valves 
contiguous in basal three-fourths; genital fissure in apical fourth 

59780—Bull. 683—09——30 


466 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


and nearly closed, margins fringed with very small sete. Inferior 
membrane not visible. . 

Habitat—Kansas (George Horn); Colorado (Denver, April, H. 
Soltau); Wyoming (Cheyenne, June and July, Hubbard and 
Schwarz). 

Number of specimens studied, 34. 

Type in the Horn collection. 

Ty pe-locality.— Kansas. 

Salient type-characters.—Thoracic margin acute, not foliaceous nor 
suddenly formed, not deeply emarginate at apex. Elytral margin 
acute, feebly reflexed, and not attaining the apex; disc flattened, and 
never concave. Epipleural limits distinctly defined (Horn). 

Diagnostic characters—The thorax is distinctly wider than long, 
the apical emargination is shallower and broader, the elytral dise is 
flat or feebly convex, and the apices never caudate; the margin does 
not reach the apex. The epipleure are usually well defined and 
even feebly convex. 

From contusum it differs in the shallower apical emargination of 
the prothorax, and by the elytral margin not reaching the apex. The 
prothoracic margin is distinctly narrower. 

It is recognized from elongatum by the shorter and broader form 
and by the elytral margin not attaining the apex. 

From depressum by the wider thoracic and elytral margins, and in 
not having the marginal line reaching the apex. 

Planum is undoubtedly very closely related to contusum and a 
very large series might show it to be simply a subspecies. I deem 
it best at present not to unite the two until further collecting has 
been done. The explanate thoracic’ margin varies in width and the 
marginal line is often indicated to apex by a line of asperities. 

General observations.—The mentum is moderate, middle lobe more 
or less semicircularly arcuate anteriorly from the subangulate sides. 
The apex may be subtruncate and the sides straight and oblique; 
surface feebly convex and more or less obsoletely foveate laterally. 

Prosternum and mesosternum as in contusum. In a few specimens 
there is a very slight tendency for the prosternum to become feebly 
prominent posteriorly. 

The abdominal process is rather small and nearly quadrate, its 
width is slightly less than the length of the post-coxal portion of 
the same segment, the latter is about equal to the third in length; 
the second is one-fifth of its length longer than the third, the latter 
scarcely twice as long as the fourth. 

The abdominal and metasternal salients are equal in width. - 

The metasternum laterally between the coxe is short, in length 
about equal to the width of a mesotibia at middle, 


REVISION OF ELEODIINI—BLAISDELL. 467 


The tibial grooves of the femora are well developed and defined ; 
surface of each plane to concave, glabrous and at times feebly rugu- 
lose; margins well defined and more or less acute, gradually con- 
verging to the femoral base in each instance, those of the profemora 
are slightly arcuate. 

The protibixw are slightly compressed and distinetly carinate ex- 
ternally, carina more or less entire, surface behind it rather narrowly 
and longitudinally grooved, groove more or less asperate. 

Tarsal grooves of the mesotibixw rather more or less evident ex- 
ternally in apical two-thirds, grooves more or less narrow, sub- 
glabrous or subasperate. External surfaces of the metatibie are 
more or less flattened, scarcely grooved and asperate. 

The tarsi are moderate in length and stoutness. 

The protarsi are about one-fourth of their length shorter than a 
mesotarsus. The first four joints are subequal in length, the first is 
slightly longer than wide, the succeeding three feebly wider than 
long. 

The mesotarsi are about one-fifth of their length shorter than a 
metatarsus. First joint scarcely as long as the next two combined: 
the second, third, and fourth subequal in length and distinctly 
longer than wide; fifth a little shorter than the preceding three 
taken together. 

The metatarsi about three-fifths as long as their tibie. First 
joint quite equal to the succeeding two taken together, the fourth 
just a little longer than the second and third combined, the latter 
two are equal to each other. 


EMBAPHION CONTUSUM LeConte. 


Embaphion contusum LeConte, Journ. Acad, Nat. Sci. Phila., 2d ser., IV, 
1858, p. 20; Smithson. Contrib. to Knowledge (Coleopt. Kans. and east- 
ern N. Mex.), 1859, p. 15, pl. 1, fig. 8—Horn, Trans. Amer. Phil. Soc. 
Phila., XIV, 1870, p. 321. 

Var. laminatum Casey, Ann, N. Y. Acad. Sci., V, Nov. 1890, p. 408. 

Elongate oblong, piceous black, more or less opaque, sides of the 
prothorax rather broadly foliaceous, elytral margins less strongly 
so, margins distinctly attaining the apex. 

Head small, less than twice as wide as long, nearly plane, broadly 
and feebly impressed laterally, sides of the frons rather prominent : 
surface very finely, sparsely, and quite indistinctly punctate, each 
puncture bearing a small, short, and curved seta. 

Antenne long and rather slender, very feebly compressed in outer 
four joints, which are feebly widened; third joint a little shorter 
than the next two taken together; fourth slightly longer than the 
fifth; the latter, sixth, and seventh subequal; eighth slightly shorter 
and subtriangular; ninth and tenth suborbicular; eleventh ovate. 


468 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Pronotum proper about as wide as long, lateral margins thin and 
moderately foliaceous, more or less reflexed and @oncave; disc rather 
moderately convex, finely and rather sparsely punctate, laterally 
more densely so, each puncture with a short curved seta; apea rather 
deeply and somewhat semicircularly emarginate, the emargination 
about three times wider than deep, not noticeably margined; sides 
evenly rounded, margin not widening at base; base quite truncate, 
and about equal to the apex, with a distinct marginal bead; apical 
angles obtuse, formed by the moderately advanced foliaceous mar- 
gins, and are about half as long as the head; basal angles broadly 
rounded, slightly more prominent posteriorly than the middle of the 
base. 

Propleure smooth and quite impunctate. 

Elytra elongate oval, nearly twice as long as wide, margins acute, 
slightly reflexed and always attaining the apea; base quite truncate, 
much wider than the contiguous prothoracic base proper; humeral 
margins rather broadly rounded, not anteriorly prominent, humeri 
proper obtuse ; sédes feebly arcuate or subparallel in basal half, thence 
to apex quite evenly and broadly arcuate, apex more or less actite 
and produced ; disc quite plane or feebly concave, sometimes biconcave 
by elevation of suture in posterior half, more or less arcuately de- 
clivous posteriorly, acutely inflexed laterally, inflexed sides quite 
straight and oblique; surface sculptured with subasperate punctures 
which may be rather densely placed and more or less irregularly 
arranged, at times somewhat serially so, occasionally the. punctures 
are not asperate and rather coarse; inflexed sides irregularly and 
more or less densely punctate. 

E' pipleurw narrow, not in the least dilated at base, very gradually 
narrowed to apex, smooth and impunctate; superior margin well 
defined in apical fourth, thence to base by a series of punctures, 
surface on the same plane as the inflexed sides of the elytra; widening 
at apex to become the inferior surface of the more or less produced 
apex. 

Sterna quite smooth and more or less impunctate. 

Parapleurw vather sparsely and more or less indistinctly punctate. 

Abdomen smooth, very sparsely and more or less indistinetly pune- 
tulate, obsoletely rugulose; third segment scarcely oblique, others 
horizontal. 

Legs moderate in length, rather slender. Profemora mutic as 
usual, tibial spurs similar and protarsi simple in the sexes. 

Male——Somewhat narrow. Antenne reaching considerably beyond 
base of the prothorax. Elytral disc more or less gradually declivous 
behind, apex produced into a short cauda, which may be angulate at 
the sides where the inflexed sides terminate; cauda more or less 
abruptly produced, rather oblique, and more or less asperate, with 


re fe 


» 


REVISION OF ELEODITINI—BLAISDELL. 469 


tips separated. Abdomen moderately convex and more or less im- 
pressed on first two segments. Protibise suddenly and briefly sinuate 
at base. 

Female.—Rather broad. Antenne reaching beyond the protho- 
racic base, but shorter than in the male. Elytral (lise arcuately de- 
clivous posteriorly, not produced at apex, but more or less acute. 
Abdomen quite strongly convex. Protibiw gradually narrowing at 
base. 

Measurements.—M ales; Length, 15-20.5 mm.; width, 7.2-8 mm. 
Females: Length, 14-19.5 mm.; width, 7-10 mm. 

Genital characters, male-——Edeagophore slender, elongate oblong- 
ovate. 

Basale elongate oval, surface moderately convex, sides moderately 
arcuate, apex deeply and triangularly emarginate to receive the base 
of the apicale. 

Apicale slender and elongately triangular; surface evenly and 
rather strongly convex, without groove; sides nearly straight, gradu- 
ally converging to apex, the latter acute; base strongly and acutely 
lobed at middle, oblique and broadly sinuate laterally. 

Sternite subparabolic in outline. Each lobe subtriangular, external 
border quite evenly arcuate to apex, the latter subacute, prominent 
and slightly produced ; internal border sinuate; surface more strongly 
convex along the external border, thence sloping inward, densely 
punctured in apical two-thirds, especially internally, setose, sete 
quite long and dense, more so on apex. Sinus semicircular, membrane 
not setose across the same. 

Female.—Genital segment (Plate 5, fig. 8) elongately triangular 
with the apices, or feidingclar- -trapezoidal without the same, superior 
surface plane, not setose. 

Valvula—Dorsal plate suboblong to feebly and elongately tri- 
angular, slightly narrowing from base to apex; surface plane, smooth, 
and not noticeably punctate; external border angulate at basal fourth, 


thence more or less sinuate and arcuate to apical margin, the latter 


subtruneate at middle two-fourths, angle rounded, in inner fourth 
continuous with the surface of apex; internal margin quite straight 
or more or less feebly arcuate. Apex chitinous, produced, and 
everted, convex above and concave beneath, rounded at tip; fossa 
at base of the external apical surface, beneath apical margin of dorsal 
plate. 

Appendage mammilliform, glabrous, with a pencil of few quite 
long sete at tip. 

Basal prominences feebly evident, lateral surfaces of valves slightly 
visible from above. 

Superior pudendal membrane attaining the apical fourth of the 
dorsal plate, finely and longitudinally rugulose. 


470 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Ventrolateral surfaces—Body with surface limes straight when 
viewed longitudinally, quite evenly convex from side to side, smooth 
and shining. Submarginal groove rather small beneath the external 
border of dorsal plate, which is feebly explanate in apical half, termi- 
nating at the apical margin. Internal margins of the valves con- 
tiguous in basal four-fifths; fissure quite short, nearly closed, mar- 
gins finely setose. Inferior membrane not visible. 

Habitat—Wyoming (Cheyenne, June, Hubbard and Schwarz; 
April, H. Soltau); Colorado (Salida, Fort Collins, Clear Creek, 
South Park Region, San Luis Valley, H. F. Wickham) ; New Mexico 
(Deming, July and August, Hubbard and Schwarz; Las Vegas, 
August, Barber and Schwarz; Coolidge, Wickham) ; Arizona (George 
Horn; Winslow, Hubbard and Schwarz; Santa Rita Mountains, 
July, Hubbard and Schwarz; Tucson, Charles Fuchs) ; Kansas (F. H. 
Snow; George Horn). 

Number of specimens studied, 48. 

Type in the LeConte collection. 

Type-locality.—F ort Laramie and Santa Fé, New Mexico. 

Salient type-characters—Opaque. Thorax with deeply emargi- 
nate apex. margin broadly explanate, moderately reflexed, apical 
angles subacute, basal angles broadly obtuse. Elytra with the dorsum 
plane, acutely margined at apex, margin reflexed, strongly so, seri- 
ately, finely and rather densely muricato-punctate. Male with elytral 
apices briefly prolonged (LeConte). 

Diagnostic characters.—In this species the thorax proper is broader, 
the prothoracic margins are moderately foliaceous and reflexed, the 
integuments are more or less opaque, finely and subasperately sculp- 
tured, the elytral margins attain the apex. The males are briefly 
caudate. 

From muricatum it differs by not having the basal angles of the 
prothorax prolonged backward over the humeri. The elytral margin 
is thinner, narrower, and usually scarcely reflexed unless it be near 
the humeri. The sides of the pronotum are rounded from apex to 
base and not broader behind, the basal angles are broadly rounded 
and slightly more prominent posteriorly than the middle of the base. 

In contusum the males have the inferior surface of the elytral cauda 
form by the dilated apices of the epipleural surface. 

From glabrum it is recognized by the dull luster and finely sub- 
asperate sculpturing, also by the acute elytral margins attaining the 
apex. 

In the U. S. National Museum collection there are two males and 
two females which were collected in New Mexico. They are the 
largest and most strongly developed specimens that I have seen. The 
elytral apices are distinctly caudate, and the elytral disc quite evenly 
concave transversely in the males and the suture is feebly raised 


ef 


REVISION OF ELEODIINI—BLAISDELL. 471 


posteriorly, indicating an approach to /aminatum Casey. The 
females have the elytral dise broadly oval and quite flat. The epi- 
pleurz are not at all defined from the inflexed sides except at apex, 
they are glabrous. In these specimens the sexes have the prothoracic 
margin slightly broader and more decidedly reflexed than the average 
number of specimens of the typical form (forma typica); in fact, 
some specimens have the margins scarcely at all reflexed. 

These large specimens may be placed as an extreme form in the 
cabinet, and labeled forma grandis. 

General observations—Mentum more or less variable and moderate 
in size. Middle lobe arcuate anteriorly, or subtruncate with the 
sides straight and converging to apex; surface very feebly concave, 
rather longitudinally prominent at middle and broadly, feebly con- 
cave laterally and more or less punctate. 

The prosternum between the coxe is quite suddenly prominent 
ventrally, antero-posteriorly arcuate, frequently more or less grooved. 
Feebly widened behind the equator of the coxe. 

The mesosternum is more or less obliquely and arcuately declivous, 
more or less broadly and feebly concave. 

The abdominal salient subquadrate and scarcely as wide as the 
third segment is long; post-coxal part of first about equal in length 
to the second segment, the latter twice as long as the fourth; third 
about one-third of its length shorter than the second. 

The abdominal and metasternal processes are subequal in width. 

The metasternum laterally between the coxe is very short and 
about equal to a mesotibia at base. 

The tibial grooves of the femora are entire, well defined and gla- 
brous; their margins are subacute and smooth. The profemoral 
grooves are more or less strongly concave; the margins converge to 
become contiguous at the femoral base. 

The mesofemora have the grooves scarcely concave; the margins 
converge to become feebly distant at femoral base. 

The grooves on the metafemora are not concave, quite flat, with 
margins gradually converging and slightly distant at base. 

The protibiz are slightly compressed and carinate externally, the 
carina is entire, the tarsal groove is immediately behind it and is 
frequently obsolete, the surface always asperate. 

The mesotibiw are more or less distinetly grooved externally in 
apical two-thirds; the surface of the groove is glabrous, with sparsely 
placed asperate punctures. 

The metatibie are simply flattened externally and asperate. 

Tarsi moderate as usual. 

The protarsi are scarcely one-fourth of-their length shorter than a 
mesotarsus; joints two, three, and four subequal in size and very 
slightly wider than long and together about equal to the fifth; the 


472 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


first is a little longer than the second; mesotars# about two-sevenths 
of their length shorter than a metatarsus; joints two, three, and 
four subequal, a little longer than wide, and together just a little 
longer than the fifth; the first about one and a half times longer 
than wide. 

The metatarsi are about one-half of their length shorter than their 
metatibia; first and fourth joints equal in length and either one is 
equal to the second and third taken together; the latter two are sub- 
equal in length, each about twice as long as wide. 


EMBAPHION CONTUSUM var. LAMINATUM Casey. 


Moderately robust, black and dull throughout. 

ITead small, very finely, sparsely, and subasperately punctate. 
Antenne slender, third joint from four to nearly five times as long 
as wide, eighth one-half longer than wide. 

Pronotum about one-half wider than long; disc proper very feebly 
convex, fully as long as wide, and having two feeble, subparallel, and 
sinuous impressions near the middle; surface throughout finely, ex- 
tremely sparsely, and subasperately punctate; apex strongly emargi- 
nate, the emargination not quite three times as wide as deep; side 
margins very widely and strongly reflexed, the outer edges parallel, 
strongly and evenly arcuate throughout; base transverse; basal angles 
broadly rounded and projecting bevond the median portion of the 
base. 

Elytra from one-half to four-fifths longer than wide; base trans- 
versely truncate; humeri rounded; sides very thin and broadly but 
not abruptly reflexed, the acute lateral edges parallel and feebly 
arcuate in basal two-thirds, then rounded to apex and slightly pro- 
longed, uniting in a prolongation of the suture; disc distinctly wider 
than the prothorax, each elytron broadly concave, the suture elevated ; 
inflexed sides nearly flat, somewhat coarsely and sparsely but un- 
evenly punctate; surface of disc with approximate, imperfectly 
defined and feebly impressed rows of rather coarse, impressed punc- 
tures, also finely and sparsely asperate. 

E’pipleure very imperfectly defined except near the apex. 

Legs very slender. 

Male.-—Somewhat slender. 

Female.-—Moderately robust. 

Measurements.—Length, 14-15 mm.; width, 6.8-7.5 mm. 

Genital characters not studied. 

ITabitat—Texas (El Paso, G. W. Dunn). 

Unknown to me in nature. 

Type in Colonel Casey’s eollection. 

Type-locality.—E1 Paso, Texas. 


REVISION OF ELEODIINI—-BLAISDELL. 473 


Casey writes that this form is related only to confusum, and I can 
not help deciding that it is only one of the extreme forms of that 
species. I do not consider that the peculiar form of the upper sur- 
face of the elytra can give specific standing in this instance. It is 
true that in the typical contuswm the elytral dise is plane, but in the 
series before me are examples with flat elytral dises; others have them 
slightly concave, with margins more strongly reflexed, until finally 
the extreme forms approach the one here described under contusum 
as forma grandis, for example; the latter is much larger than /ami- 
natum, the male having a length of 20.5 mm., width 10.5 mm.; the 
female, length 19 mm., width 10.5 mm., and is distinctly a much 
larger form. 

The elevation of the elytral suture is observed in some examples 
of all of the species known to me. 

The sculpturing is quite variable and in the extremes of a series 
quite in contrast. Furthermore, the elytral punctures referred to 
in the description are merely impressed fovezw, and according to 
the original remarks do not appear to be in the least setigerous: * the 
finer, but strong and sparse, asperities are distributed over the sur- 
face without regard to the punctures, and each bears a strong, thick, 
semi-erect seta.” 

After a careful examination of the sculpturing under high power, 
I consider that the asperities represent the true punctures, whose 
margins have been produced into the minute eminences, each bearing 
a seta. 

The minute fovee or dents are not the true punctures. The sculp- 
turing is quite similar in all of the species before me; even the minute 
fovee are scattered here and there. The sculpturing of /aménatum 
is peculiar only in the apparent abundance of fovee. 

A comparative study of the elytral sete shows that in g/abrum the 
sete are the most minute and the stoutest in contusum, the longest 
and most slender in planum. 

Not having seen the types nor an authentically identified specimen, 
my views are, of course, founded upon the series at hand and the laws 
governing homology and analogy. 


EMBAPHION MURICATUM Say. 


Akis ’? muricata Say, Journ, Acad. Nat. Sci. Phila., IIT, 1824, p. 251. 

Embaphion muricatum LeConte, Complete Writings of Thomas Say, II, 
1859, p. 149; in Thomson’s Arcana Nat., I, 1859, pl. xu, fig. 10.— 
Horn, Trans. Phil. Soc. Phila., XIV, 1870, p. 320. 

Embaphion concavum LeConte, Proc. Acad. Nat. Sci. Phila., VI, 1853, p. 
446. 

Oval to oblong-oval, brownish to piceous black, thoracic and elytral 
margins very broad and foliaceous, strongly reflexed. 


474 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Head small, less than twice as wide as long, plane sides of the frons 
slightly prominent, punctate, punctures very feebly subasperate, fine, 
not dense, each with a small curved and short seta, frontal suture 
usually not visible. Antenne rather long, quite slender, outer four 
joints very slightly compressed and scarcely widened, third joint 
shorter than the next two taken together, fourth scarcely longer than 
the fifth, the latter and sixth subequal, seventh shorter, eighth feebly 
shorter than the seventh and slightly triangular, ninth and tenth 
suborbicular, eleventh subovate. 

Pronotum with margins very broadly foliaceous, the margin more 
than one-half wider than the disc, the latter comparatively narrow, 
longer than wide at middle, very feebly convex, usually with irregular 
depressed areas; finely, more or less subasperately and sparsely pune- 
tate; reflexed margins wider posteriorly and more or less concave, 
a little more distinctly punctate, punctures less sparse, each with a 
short curved seta; apex deeply and feebly subquadrately emarginate, 
the emargination about one-half wider than deep, sides almost 
parallel, and scarcely margined; sides evenly but not strongly arcu- 
ate, moderately converging from base to apex; base proper feebly 
arcuate, not margined and about equal to the length, laterally sinuate; 
apical angles rather narrowly rounded and formed by the advanced 
foliaceous margins and nearly as long as the head; basal angles are 
posteriorly prominent, subacute, and projecting backward over the 
basal angles of the elytra. 

Propleure smooth and impunctate; inferior surface of the foli- 
aceous margins obsoletely punctate. 

Elytra oval to elongate oval; margins broad and reflexed, angles 
at humeri nearly rectangular and more or less truncate at base, pos- 
teriorly extending beyond the apex, the two meeting on a line with 
the suture above the true elytral apex, and defined from the same by 
a slight groove, borders evenly arcuate from base to apex or more or 
less parallel basally; base evenly but not strongly emarginate; 
humert proper broadly rounded and not prominent; sides proper 
more or less evenly arcuate; apex proper not produced and narrowly 
rounded; disc plane, feebly convex, at times slightly coneave, the 
inflexed sides nearly straight and oblique, gradually and not strongly 
arcuately declivous behind; surface sculptured with approximate 
series of fine asperate punctures, which become more irregular and 
slightly denser laterally. Each puncture bears a short and rather 
robust curved seta. 

Epipleure narrow, not attaining the humeral margin and not 
dilated, but gradually narrowing to apex, not defined from the in- 
flexed sides of the elytra and on the same plane; superior margin 
obsolete, except near apex; elsewhere represented by a line of punc- 
tures or a faint groove. 


REVISION OF ELEODIINI—BLAISDELL. 4 


Sterna more or less dull, finely and not distinetly sculptured. 

Parapleure smooth, rather sparsely but not very distinctly punctate. 

Abdomen horizontal, very finely and sparsely punctulate, obso- 
letely rugulose and quite evenly convex. 

Legs rather slender, moderate in length. Anterior femora mutic, 
protibial spurs similar in the sexes, the anterior slightly longer than 
the posterior. Protarsi simple. 

Male—Abdomen moderately convex and not noticeably impressed : 
anterior tibiw quite suddenly and briefly constricted at base. 

Female.——Abdomen rather strongly convex and the protibiwe grad- 
ually narrowed at base. 

Measurements.—M ales: Length, 16-18 mm.; width, 9.5-10 mm. 
Females: Length, 15-18 mm.; width, 9-10.5 mm. 

Genital characters, male—Edeagophore elongate oblong-ovate, 
rather slender, feebly arched and slightly depressed. 

Basale oblong, very moderately convex, sides feebly arcuate. 

Apicale triangular, slightly depressed; surface rather feebly con- 
vex and without groove; sides nearly straight or feebly and broadly 
sinuate to apex, the latter acute and not attenuated; base distinctly 
subacutely lobed at middle, feebly and broadly sinuate laterally. 

Sternite quite parabolic in outline. Each lobe rather long, with 
external border more or less evenly arcuate or straight in basal half, 
thence feebly arcuate to subtruncate to apex, the latter narrowly 
rounded and rather prominent; internal border oblique and short; 
surface scarcely convex, rather densely punctate in apical two-thirds, 
setose, setee rather long and not dense. 

Female.—Genital segment (Plate 5, fig. 7) triangular, rather elon- 
gate, including apices, or subtrapezoidal without the same, surface 
quite plane and not noticeably setose. 

Valvula.—Dorsal plate rather elongate-oblong or oblong-oval ; sur- 
face scarcely concave, smooth, very sparsely and somewhat coarsely 
punctate; external border more or less feebly arcuate and slightly 
explanate; internal border arcuate; apical margin more or less trun- 
cate, angle rather broadly rounded, internally continuous with the 
surface of apex, not setose. Apex chitinous, produced, slightly 
longer than the width of dorsal plate at basal third, everted, convex 
above and concave beneath, tip rounded, external border feebly ex- 
cavated at base and slightly beneath the apical margin of dorsal 
plate, forming the fossa. 

Appendage somewhat spherically mammilliform, glabrous, with a 
pencil of quite long sete at tip. 

Basal prominences obsolete. 

Superior pudendal membrane attaining the middle of the dorsal 
plate, finely and longitudinally rugulose. 


476 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Ventrolateral surfaces—Body somewhat elomgately triangular, 
obliquely truncate laterally at base, quite acute at the internal mar- 
gins of valves in the median line at base; surface lines straight when 
viewed longitudinally, evenly convex from side to side, smooth and 
shining. Submarginal groove well defined beneath the feebly ex- 
planate external border of the dorsal plate, terminating at the apical 
margin. Internal margins of the valves contiguous in basal four- 
fifths; fissure narrowly fusiform in apical fifth, margins finely setose, 
inferior membrane not visible. 

Habitat.—Texas (Mobeetee, July, H.S. Barber) ; Colorado (Pueblo, 
October 27, H. Soltau; Colorado Springs, Greeley, June, elevation 
6,000-7,000 feet, Wickham ; Fort Collins, Bellevue, West Las Animas, 
Wickham); Kansas (Wallace County, elevation 3,000 feet, F. H. 
Snow; Rice County, Clark County, June, elevation 1,962 feet, F. H. 
Snow) ; Nebraska; South Dakota (Alexandria). 

Number of specimens studied, 40. 

Type destroyed. 

T ype-locality—‘ Arkansas at the Rocky Mountains,” (Say.) 

Salient type-characters—Maregins of the thorax and elytra broadly 
foliaceous and strongly reflexed; basal angles of the prothoracic mar- 
gins projecting strongly backward over the basal angles of the elytra. 

Diagnostic characters.—Distinct from all other species as indicated 
by the salient type-characters. 

The concavum of LeConte is simply a larger, longer, and narrower 
form, with the broad elytral margins more suddenly and almost 
perpendicularly reflexed. 

General observations —The mentum is moderate in size and vari- 
able. The middle lobe may be more or less arcuate anteriorly, or 
truncate, evenly slightly emarginate at apex; sides more or less 
straightened, and converging anteriorly. The form anterior to the 
lateral and subbasal angles is subparabolic or subtriangular. 

Prosternum arcuate antero-posteriorly and somewhat prominent 
ventrally with the cox; sometimes more or less oblique and feebly 
compressed posteriorly. 

Mesosternum variable as to the degree of obliquity; usually sub- 
vertically arcuate and broadly, feebly concave. 

The abdominal process is subquadrate and just the least transverse, 
and equal in width to the length of the third segment; the post-coxal 
part of the first segment about equal in length to that of the third; 
the second is twice as long as the fourth; the third is about one-third 
of its length shorter than the second. 

The abdominal and metasternal salients are quite equal. 

The metasternum laterally between the cox is about equal in 
length to the width of a mesotibia at apex. 


. 


se 


REVISION OF ELEODIINI—BLAISDELL. 477 


~ The tibial grooves of the femora are entire and well defined, gla- 
brous, concave on the profemora, less so on the meso- and meta- 
femora; the margins are acute and smooth, gradually converging to 
the femoral base, where they meet. 

The protibiw are scarcely at all compressed nor carinate exter- 
nally, simply flattened or feebly concave in apical half, forming : 
slight tarsal groove, which is moderately asperate. 

The meso- and metatibizw are very feebly flattened externally in 
apical half. 

The tibiw appear to be quite variable as to length. In some ex- 
amples the protibiz are scarcely more than one-half the length of the 
metatibiwe; while in others they are at least three-fifths as long. This 
seems to be an individual variation observed more or less throughout 
the tribe Eleodiini. 

The protarsi are about one-fourth of their length shorter than a 
mesotarsus. The four basal joints are small, the second, third, and 
fourth subequal in size and length and about as wide as long; the 
first is about one-half longer than wide; the fifth about equal to 
the three preceding joints taken fopethier, 

The mesotarsi are about one-fifth of their length shorter than a 
metatarsus. Joints two, three, and four are subequal in length and 
slightly longer than wide; the first is distinctly more than one and 
a half times longer than wide; the fifth about equal in length to the 
preceding three combined. 

The metatarsi are about half as long as a metatibia. The first joint 
is about equal to the fourth; the second and third are subequal, the 
second apparently just a little longer than the third, and both taken 
together about equal to the fourth. 


Genus ELEODIMORPHA, new. 


Body rather robust and blapyliform. 

Head not deeply inserted, epistoma distinctly emarginate, sides of 
the front straight and converging anteriorly and not in the least 
dilated, frontal suture scarcely evident; eyes narrow, transverse, and 
subreniform; antennie 11-jointed, outer three joints feebly com- 
pressed. Mentum moderate, trilobed, lateral lobes rather small and 
inflexed, middle lobe trapezoidal, narrowing at base as usual and 
attached toa short gular pedune ‘le, surface concave. 

Maxillary palpi moderate in length, last joint triangular and mod- 
erately broad; third subtriangularly obconical, a little longer than 
wide and about two-thirds as long as the second, the latter elongate 
and obconical. 

Labial palpi small, last joint more pyriform than triangular, con- 
stricted at base on internal and lateral surfaces. 


478 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Buccal processes of the gene strongly and subacutely produced 
anteriorly over the base of the maxille. Mandibles emarginate at 
tip, cusps apparently equal. 

Prothorax moderately transverse and margined; pronotal apex 
deeply and broadly emarginate (Plate 12, fig. 39), base truncate; pro- 
sternum longitudinally impressed before the coxe, prominent in the 
median line and continuously so with the intercoxal portion, which 
is mucronate posteriorly ; anterior margin of the prosternum arcuate 
at middle and broadly sinuate laterally. 

Elytra with the dise not strongly deflexed laterally, distinctly 
margined by the superior border of the wide epipleuree which occupy 
the whole of the inflexed side, the superior epipleural margin visible 
from above throughout its entire length. 

Mesosternum and metasternum short and not prominent ventrally. 
Anteriorly the mesosternum is concave, posteriorly meeting the meta- 
sternal intercoxal salient at middle of the coxe, suture slightly arcu- 
ate. Metasternal epimera very distinct. Intercoxal process of the 
first abdominal segment rectangular and broad, as usual. 

Legs rather short, femora moderately compressed and widest near 
the middle; tibiz slightly but distinctly dilated apically, feebly but 
characteristically arcuate, spurs small and normal; tarsi moderate in 
length and thickness, joints proportioned as in the other members of 
the tribe, spinous beneath with the plantar grooves distinct. 

The tribal characters are distinct and the species has a very 
Blapylis-like form and the general structure is the same. The types 
were for years associated with Elcodes parvicollis, scabrosa, and 
cordata in my collection. 

The produced buceal processes of the gene are as marked as in 
Nyctoporis. Eleodes blanchardi presents the same condition, but to 
a much less degree. 

The maxillary galea is evidently quite different from that observed 
in Eleodes and apparently two-jointed. I could not determine posi- 
tively without dissection. 

Salient generic characters.—Epistoma emarginate, sides of front 
not dilated, buccal processes of the gene produced (Plate 8, fig. 24%), 
mentum concave; impressed sides and prominent middle of the pro- 
sternum, the latter with the anterior margin arcuate at middle and 
sinuate laterally (Plate 8, fig. 24°); wide epipleura, elytral dise 
not inflexed at the sides; femora compressed and widest at middle, 
tibize arcuate and evidently dilated apically. 

These characters define a very distinct genus, very unlike anything 
else found in the United States. 

From its southern habitat I suspect that it may be related to some 
Mexican form and have carefully searched all the literature to de- 
termine that fact, but without avail. At present I have no way of 


os 


REVISION OF ELEODIINI—BLAISDELL. 479 


verifying the results of my previous search and consider it a new and 
very remarkable addition to our fauna. 
Type of the genus.—Eleodimorpha bolean. 


ELEODIMORPHA BOLCAN, new species. 


Feebly ovate, somewhat robust; a little more than twice as long as 
wide; black, somewhat opaque; feebly setigerous beneath; dorsally 


the very minute sete scarcely project out of the punctures. 


Head somewhat wide, almost plane, punctate, punctures coarse. 
confluent on the epistoma and laterally, irregular and sparse centrally. 
scattered on the vertex; sides of the frons scarcely prominent and 
feebly impressed within; frontal suture slightly evident laterally. 

Antewne moderate in length and stoutness, outer three joints feebly 
compressed and slightly dilated, third joint not five times as long as 
the second, scarcely as long as the fourth and fifth taken together; 
joints 4 to 8 rather short, somewhat obconical, the fourth just the 
least longer than the fifth, the latter, sixth, seventh, and eighth sub- 
equal; ninth and tenth slightly transverse and oval; eleventh short, 
ovate. 

Pronotum widest just in advance of the middle, transverse, about 
one-half wider than long; disc evenly and moderately convex, rather 
coarsely punctate, punctures simple, irregular, and sparse centrally, 
becoming denser and submuricate laterally; apex broadly and deeply 
emarginate, bottom of the emargination rather straight at middle 
three-fifths, about equal to the length, marginal bead not very coarse; 
sides evenly and broadly arcuate, moderately sinuate in basal fifth, 
very briefly and feebly so just behind the apical angles, marginal bead 
fine; base truncate, rather coarsely beaded, and about one-fifth wider 
than the apex; apical angles anteriorly prominent and subacute; 
basal angles rectangular. 

Propleure sparsely and submuricately punctate, and more or less 
rugulose, moderately convex, not in the least concave beneath the pro- 
notal margins, except at the apical angles. 

Elytra oblong oval, widest at the middle, a little more than one-half 
longer than wide; base truncate, scarcely to a little wider than the 
contiguous prothoracic base; huwmeri not in the least prominent, 
obtuse, angle almost distinct; s#des evenly, moderately, and broadly 
arcuate; apex obtuse; disc evenly convex from side to side, less 
strongly so toward the base, more strongly so toward apex, where it is 
arcuately declivous, not inflexed laterally; surface evenly and not 
very densely muricately punctate, punctures evenly arranged in evi- 
dent series, each puncture distinctly defined from the others; the strial 
are smaller and less muricate, the interstitial are larger and more 
evident as small shining muricate tubercles, both series less strongly 


480 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


developed centrally, at the periphery both stronger and subequal in 
size, becoming irregular about the apical declivit? 

E pipleure quite broad, occupying the whole of the inflexed sides of 
the elytra; surface smooth and opaque, sparsely and finely sub- 
asperately punctate; scarcely narrowing in basal two-thirds, thence 
gradually to apex; superior margin visible from above throughout 
its entire length, forming a distinct and subacute bead for the elytra. 

Sterna.—Prosternum impressed before the cox, prominent at 
middle and continuously so with the intercoxal. portion, continuing 
forward to the apical arcuation and posteriorly ending in a conical 
mucro; surface sparsely and subasperately punctate and feebly 
rugulose laterally, on the central prominent portion coarsely muricato- 
tuberculately punctate, and with erect brownish hairs. 

Meso- and metasterna irregularly and sparsely punctate and 
more or less rugose. Mesosternum with brownish hairs. 

Parapleure coarsely but not densely punctate. 

Abdomen somewhat shining, first three segments rather densely 
punctate and more or less rugose, the small asperities are shining 
and larger centrally; last two segments more finely and densely 
punctate in their apical moieties, quite impunctate basally. 

Legs moderate in length and thickness. FFemora moderately com- 
pressed, widest near the middle, mutic in both sexes; tibize moderately 
arcuate in both sexes; spurs small and similar. Tarsi with the plan- 
tar grooves entire, marginal and apico-marginal spinules as usual 
and ferrugineous in color. 

Male—Slightly narrower. Elytra arcuately and just the least 
obliquely declivous posteriorly. Abdomen subhorizontal, moderately 
convex, rather strongly, broadly and longitudinally impressed at 
middle of the first three segments; intercoxal process concave; punc- 
tuation finely muricato-tuberculate. Legs stouter, and the protarsi 
moderately stout. 

Female—Slightly broader and a little more robust. Elytra areu- 
ately and vertically declivous posteriorly. Abdomen evenly convex, 
intercoxal process feebly impressed, segments rather sparsely and 
finely subasperately punctate. Legs less stout, anterior tarsi slender. 

Measurements—Male: Length, 13 mm.; width, 5 mm. Female: 
Length, 13 mm.; width, 5.8 mm. 

Genital characters, male-—Edeagophore (Plate 6, fig. 11) obtusely 
oblong-ovate in form, well chitinized, moderately elongate. 

Basale oblong, about two and a half times longer than wide, rather 
evenly convex, slightly depressed along the middle, with a faint indi- 
cation of a median longitudinal linear impression. 

Apicale rather broadly subpyriform; apical third strongly and 
quite suddenly deflexed, becoming immediately horizontal and lami- 
nate, broadly rounded at apex; dorsal surface glabrous with basal 


REVISION OF ELEODIINI-——BLAISDELL. : 481 


two-thirds somewhat depressed, laterally more arcuate and declivous, 
circularly membranous at center; sides arcuate at basal third, thence 
sinuate to apical third, where it is arcuate and continuously so with 
the broadly rounded apex; base feebly lobed at middle, broadly and 
feebly sinuate laterally. 

Clava (Plate 7, fig. 8) comparatively broad, flattened, and of 
nearly equal width throughout, obtuse and rounded at apex, slightly 
arcuate, with convexity outward, not contiguous in the median line, 
except. at base, ventrad to the edeagus; ale of the apicale evidently 
differentiated as separate elongate sclerites (subalew), but united by 
a suture to the apicale. . 

Sternite transversely parabolic in outline, bilobed. Each lobe sub- 
triangular; surface not strongly but slightly unevenly convex, gla- 
brous, rather sparsely punctate and pubescent in posterior two-thirds, 
pubescence denser in the apical area, hairs yellowish; external bor- 
der straight and oblique in basal three-fifths, thence straight and 
obliquely subtruncate to apex, the latter narrowly rounded; internal 
border quite straight; interlobar membrane not pubescent. Sinus 
triangular. 

Female.—Genital segment (Plate 6, fig. 12) triangular, about as 
long as wide at base, superior surface quite plane, chitinized through- 
out, apically véry finely pubescent. 

Valvula.—Dorsal plate not defined from the apex; surface gla- 
brous, plane, very sparsely and finely pubescent apically; external 
border rather straight, rounded at the angle and directly continuous 
with the feebly and broadly sinuate superior margin of the fossa; 
internal border broadly sinuate in apical half, thence feebly arcuate 
to the base. 

Apex short, rounded at tip, chitinous, fossa on its external surface, 
finely pubescent, hairs short but noticeable. 

Appendage mammilliform, short, penicillate, hairs quite long. 

Superior pudendal membrane long, reaching to the base of the 
apex, very minutely longitudinally rugulose. Basal prominences 
noticeable. 

Ventrolateral surfaces quite evenly convex in basal two-thirds, 
glabrous, here and there a minute hair, very feebly impressed before 
the apex. Submarginal groove distinct beneath the feebly explanate 
external border of the dorsal plate, subobsolete in basal fourth. 
Apex feebly convex beneath. Appendage and fossa distinctly visible 
from below. Internal margins of the valves contiguous, genital fis- 
sure apical, small, and fusiform. 

Habitat—Bolean Mountain, between Warner’s Ranch and Julian, 
San Diego County, California. "Taken from beneath a log. 

Number of specimens studied, 2 (male and female). 

597TSO—Bull. 683—09——31 


482 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Sexitypes in my own collection. " 

Type-locality—Bolean Mountain. 

Salient type-characters.—Body blapyliform, surface dull, buccal 
processes of the gene produced; prothorax deeply emarginate at 
apex, angles anteriorly prominent and subacute, base truncate; pro- 
sternum impressed before the coxee, prominent at middle and pubes- 
cent. Elytra truncate and more or less coarsely margined at base, 
adapted to the prothorax; sculpturing of evident series of small 
shining muricate tubercles; epipleuree wide, forming the inflexed 
sides of the elytra. Legs short, femora compressed, tibiz arcuate, 
tarsi simple. 

Diagnostic characters.—Distinct as defined by the salient type 
characters, and can not be confused with any other species in our 
fauna. . 

General observations—The mentum is trapezoidal, coarsely 
punctate and setigerous, surface concave. 

Prosternum between the coxe feebly grooved, not prominent 
ventrally and with the anterior median prominence forming a con- 
tinuous and feeble arcuation from the apical margin to the tip of the 
conical mucro. 

Mesosternum deeply concave to receive the prosternal mucro, sides 
prominent around the acetabula, pubescent. 

The intercoxal process of the first abdominal segment is rectangu- 
lar, slightly transverse, equal in length to the post-coxal portion, its 
width searcely equal to the combined lengths of the third and fourth 
segments. 

In the male the post-coxal part of the first segment is about equal 
in length to that of the third, the second is a little longer than the 
third. 

In the female the post-coxal part is about equal in length to the 
third, the second scarcely twice as long as the fourth. 

The metasternal salient is about a fourth (male) to a third 
(female) narrower than the abdominal salient. 

The metasternum laterally between the cox is about as long as 
the width of a mesotibia at middle. 

The superior and inferior surface lines of the femora are distinctly 
arcuate, most noticeable in the inferior at middle; the tibial grooves 
are well defined and subentire, the floor of each is flat, smooth, and 
abruptly limited by cariniform margins, they are longitudinally 
convex, the convexity is adapted to the tibial arcuation. 

In the male the lateral surfaces are longitudinally impressed along 
the inferior margins, not so in the female where the femora are a little 
less stout and the grooves limited by thinner margins. 


REVISION OF ELEODIINI——BLAISDELL. 483 


The tibiw are arcuate and each is somewhat rounded in basal half 
and thence to apex somewhat compressed and distinctly but not 
strongly dilated. 

The protibiw are the most compressed and scarcely carinate exter- 
nally, the tarsal grooves are evident but feeble and the posterior 
border of each more prominent than the anterior. 

The tarsal grooves of the mesotibiz are the best developed, their 
margins are equal. 

The external surface of the metatibiz are flattened and more or less 
grooved, 

The articular cavities of the protibizw are nearly open, on the other 
tibiw they are shut off from the grooves by well marked but somewhat 
atrophic edges. 

The relative lengths of the tarsi and also of their joints are quite 
similar in the sexes. 

In the protarsi joints two to four inclusive are quite equal and 
short, the first is longer than wide, the fifth about equal in length to 
the preceding three taken together. 

In the mesotarsi joints two to four inclusive are subequal and short, 
the first is scarcely as long as the second and third taken together, the 
fifth is not as long as the combined lengths of the preceding three. 

In the metatarsi joints two and three are equal and together about 
equal to the fourth; the first is a little longer than the fourth. 

Tarsal formula: 


Pro. Meso. Meta. Metatibie. 
Male—2. 2} at ee 
Female.—1% 24 23 4) 


Genus TROGLODERUS LeConte. 
Trogloderus LeContr, N. Amer. Entomologist, I, no. 1, July, 1879, p. 2. 


Body elongate, rough, opaque, eyes slightly oblique; sides of the 
frons moderately dilated, distinctly more prominent laterally than 
the eyes; epistoma moderately dilated laterally, scarcely in the least 
reflexed, margin arcuate laterally and emarginate anteriorly; labrum 
moderately small, transverse, somewhat short, sinuate, strongly re- 
tractile beneath the epistoma ; mandibles not at all visible from above; 
antenne with the third joint as long as the two following combined, 
eighth, ninth, and tenth distinctly transverse, somewhat perfoliate, 
eleventh narrower than the tenth; mentum rounded in front, trape- 
zoido-triangular to parabolic in outline, surface more or less im- 
pressed laterally; prothorax deeply emarginate in front, rounded at 
the sides, then briefly and strongly constricted or simply sinuate, 
apical angles stout and anteriorly prominent, basal angles small and 
more or less prominent and acute, disc more or less foveate in the 


484 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


median line; scutellum very feebly entering between the elytra, the 
latter not margined at base nor adapted to the prothoracic base, 
suture and four costz each side acutely elevated; epipleure narrow, 
entirely overlapped by the metasternal side pieces and first three 
abdominal segments, exposed and horizontal opposite the last two 
segments; prosternum quite strongly protuberant ventrally before the 
coxe and produced behind; mesosternum deeply concave; metaster- 
num and first two abdominal segments at middle, broadly and shal- 
lowly concave; intercoxal process of first abdominal segment broader 
than long; profemora more or less armed with a tooth at outer 
fourth; anterior tibiz more or less compressed and arcuate. 

The above diagnosis differs in several important points from that 
given by LeConte or Horn. 

The laterally dilated epistoma with sides arcuate is in contrast to 
the condition in the other genera of the present tribe, where the 
sides of the epistoma are sinuate or straight and convergent ante- 
riorly. 

The epistoma in the present genus appears longer on account of the 
dilatation and less broadly sinuate anterior margin. In most of the 
specimens before me the labrum is not visible, but in others it is 
entirely uncovered as in ZJeodes, and in all probability it is capable 
of greater retraction than in the other genera of the tribe, where it 
is retractile to a less degree. 

The mandibles are not visible from above—except slightly in the 
sinuation of the labrum—as in Lleodes, Embaphion, and Eleodi- 
morpha, and this is no doubt due in part to the fact that the mouth 
parts are not so strongly developed or prognathic as in the above 
genera. 

In the Seaurini (Cerenopus and Argoporis) the labrum is entirely 
invisible from above and apparently deeply concealed beneath the 
produced epistoma, at least I have never seen it protruded. 

The scutellum does not enter between the elytra to any extent in 
costatus, but more so in tuberculatus. 

The epipleure while narrow, are nearly entirely inflexed above and 
overlapped by the metasternal side pieces and first three abdominal 
segments, but they are visible beneath the humeri opposite the meso- 
sterum, horizontal and exposed near the apex opposite the last two 
abdominal segments. I have not observed such an extreme condi- 
tion elsewhere in the Eleodiini, although it is more or less indicated 
in sporadic instances. 

All specimens that I have examined are identical in this respect. 

The habitus of the insect might be considered somewhat discordant 
in the present tribe, but no more so than in the cases of Lleodes 
granosa and veterator in that genus. 


REVISION OF ELEODITINI—BLAISDELL. 485 


I do not consider it in harmony with the Scaurini as defined by 
Casey.*. On account of the short, transverse head, with eyes near the 
prothorax, it is more closely related to the Eulabini. 

In many respects it is more closely in harmony with /'leodimorpha 
than any other heretofore mentioned. 

The genital characters appear to show greater affinity with the 
Eleodiini. 

Genital characters, male-—Edeagophore oblong-ovate and acutely 
pointed. Apicale triangular, evenly convex above and acute at apex. 
Sternite bilobed, lobes separate and united by membrane. 

Female—Genital segment moderately depressed, subequilaterally 
triangular, valves more or less reflexed and fully chitinized. The 
dorsal plate is not at all defined from the apex, surface more or less 
concave, with scattered sete in apical moiety. The appendage is 
very small, semi-oval in outline, subapical and feebly penicillate. 
Superior pudendal membrane moderately exposed and reaching at 
least to the apical third of the dorsal plate. 

Ventrolaterally the submarginal groove is very feebly defined, the 
surface lines are straight longitudinally. Internal margins of the 
valves contiguous in basal half, fissure narrow in apical half; inferior 
membrane not visible. 

It can be seen by comparing figs. 4, 7, 8, 13, and 14 of Plate 6 
with those of the different species of /Jeodes, and then with figs. 5 
and 6 of the same plate, that 7rogloderus is more Eleodiinine than 
Seaurinine. Hulabis agrees with Cerenopus in having the two lobes 
of the sternite coalescent into one piece, emarginate at apex, and this 
is distinctly in accord with what appears as true Scaurinine charac- 
ters. This investigation is in its infancy and not mature for dis- 
cussion at the present time. 

For the present, at least, I agree with Dr. George Horn in re- 
taining 7'rogloderus in the Eleodiini. Future studies might indicate 
the advisability of separating it, possibly with Lleodimorpha, as a 
distinct tribe, and it will indeed be interesting to discover some other 
closely related species that might indicate its genealogical history. 

General characters—The profemora are stout and subclavate, oval 
in transverse section; the tibial grooves are shallow and feebly defined 
by asperate margins, best defined in the males, but in either sex more 
or less evanescent toward the base, narrower in costatus, especially 
in the males. 

The meso- and metafemora are comparatively slender, slightly and 
very gradually increasing in width from base to apex, but almost 
parallel, quite straight, except for the normal curve by which they 
are adapted to the sides of the body, oval in transverse section; tibial 


@ Annals N. Y. Acad. Sci., V, Nov., 1890, p. 391, 


486 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


grooves more or less evanescent. in basal half and feebly defined 
apically, very shallow but rather smooth. 

Protibiew distinctly compressed, especially along the inner side; 
external border arcuate in basal half, thence straight and witha tarsal 
groove, that evenly and gradually widens to the articular cavity 
with which it is continuous; the anterior margin of the groove is 
‘ather smooth, the posterior more or less denticulate; the anterior 
and posterior sides of the articular cavities are feebly dilated. Le- 
Conte says that the apical angle is produced, but I do not see that it 
is in the least so. In the male of costatus the protibiz are briefly and 
very abruptly constricted at base so that a right angle is formed; the 
tibize thence to apex quite even in width, as a whole slightly arcuate, 
ach internal edge very broadly and feebly sinuate. 

The meso- and metatibie are slender, straight, gradually and feebly 
increasing in size to apex, where they are somewhat. dilated, circular 
in transverse section, and more or less grooved or flattened externally 
for the tarsi; articular cavities closed. 

The protuberant prosternum is very obvious in costatus, less so in 
tuberculatus, in each it is deeply grooved around the anterior border 
of the acetabula; in ¢uberculatus the anterior margin is distinetly 
defiexed. 

Distribution.—Confined as far as known to the arid regions of 
eastern California and western Nevada, northward into Idaho. 

The two known species may be separated as follows: 

Pronotum cribrately punctate. oo = 2 eee costatus, 
Pronotum “tuberculates- eS = sree 2S eee tuberculatus. 


The synopsis of genital characters may be formulated as follows: 


Superior pudendal membrane long, reaching to the apical fourth 
of the dorsal: plates 32244 2h ee eee costatus. 
Superior pudendal membrane shorter, reaching to apical third_-___ tuberculatus. 


TROGLODERUS COSTATUS LeConte. 


Trogloderus costatus, LeContTrE, N. Amer. Entomologist, I, No. 1, July, 
1879, p. 3, pl. 1, fig. 3. 

Elongate, oblong-oval, opaque, dirty black, integuments dense; 
thorax cribrately punctate; elytral coste acutely elevated. 

Head \ess than twice as wide as long, feebly convex, more or less 
irregularly impressed, frontal suture not evident, but indicated by 
the abrupt termination of the frontal impression, at times slightly 
elevated; coarsely and confluently cribrate, the punctures at times 
more or less corroded. 

Antennw short, scarcely reaching beyond the middle of the pro- 
thorax, somewhat clavate, feebly compressed, second joint nearly one- 
half as long as the third and not quite as long as wide, third joint 
about equal to the next two taken together, fourth the least longer 


REVISION OF ELEODIINI—BLAISDELL. 487 


than the fifth, the latter, sixth and seventh subequal and each about 
as long as wide, eighth, ninth, and tenth transverse and gradually 
increasing in width, eleventh narrower than the tenth, triangulo- 
ovate and about as long as wide. 

Pronotum one-third to one-half wider than long, widest at the 
middle; disc rather strongly convex and moderately declivous at the 
apical angles, coarsely, more or less confluently and cribrately punc- 
tate, with two deep oval fovew occupying nearly the whole of the 
median line, sometimes forming a deep median groove, or frequently 
entirely absent; apex deeply and broadly emarginate, beaded laterally, 
obsoletely so at middle three-fifths; s‘des moderately strongly arcuate 
to basal ninth, thence briefly and strongly sinuate to base, here the sides 
are divergent or subparallel, scarcely at all sinuate behind the apical 
angles, marginal bead distinctly serrulate anteriorly, posteriorly 
obsoletely so; base transverse, scarcely beaded; apical angles large, 
prominent anteriorly, subacute; basal angles minute, acute, and promi- 
nent externally or rectangular. 

Propleure more or less convex, but not prominently so, smooth, 
more or less granulate on the acetabular convexities and posteriorly, 
sometimes rugulose. 

Elytra elongate oval, less than twice as long as wide, widest at the 
middle; 4ase not margined nor adapted to the contiguous prothoracic 
base, which is slightly wider; Awmeri obsolete; sides evenly arcuate 
and quite evenly and gradually narrowing from the middle to apex, 
the latter slightly produced and rather narrowly rounded; disc quite 
evenly and rather strongly convex from side to side, somewhat quickly 
and arcuately declivous posteriorly, but slightly and broadly sinuate 
before the moderately oblique apex (viewed from the side), the suture 
and four smooth cost each side acutely elevated, sutural costa feeble 
and evanescent on apical declivity, the humeral and subhumeral meet- 
ing anteriorly on humeral region, the subhumeral free and obsolete 
before the apex, first discal free at both extremities being evanescent 
on apical declivity, second discal uniting with the humeral before 
the apex, point of union not tuberculate nor prominent; each in- 
terval broad and apparently biseriately subfoveolate, the foveole 
more or less indistinctly marked, more or less feebly transversely 
coalescent, irregular in the outer two intervals, the minute subsquami- 
form set scattered independently of the pits. 

Epipleviw narrow, overlapped at middle by the metasternal pieces 
and first abdominal segment, horizontal in apical two-thirds; surface 
smooth, obsoletely punctate. 

Sterna sparsely, coarsély, and more or less moderately punctate; 
mesosternum deeply and broadly concave, sides prominent; metaster- 
num more or less feebly and broadly concave. 

Parapleure coarsely and more or less obsoletely punctate. 


488 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Abdomen horizontal, moderately convex, coarsely and more or less 
obsoletely punctate; intercoxal process and first two segments at 
middle third, distinctly and broadly concave, the concavity extending 
more or less upon the third segment, where it is rather semilunar in 
outline, the concavity is limited laterally. by a convex ridge, well 
marked on first segment and there continuing the internal line of the 
acetabula, Jess evident on the second segment. 

Legs rather short and scabrous; femora punctate, tibize muricate; 
anterior tibix: more or less curved and slightly compressed; tibial 
spurs similar in the sexes. Tarsi rather short; first joint of the ante- 
rior with a distinct process beneath, plantar groove not well defined ; 
apical margin of each joint somewhat prominent beneath; spinules 
flavate, short and stout. 

Male.—Somewhat narrow. Elytra arcuately and slightly obliquely 
declivous behind, abdomen strongly concave. Profemora with a 
small and broad tooth at outer fourth. Protibiz distinctly and 
briefly constricted at base, distinctly angulate at point of constriction. 
First joint of the protarsi with a distinct and more or less rounded 
process beneath, which is about a third longer than wide, truncate at 
tip, where it is very inconspicuously spinulose. 

Female.—Slightly broadened. Elytra arcuately and rather verti- 
cally declivous, thence feebly and broadly sinuate before the moder- 
ately oblique apex. Abdomen less strongly concave. Profemora 
obsoletely dentate, tooth sometimes minute. Protibize feebly and 
briefly but not suddenly constricted at base. First joint of the pro- 
tarsi with a distinct process beneath which is about a fifth longer 
than wide. 

Measurements —Males: Length, 11-12 mm.; width, 44.5 mm. 
Female: Length, 12.8 mm.; width, 5 mm. 

Genital characters, malé—FKdeagophore (Plate 6, fig. 7) oblong- 
ovate, acutely pointed, not arched. 

Basale oblong-oval, twice as long as wide, evenly convex; sides 
feebly arcuate. 

Apicale triangular, slightly elongate; dise evenly convex, without 
a groove; sides feebly arcuate at basal third, thence slightly and 
broadly sinuate at tip, the latter acute: base arcuately lobed at 
middle, laterally feebly sinuate. 

Sternite transversely trapezoidal. Each lobe triangular and rather 
long; sides rather gradually and mutually converging to apex, the 
latter subacute although narrowly rounded; surface more or less 
evenly convex, glabrous at base and impunctate, apical two-thirds 
more or less densely punctate and setose, sete quite long, denser and 
longer about apical margin. Sinus broad, membrane not setose. 

Female.—Genital segment (Plate 6, fig. 13) subequilaterally trian- 
gular, valves reflexed and fully chitinized, moderately depressed. 


REVISION OF ELEODITINI—BLAISDELL. 489 


Valvula—Dorsal plate triangulo-oblong and not at all defined 
from the apex, external margin moderately convergent apically ; 
surface more or less concave and with scattered punctures externally 
and apically, each puncture with a moderately short seta; external 
margin straight to feebly and broadly sinuate, continuous with the 
subacute margin of the apex, the latter slightly emarginate exter- 
nally for the minute fossa; internal margin quite straight; apical 
margin not in the least evident, apex not at all defined from the rest 
of the valve, obliquely conforming to the plane of the dorsal plate, 
more or less rounded at tip, which is subacute and finely setose within. 

Appendage minute and punctiform, apparently slightly com- 
pressed, calvous. 

Superior pudendal membrane long, attaining the base of the apex, 
finely longitudinally rugulose, narrowly exposed. 

Basal prominences not evident. 

Ventrolateral surfaces not longitudinally convex, moderately so 
laterally; surface very glabrous and not defined from apex; sub- 
marginal groove very feebly defined for a short distance cephalad 
to the appendage; apex with few scattered sete. Internal margins 
of the valves contiguous in basal moiety, fissure narrow in apical 
half, membrane not visible. 

Habitat.—Western Nevada and the contiguous eastern California ; 
Idaho (Rock Creek). 

Number of specimens studied, 3. 

Type in the LeConte collection. 

Type-locality.—Rock Creek, Owyhee County, Idaho. 

Salient ty pe-characters—Elongate, dirty black, opaque. Head and 
thorax coarsely and confluently cribrate, the latter with two deep 
oval fovee occupying nearly the whole of the median line. Elytra 
elongate oval, with suture and four discoidal costae acutely elevated, 
intermediate furrows deep and broad, subreticulate (LeConte). 

Diagnostic characters —The distinctly cribrately punctate prono- 
tum distinguishes costatus from tuberculatus; in the latter species 
the elytral coste are asperate and not smooth as in the former. 

The mentum is somewhat small and more or less trapezoido-trian- 
gular, foveate laterally, not noticeably strongly sculptured. 

The prosternum is strongly protuberant ventrally before the cox, 
arcuately declivous anteriorly and quite horizontal between the 
acetabula, feebly longitudinally grooved at middle from anterior 
margin; surface asperately punctate end sparsely setose, sete short; 
produced and acute posteriorly. 

Mesosternum deeply and broadly concave, sides prominent. 

Metasternum laterally between the coxe as long as the width of a 
mesotibia at middle, 


490° BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


The abdominal intercoxal process is slightly transverse, about a 
third of its width broader than the metasternal Salient and equal in 
length to the post-coxal part of the first ventral segment: second seg- 
ment is as long as the third and fourth taken together, and nearly 
three times as long as the fourth. 

The tarsi are rather short and slender. 

ry 

Tarsal formula: 


Pro. Meso. Meta. Metatibiz. 
Male.— 1} 12 1} 3h 
Female.— 1! 14 13 3% 


TROGLODERUS TUBERCULATUS, new species. 


Elongate, oblong-oval in outline, twice as long as wide, integuments 
dense and opaque, pronotum tuberculate; elytral coste acutely ele- 
rated and subserrulate, apex bituberculate. | 

Head less than twice as wide as long, plane, interocular region 
with three impressions, two lateral and one median, intervening sur- 
face convex; epistoma moderately prominent and feebly convex, con- 
tinuously so with the supra-antennal convexities, frontal suture not 
evident, but indicated by the rather suddenly convex epistoma: sur- 
face punctato-tuberculate. Antenne short, evidently not reaching be- 
yond the prothorax, feebly compressed and somewhat clavate, third 
joint about equal to the next two taken together, fourth and fifth 
about as wide as long, sixth and seventh evidently just the least wider 
than long, eighth feebly transverse, ninth and tenth distinctly so, 
eleventh small, narrower, and about as long as wide, orbiculo-ovate. 

Pronotum widest at the middle, somewhat explanate at the sides, 
one-half wider than long; disc moderately convex, distinctly arcuately 
declivous at the apical angles, rather densely tuberculate, tubercles 
small and rounded, with two deep oval fovew occupying the whole of 
the median line and about a third of the width, the fovez limited 
laterally by raised and rounded sides and separated by a similar 
transverse convexity, the concavity of each reaching the apical and 
basal margins, respectively; apex deeply emarginate in circular are 
(viewed from the front), the emargination appearing angulate 
laterally when viewed from above, without a bead; sides evenly, 
broadly, and rather strongly arcuate, quite suddenly sinuate at basal 
tenth, thence briefly straight and almost parallel to base, margin 
subserrulate; base, transverse, without bead; apical angles subacute, 
triangular, stout, and anteriorly prominent; basal angles minute, 
subacute and feebly prominent posteriorly. 

Propleure moderately convex antero-posteriorly and more or less 
concave between the somewhat explanate sides of the pronotum and 
the acetabular convexities, irregularly granulate and more or less 
rugulose, 


REVISION OF ELEODIINI—BLAISDELL. 491 


Elytra oval, about a third longer than wide, widest at the middle; 
base not margined nor adapted to the contiguous prothoracic base, 
not narrower; Auwmeré rounded or feebly subangulate at the meeting 
of the humeral and subhumeral coste; s/des evenly and moglerately 
arcuate, apex not noticeably produced when viewed from above, 
rather broadly rounded; disc rather strongly convex from side to 
side, slightly depressed on the dorsum, arcuately declivous poste- 
riorly, but sinuate before the slightly oblique apex ; surface sculptured 
with a sutural and four costw on each side, each acutely elevated, 
sutural costa feeble and evidently distinct to the apex, subhumeral, 
humeral and outer discal costw unite on each elytron near apex in 
a distinct tubercle, the internal discal costa becoming evanescent on 
the declivity; each costa is distinctly subserrulate on crest from a 
series of impressed and rather distantly spaced punctures, each 
puncture with a minute seta; intervals broad and apparently biseri- 
ately subfoveolate, the foveole rather distinctly marked and more 
or less transversely coalescent, giving a subreticulate appearance ; 
each interval with sparsely placed minute elevated granules, which 
bear a minute seta projecting from their base behind; sculpturing 
irregular in the outer intervals. 

Epipleure narrow and entirely overlapped by the sternal side 
pieces and first three abdominal segments, exposed apical portion 
horizontal, also briefly exposed beneath the humeri. 

Sterna and parapleurw more or less scabrous and somewhat tuber- 
culate. Sterna feebly concave. 

Abdomen horizontal, moderately convex, first segment distinctly 
flattened and slightly concave between the coxe, second segment 
very feebly flattened at middle third; surface more or less scabro- 
tuberculate. 

Legs short and scabrous. Femora tuberculately punctate, with a 
minute tooth at outer fourth on the profemora. Tibiw muricato- 
tuberculate, protibiz somewhat constricted at base. Tarsi short and 
feebly grooved, first joint of the protarsi moderately produced at tip 
beneath. 


Measurements —Male: Unknown. Female: Length, 12  mm.; 
width, 6 mm. 


Genital characters, male——Not studied. 

Female.—Genital segment (Plate 6, fig. 14) triangular, a little 
longer than wide, fully chitinized, valves slightly reflexed. 

Valvula.—Dorsal plate elongate, oblong-triangular; surface gla- 
brous and slightly longitudinally concave, very sparsely punctate and 
setose apically, sete short; external border more or less feebly sinu- 
ate, somewhat prominent toward base; internal margin straight. 
Apex not defined from the dorsal plate, briefly sinuate externally, 
forming a fossa, tip rounded, but not broadly so; internally finely 
setose on the valvular membrane. 


492 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Appendage small, flattened, and semi-elliptical with two or three 
short sete at tip. 

Superior pudendal membrane attaining the apical third of the 
dorsal plate and not rugulose. 

Basal prominences not evident. 

Ventrolateral surfaces with surface lines straight when viewed 
longitudinally, moderately convex transversely; surface glabrous, 
sparsely punctate apically and feebly concave laterally before the 
apices, the latter concave beneath. Submarginal groove searcely evi- 
dent. Internal margins of the valves contiguous in basal two-thirds, 
fissure subapical, with margins feebly setose. Membrane not visible. 

[abitat—California (Los Angeles County). 

Number of specimens studied, 1. 

Monotype a female (Cat. No. 12236) in the U. S. National Museum. 

Type-locality—lLos Angeles County, California; collector, D. W. 
Coquillett. 

Diagnostic characters.—The tuberculate pronotum, which is some- 
what dilated at the sides, subserrulate coste of the elytra, outer 
three coste meeting in a tubercle upon the apex, sufficiently char- 
acterize this species as distinct. 

The female genital characters, if constant in a series, will also 
prove of value, as the superior pudendal membrane is much shorter 
and not longitudinally rugulose in tuberculatus. 

General observations.—The mentum is moderate in size and sub- 
parabolic in outline, obsoletely foveate laterally, and not noticeably 
scabrous. 

The anterior margin of the prosternum is quite strongly deflexed, 
the surface is gradually and not strongly arcuate between the coxe, 
grooved only between the same and mucronate behind. The surface 
is transversely flattened in front of the coxe at middle and very 
decidedly in contrast to the protuberant condition observed in 
costatus. 

The mesosternum is rather narrowly and deeply concave and 
moderately prominent laterally. 

The metasternum laterally between the cox is about as long as 
the width of a mesotibia at middle. 

The abdominal intercoxal process is subquadrate and about as 
wide as the metasternal salient, and as long as the post-coxal part 
of the first ventral segment, quite equal to the length of the second, 
which is subequal to the third and fourth segments taken together; 
the fourth is about one-half as long as the third. 

The tarsi are rather short and somewhat stout, at least stouter 
than in the female of costatus before me. 


Tarsal formula: 
Pro. Meso. Meta. Metatibia. 

Male,— 
Female.— 1] 14 13 33 


~~ 


REVISION OF ELEODIINI—BLAISDELL. 493 
SPECIES NOT RECOGNIZED. ' 


I have not been able to recognize nor to correlate the following 
species in the ample material at hand: 


ELEODES VICINA LeConte. 


Nigra, nitida, thorace subtiliter punctulato, latitudine haud longiore, lateribus 
modice rotundatis, postice subangustato, elytris discrete minus subtiliter seria- 
tum punctatis punctis paucis interjectis, femoribus muticis. Long. .7. 

Habitat ad flumen Gila. : 

?. quadricollis attinis, at elytris punctis majoribus magis discretis distinetius 
seriatis differt. Mas, subcylindricus, elytris dorso convexis, postice attenuatis. 
Femina, elytris latioribus, dorso minus convexis, postice vix attenuatis. 

The type is from Sonora, which is the same as that of Arizona. 
Mr. Blanchard writes me that the two males in the LeConte collee- 
tion are more like carbonaria than quadricollis, the side margins of 
the elytra being stronger and the punctures at the sides more 
substriate. 

Colonel Casey, who has no doubt seen the types, writes: ¢ 


Vicina is distinct from quadricollis in having much finer punctures, which 
are sparser and much less asperate towards the sides, and also in several other 
characters, among which may be mentioned the form of the protherax, widest 
at anterior third in quadricollis and just before the middle in vicinus, the much 
longer posterior tarsi in the male of vicinus. Vicinus is peculiar to the Gila 
Valley of Arizona. 


A male from M. L. Linell’s collection, bearing a label with vicinus 
written upon it, evidently in his handwriting, corresponds to Le- 
Conte’s description, and very closely to two males in my own col- 
lection from Arizona and without definite locality. 

Linell was considered a careful student and his label bears some 
weight. His specimen is quite different from one which Fall ques- 
tionably refers to vicina. , 

I do not believe vicina to be distinct, and it must be a race or form 
of carbonaria, or quadricollis near the varieties anthracina and 
lustrans: the form of the protibial spurs in the female must decide 
which. 

ELEODES NITIDUS Casey. 


The following is Colonel Casey’s description : 


Rather slender, convex and subeylindrical, black throughout and strongly 
shining, glabrous. 

Head rather sparsely punctate, moderately coarsely so toward apex: an- 
tenn robust, nearly as long as the head and prothorax, the third joint four 


"Ann, New York Acad. Sci., V, Nov. 1890, p. 395. 


494 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


times as long as wide and fully as long as the next tyo together, last three 
joints distinctly dilated, forming a club, the ninth and tenth strongly trans- 
verse. 

Prothorax quadrate, very nearly as long as wide; disc just visibly wider 
at apical third than at base, evenly convex, externally, minutely, and sparsely 
punctate; apex transverse, broadly and feebly bisinuate, equal in width to the 
base; sides parallel, very feebly arcuate; base broadly and rather strongly 
arcuate; apical angles right, narrowly rounded; basal angles broadly obtuse 
and not prominent nor at all rounded. 

Elytra two and three-fourths times as long as the prothorax, and in the 
middle nearly one-half wider, about twice as long as wide; base broadly emar- 
ginate; humeri right, not noticeably exposed; sides broadly arcuate; apex 
acutely rounded; disc obliquely declivous behind, having feebly marked, distant, 
completely unimpressed series of extremely minute punctures; intervals mi- 
nutely, sparsely, and more or less confusedly punctate, the punctures not 
distinctly larger or denser laterally. 

Femora all slender and completely unarmed; spurs of the anterior tibize 
very unequal, the anterior slender, acutely pointed, and moderate in size; 
posterior tarsi slightly compressed, nearly three-fourths as long as the tibie. 

Prosternum slightly prolonged but not at all reflexed, the apex vertical. 

Length, 18 mm.; width, 6.7 mm. 

Habitat— Arizona. 

Casey’s specimen was a unique and he compares it with longicollis. 
He writes that it differs from that species in its flatter, rather shorter 
prothorax, which is less rounded on the sides, and in its shorter, 
broader elytra, broadly and rather strongly emarginate at the base 
and having widely distant rows of punctures, and in having a shorter, 
stouter, and much more clavate antenne. 

It is possible that the species described by me as dissimilis may be 
nitidus. Dissimilis is very variable. (See p. 398.) 


ELEODES REFLEXICOLLIS Mannerheim. 


I have failed to satisfactorily place this species of Mannerheim’s, 
the type of which was taken at Ross, California. This locality is 
undoubtedly Fort Ross, near the mouth of the Russian River on the 
coast. 

Mannerheim says that it is close to Hleodes planata Eschscholtz, 
and that the sides of the thorax are unusually reflexed, and the base 
abruptly narrowed. Peculiar and aberrant forms are occasionally 
met with. (See scabricula for remarks upon a unique specimen 
taken in Napa County, California, and in the collection of A. 
Koebele.) 

BARLY STAGES OF THE ELEODIINI. 


Comparatively little has been written upon the early stages of the 
Tenebrionide of the United States, and also little upon the tribe, 


See. 


REVISION OF ELEODIINI——BLAISDELL. 495 


treated in this paper. The only articles accessible are those of Carl 
F. Gissler*. Some of the figures are so very poor in these papers 
before me that they are of little use. 

Mr. Gissler writes that in studying the larval states of the Tene- 
brionidw we find that there is scarcely a coleopterous family whose 
members are homologically so closely allied and correlatively alike, 
and are all referable to three typical larval forms: 

1. Those having homogeneous corneous segments, cylindrical in 
longitudinal series, the stigmata situated in the pleurites. Example, 
that of Tenebrio. 

2. Form more compact, with only the prothoracic segments corne- 
ous and abdominal segments slightly arcuate. The dorsal segments 
do not overlap, forming a marginal bead, above which (on the tergal 
part of the segment) the abdominal stigmata are situated. Example, 
that of Bolitophagus and Bolitotherus. 

3. Clypeus not entirely separated, mandibles more strongly dentate, 
terminal joint of labial palpi large and truncate, pygidium without 
articulated spines. 

The larve of the /Jeodiini belong to the first form. They are elon- 
gately cylindrical, resembling those of the Elateride, which are com- 
monly called wireworms. 

The former differ in having the cardo of the maxille connate and a 
transverse impression immediately behind the clypeus, thus appar- 
ently separating the same from the front, and also by the better de- 
veloped labrum (Gissler). 

Very young larve have their antennal joints more globular than the 
older ones. In full-grown larve of the Eleodiini the first joint of 
the antenn is much shorter than the second, its substance soft, non- 
chitinized and retractile; its position is in a minute cavity, which is 
surrounded by a semi-circular ridge (Gissler). 

William R. Erichson has described this as the first joint in the larva 
of Tenebrio.” : 

From my dissections I regard this simply as the basal articulation 
with its membrane. Compare figs. 9 and 10, Plate 13. It is quite 
evident that the antenne are retractile, but when dissected out it is 
composed of only three joints. 

Mr. Gissler obtained eggs of /leodes gigantea from specimens kept 
in captivity. These were discovered in October, and measured in 
length 0.8 mm.: width, 0.85 mm.;: elongate oval and milk white in 

* Bull. Brooklyn Ent. Soc. for 1878-1879, I, No. 1, p. 11; No. 3, p. 18; No. 11, 


p. S5. ; 
>See Archiv fiir Naturgeschichte, 1841, p. 365. 


496 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


color. In November (20th, 1877) he noticed the first larvee; none 
appeared after December. 

Mr. Gissler describes the larva of “'Jeodes gigantea and dentipes as 
follows: 

Larva (Lleodes gigantea).—Head convex above, gular region con- 
cave; all the mouth parts like Z'enebrio; antenne differing by the 
stouter second and third joints. Pygidium with 18 lateral spines, 
terminal obtuse tip, sparsely frimbriate. Color of small specimens 
(6 mm.) entirely white. 

No trace of ocelli or even dark spots on the ophthalmic region. 
The small larve have the abdominal segments nearly all flat and 
gradually after several exuviations acquiring the typical wire-shape 
and darker color. The legs, thoracic and abdominal segments as in 
Tenebrio. . 

Larva (Lleodes dentipes).—Pygidium with but ten spines, the two 
terminal are slightly longer and run parallel over the tip. The front 
legs also differ from those in the larva of gigantea; the head, mouth 
parts, antenne, middle and posterior legs are as in the latter species. 
A larva but a few hours old (Plate 13, fig. 15) has two small blunt 
terminal spines, at the sides of which are two stout, long bristles; 
there are no lateral spines. The latter appear after the first moult. 
Length, 2-5 mm. Mr. Gissler also writes that a number of L/eodes 
dentipes began to deposit their eggs in April, and the larvae grew to 
30 and 35 mm. 

The above descriptions are very unsatisfactory and give no clear 
nor complete idea of the structural characters of these larve. 

Mr. Gissler kept his larve in a breeding jar with earth and decayed 
wood. 

I have kept in captivity the several species of /eodes found about 
San Francisco and succeeded in obtaining the eggs of several species. 
In general form and color they were all alike, milky white and oval 
in form, differing only in size and this in a few fractions of a milli- 
meter; the majority were about a millimeter in length. I examined 
them carefully under high magnifying power, but the enveloping 
tunic was alike in all, no sculpturing. The following were exam- 
ined: Eggs of dentipes, quadricollis, cordata, parvicollis, and 
scabrosa. 

I succeeded in obtaining larve from only one species and that was 
dentipes; they perished during the excitement following the San 
Francisco disaster. The larve had reached the length of 9.5 and 10 
mm. I discovered them in July and I did not know how long they 
had been dead. I had not discovered them before April 18. . 

Between these larvee and those mentioned by Mr. Gissler I find 
very little resemblance. This probably depends upon a different 
age of the larve, My study and dissections were made upon two 


REVISION OF ELEODIINI—BLAISDELL., 497 


larve 10 mm. in length (Plate 13, figs. 4-14), supplemented by 
eight others. The illustrations give the characters better than a 
labored description. 


LARVA OF ELEODES DENTIPES. 


Length, 10 mm.; color, castaneo-testaceous, at times fuscous. Sur- 
face glabrous. Form elongately cylindrical, flattened ventrally. * 

The ultimate and posterior border of the penultimate segments, 
both above and beneath, sparsely clothed with long flying hairs; the 
under surface of the thoracic segments are similarly clothed, the 
hairs being a little longer and rather denser; the sides of the head 
beneath are rather more conspicuously pubescent. Similar but 
shorter hairs are scattered here and there over the dorsal surface of 
the tergites, apparently without a definite orderly arrangement, ex- 
cept laterally just a little in advance of the posterior border on each 
segment there is a longer hair, and these are observed on all the seg- 
ments, forming a series on each side; on the prothorax there is a 
hair at each angle. There is also an evident series on the sternites 
just within the lateral border on each side; four or five hairs were 
counted; the number varied, possibly from being broken off; the 
series are definite and continuous on the abdominal sterna. 

Head (Plate 13, fig. 9) somewhat circular in outline, but truncate 
at base, transverse without the ¢lypeus and labrum, slightly de- 
pressed; just behind each antenna are two ophthalmic spots; all are 
small and transverse, the anterior a little more external than the 
posterior, separated by about five times their own length; imme- 
diately in front of the anterior are three or four setigerous punctures. 

Antenne three-jointed, situated immediately behind the mandibles 
in articular cavities with rather prominent borders; joints one and 
two about equal in length and about twice as long as wide, terminal 
joint very small and bearing a seta at tip; retractile. (See figs. 10 
and 9, Plate 13.) 

Mandibles short, somewhat irregularly triangular in outline, apex 
emarginate, superior cusp longer than the inferior; superior surface 
feebly convex, slightly irregular; external surface arcuate as viewed 
from above, more strongly so just apicad to the middle when viewed 
from below; external margin of superior surface with a cluster of 
some three or four set at middle, a couple of setee more basad; in- 
ternal margin with a small triangular and acute dentiform process 
slightly basad to the mid-point between apex and molar. 

Inferior surface irregularly convex, excavated within the cuspi- 
date apex, distinctly evident as viewed from below. Internal surface 
with a prominent molar (figs. 13 and 14, Plate 13). Membranous 
lobes so evident in the adult, obsolete. 

59780—Bull. 683—09——32 


498 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Maxille rather stout. Cardo and stipes rather distantly sepa- 
rated from the mentum by a membranous structure. Palpi three- 
jointed, second joint slightly broader than the first, third distinctly 
smaller. 

Mentum subquadrate, sides slightly converging to base, apex sin- 
uato-truncate, the articulating membrane attaching it to the pal- 
piger more or less distinct; ligula a delicate median process between 
the palpi, bearing two setz at tip. 

Labial palpi short and two-jointed; first joimt about as wide as 
long, possibly a little longer if fully extended; second joint much 
narrower and smaller. 

Submentum (figs. 8s and 11s, Plate 13) trapezoidal, base moderately 
arcuate, apex truncate, sides feebly, broadly but distinctly sinuate. 
Both the mentum and submentum are sparsely clothed with rather 
long flying hairs. 

Gula subquadrate, sides somewhat arcuate basally and feebly sin- 
uate apically. Suture distinct. 

Labrum transverse, a little more than twice as wide as long, anterior 
margin slightly arcuate, more strongly rounded laterally. There is 
an apical series of about eight sete, and a basal series apparently the 
same in number. 

Clypeus with a feebly transverse impression which shows at the 
sides as an emargination, just back of which is a setigerous puncture, 
another more internal, the sete of the two sides being in line; sepa- 
rated from the frons by a distinct transverse groove which ter- 
minates at the anterior border of the antennal socket. 

Prothorax nearly quadrate on the dorsum, slightly transverse, in 
some specimens transversely and feebly concave. J/eso- and meta- 
thorax much shorter and together scarcely as long as the prothorax. 

Abdominal segments subequal in length. 

Spiracles—There are nine pairs. The anterior is situated under 
the margin of the prothorax in a depression external to the coxa, 
larger than the others and with the margins more prominent and 
chitinous. The abdominal are in segments 1-8 inclusive and situated 
at the anterior margin of the pleurites; they are circular and flat, 
margins not prominent. 

Pygidium (Plate 13, fig. 5) triangular, sides moderately arcuate, 
apex rather broadly rounded. The marginal series of spines con- 
sists of eight on each side, the terminal two are just noticeably a 
little stouter and longer than the others. 

The specimen from which fig. 5 was drawn shows three spines dis- 
placed inward from the proximal extremity of the marginal series; 
another specimen had them present only on one side and in others 
they were entirely absent. I was unable to determine if this was the 
result of wear. 


afc 


YF ———— 
. Pe = 


REVISION OF ELEODIINI—BLAISDELL. 499 


Mr. Gissler notes that in one specimen studied by him there were 
two spines articulated in one cavity, while in another specimen they 
were asymmetrically placed, nine on one side and eight on the other. 
In the sculpturing of the pygidia a few smooth spots were observed 
in Eleodes. ® 

Legs similar in form, the anterior larger and stouter, claws of the 
middle and posterior are distinctly more slender; the anterior are 
more strongly chitinized. The coxe are quite conical and transverse, 
also prominent. 

The anterior has spines arranged as follows: At inner side of base 
of claw a spine points inward, tibiw with three spines on inner sur- 
face, femur with three, trochanter with two, each podomere with 
seattered flying hairs of varying lengths. 

The middle and posterior legs also have spines somewhat differ- 
ently arranged and more difficult of description. 

Mr. Gissler writes that in 7'enebrio and Fleodes (Plate 13, fig. 15) 

the first post-embryonic stages are alike in the form of the pygidium. 
The integumentary appendages of larval pygidia may be regarded as 
the homologues of the dorsal parapodia of their ancestors—the 
Annelides. 
_ Two very minute tubercles can be seen under strong microscopic 
power at the middle of the base of the pygidial tips in //codes. 
Whether they are orifices or not I can not decide. The larve of £7/e- 
odes when disturbed and handled violently jerk their abdomen and 
spurt a turbid, odorless, watery fluid out of two lateral orifices situ- 
ated under the tergal fold near the posterior margin of the seventh 
abdominal segment. These structures are a peculiar adaption to the 
larval body, since the excretion of the repugnatorial glands of the 
imago is spurted out through the anal opening. The higher develop- 
ment of this gland is certainly attained during the pupal period (Giss- 
ler). 

Mr. Gissler found that in Blaps mortisaga the pygidium had usually 
twelve lateral and two latero-terminal spines (Plate 15, fig. 18). Out 
of ten specimens two had sixteen in all, the rest only fourteen. In 
Eleodes gigantea and FE. dentipes he gives as the usual number of 
spines sixteen lateral and two latero-terminal. A number of either 
species had only sixteen in all. (See Plate 15, fig. 17.) 

The larve of LJeodes are abundant, but I have never had time to 
successfully rear them, to determine the species to which they be- 
longed. From the sand dunes along the coast the larve of 2. clavi- 
cornis and scabrosa have been taken. Two pupe were obtained; one 
developed a clavicornis, the other being identical was studied, figured, 
and preserved in spirits, 


500 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


PUPA OF ELEODES CLAVICORMIS. 
(Plate 138, figs. 1, 2, and 3.) 


Length, 10.5 mm.; width of body, 4.5 mm.; greatest width from 
knee to knee, 7 mm.# Oblong-oval, moderately arcuate, head flexed 
against the prosternum; abdomen moderately depressed ; color yellow- 
ish white, appendages semitranslucent. 

Head exposed, vertex visible from above, clypeal region convex, 
frons somewhat transversely impressed, vertex rather more promi- 
nent; antenne curving backward against the sides of the prothorax, 
over the profemora, at which point “they are not visible when viewed 
from below. 

Pronotum evenly convex; base almost truncate, basal angles rather 
broadly rounded, sides arcuate, apex feebly and broadly emarginate, 
with angles somewhat narrowly rounded. 

Mesonotum short, transverse, and moderately convex; elytral pads 
passing obliquely backward and ventrally above the pro- and meso- 
crura, apical fourth between the meso- and metacrura, visible ven- 
trally only at this point. Epipleural margins apically contiguous to 
basal half of the mesotarsi. 

Metanotum a little longer than the mesonotum, moderately convex, 
and broadly sinuate behind. 

Legs prominent laterally, not appressed against body, distinctly 
compressed and comparatively broad; tarsi distant from each other 
in median line, except the metatarsi, which are in contact in apical 
half; coxe and sterna visible in the median line. 

Abdominal segments convex dorsally, less so ventrally; pleural 
region of segments one to seven produced laterally into subquadrate 
laminiform processes; last two segments without lateral processes 
and more evenly convex from side to side. Fourth ventral segment 
deeply and broadly emarginate at apex; fifth smaller, rather arcuate 
at apex and within the emargination of the fourth; third segment 
broadly and less deeply emarginate. 

Abdomen at apex terminating in two elongate processes, each grad- 
ually tapering from base to apex, the latter chitinous; beneath the 
bases of these cerci there is a small segment which is deeply emar- 
ginate, with angles prominent posteriorly. 

The lateral processes of segments one to seven are limited at base 
dorsally by a longitudinal impression on either side of the dorsum; 
the impression begins on segment one and ends on base of the seventh ; 
ventrally the impressions are less strongly marked. 

In the specimen at hand the processes of segments two to six are 
deeply and semicircularly emarginate at middle third, each anterior 
and posterior third is squarely truncate at apex, with angles sub-_ 
acute, those at the emargination are chitinous and denticulate, each” 


- we el 


REVISION OF ELEODITNI—BLAISDELL. 5Ol 


dentate angle has a small seta at middle of its side toward the 
emargination; contiguous sides of the processes of each adjoining 
segment narrowly chitinous, very minutely and irregularly denticu- 
late; posterior third of the process of first segment not developed ; 
processes of seventh segment imperfectly developed. Anal cerci with 
a few scattered sete. 

Pronotum with a lateral marginal row of comparatively closely 
and evenly spaced chitinous set, internal to which is a submarginal 
series of distantly spaced sete, often broken off and appearing as 
minute chitinous tubercles; there is also a post-apical series of very 
distantly spaced and similar sete, as well as an antebasal series simi- 
larly spaced. 

In Plate 13, fig. 2, it will be seen that the post-apical series appear 
as marginal when viewed from above. 

On the dorsal surface of the head there are four to six sete; dor- 
sal surface of abdominal segments three, four, five, and six with a 
seta each side near basal and external angle. Legs with a short seta 
on outer anterior femoral region near the knee; fourth ventral ab- 
dominal segment with a subapical row of sete. 


Tribe BLAPTINI. 


In (Blaps) the globular outer antennal joints, wide epipleure 
which terminate upon the external surface of the cauda when present, 
the intercoxal tubercle of the first abdominal segment and the pubes- 
cent pit at base of the second abdominal segment the present tribe 
stands in marked contrast to anything observed in the genus /’/eodes, 
and in fact all of the genera of the EJeodiini, except in the wide 
epipleure which forms the whole inflexed sides of the elytra in 
Eleodimorpha. 

The genital characters are certainly very distinct and more primi- 
tive than is to be seen anywhere in the LVeodiini. 

Genital characters, male—Lobes of the sternite strongly connate, 
clave connate, lying dorsad to and forming a pallium for the edeagus, 
more obsolete ale, more embryonic condition of the edeagophore, 
there being a complete solution of continuity in the chitinization 
along the median line of both the basale and apicale, these segments 
being longitudinally subdivided by an evident membranous line into 
lateral sclerites. 

Female.—Valves of the genital segment (Plate 5, figs. 16 and 17) 
completely chitinized, apex of each undifferentiated, appendage ab- 
sent. The facies of the segment is quite different from that observed 
in the Zleodiini, but a phylogenetic relationship is evident. 

I am not desirous to repeat what may already be known in Europe, 
but simply to indicate my reasons for not retaining the genus 4laps 
in the same tribe with the others dealt with in this paper. 


502 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


It is only by the assiduous working of data and in the making of 
deductions in accordance with the laws governing ontogenesis and 
variation that our knowledge will be complete or as full as scientific 
progress will permit. 

The student must bear in mind that the characters of Blaps given 
have been drawn up from a very limited number of species, and that 
it is to be expected that a genus so rich in species will present varia- 
tions analogous to those observed in /leodes. I wish to point out 
that the genus is undoubtedly a divergent stem from the ancestral 
phylogenetic trunk from which /7eodes also had its origin. A de- 
tailed study along the lines which I have laid down might be the 
means of greatly modifying our view of relationships as at present 
accepted. ° 

Genus BLAPS Fabricius. 


This well-known genus of the Old World needs no treatment at 
my hands and is given a place here on account of two species having 
been introduced into the United States and therefore are to be con- 
sidered as a part of our fauna. 

In many characters it resembles H7eodes and also presents several 
very distinct and divergent characters. 

Mentum transverse, plane and more or less variable in form, some- 
times trapeziform, sometimes rounded at the sides and in front, at 
times subtruncate, rarely sinuate, always narrowing at base for a 
short distance. The lateral lobes are small and invisible. 

Mawillary palpi elongate, last joint securiform, longer than wide. 

Labrum prominent, transverse, rectangular or subcordiform, angles 
rounded and more or less sinuate at middle of the anterior border. 

Head prominent, not deeply inserted, frontal suture distinct or 
obsolete; epistoma with sides more or less straight and convergent 
anteriorly, apex truncate or feebly sinuate. 

Eyes transverse, narrow, subreniform or sublunate. 

Antenne moderate, 11-jointed, third joint long, 4 to 7 of variable 
length, obconical, 8 to 10 globular, 11th oval, pointed at tip. 

Prothorax variable; sometimes almost plane and rectangular, at 
others more or less convex and rounded at the sides, the latter always 
margined. 

Scutellum of variable size, sometimes not visible. 

Elytra at base not as wide or wider than the contiguous protho- 
‘acic base; form and length variable, frequently attenuated poste- 
riorly and caudate at apex. 

Epipleure attaining the humeral angle where the superior margins 
are visible from above, gradually narrowing to elytral apex. Legs 
generally long, especially the posterior; anterior femora more or less 
thickened; tarsi moderate, spinous beneath, plantar grooves distinct. 

Prosternum mucronate or not. 


REVISION OF ELEODIINI—BLAISDELL. 503 


Male characters.—Narrower, finer punctuation, greater length of 
elytral cauda when present. Abdomen more or less moderately 
convex, first abdominal suture more or less impressed with base of 
second segment and apical moiety of the first, giving the abdomen 
the appearance of being oblique and the sterna somewhat promi- 
nent; first suture with a fossa at middle, densely clothed with golden 
pubescence on the segmental margins; sometimes a truncated tubercle 
on the intercoxal process of the first abdominal segment. 

Female characters.—Broader form, somewhat coarser punctuation. 
Abdomen rather strongly convex. 

Genital characters —The characters observed in the specimens at 
hand are as follows: 

Male.—Edeagophore, length 5.5 mm.; edeagus 1.5 mm. Very 
elongate, somewhat flattened, strongly arched, somewhat oblong- 
ovate and well chitinized. 

Basale, elongate oblong, about four times as long as wide, mod- 
erately convex from side to side, divided by a median membranous 
groove which extends from base to apex, apically continuous with 
the groove of the apicale. 

Apicale elongately triangular, at least twice as long as wide, some- 
what depressed, disc feebly convex, narrowly and more strongly so 
laterally, a long median groove extending from base to apex: base 
quite broadly and strongly lobed at middle three-fifths, laterally 
broadly and feebly sinuate; sides slightly and evenly arcuate, regu- 
larly converging to the subacute apex, the latter rendered minutely 
emarginate by the median groove. 

Sternite transversely suboblong, lobes strongly connate and con- 
tinuously chitinized, transversely prominent at middle, the surface 
sloping apically and basally; apical moiety broadly concave; basal 
half less broadly concave at about middle third, laterally the surface 
is convex and gradually diminishes toward the lateral angles of the 
apex; apical margin broadly and evenly emarginate: external 
borders moderately arcuate; apical angles somewhat prominent and 
rather narrowly rounded. 

Apical moiety densely pubescent, hairs of moderate length and 
rather coarse, basal portion glabrous. 

Clave absolutely not evident as distinct structures. They are 
present but connate forming a dorsal pallium for the edeagus. The 
margins of the pallium gradually attain the midline ventrad to the 
base of the edeagus, becoming contiguous but not connate; beyond 
the middle of the edeagus the pallium is entirely dorsal, with sides 
distinctly chitinous and semimembranous at middle. The apex of 
the pallium projects just a little beyond the caput edeagi. Alw 
strongly inflex and somewhat obsolete. 


504 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Female.—Genital segment (Plate 5, figs. 16 and 17) triangular, elon- 
gate, general dorsal surface plane and glabrous, appendages absent. 

Valvula fully chitinized, apex not differentiated from the dorsal 
plate, surface slightly irregular, at middle minutely and ocellately 
punctured, a minute seta arising from each puncture, apex slightly 
deflexed and subacute, very feebly convex above; external border 
broadly and feebly sinuate at middle, arcuate basally and apically; 
internal margins rather straight in apical two-thirds, arcuate and 
converging from base where they are moderately distant to become 
contiguous at middle third. 

Superior pudendal membrane long, reaching to the base of the 
apical region, very finely longitudinally rugulose. 

Ventrolateral surfaces convex basally, less so towards apex, mod- 
erately concave beneath the slightly deflexed apices; internal margins 
of the valves contiguous basally and apically, genital fissure fusiform 
and just basad to the middle: submarginal groove obsolete, some- 
what evident beneath the apical margin. Dorsal plate not at all 
explanate. Apical two-thirds very sparsely and finely pubescent. 

The following two species can be credited to our fauna: 

Blaps similis Watreille. 

Blaps mucronata Latreille. 

T shall not attempt a specific diagnosis. Doctor George Horn com- 
pared native specimens with the European and dispelled any previous 
doubts about their correct identification. 

Habitat—A few specimens of mucronata occurred near Baltimore, 
Maryland. Similis occurs abundantly at Alexandria, Virginia. 

I have had the privilege of studying a series in the collection of the 
U.S. National Museum, said to have come from West Virginia (Riley 
Coll.) ; figs. 16 and 17, Plate 5, were taken from a broad female from 
that series. A small series was kindly loaned me by Mr. Charles 
Waterhouse, of the British Museum, including what is supposedly 
correctly identified specimens of sémilis, mortisaga, and mucronata. 
Besides these, I examined a series rather recently obtained from 
London (British Museum), England, by Miss Julia Wright. 

The specimens in our American collections do not seem to be defi- 
nitely labeled. Mr. Liebeck, of Philadelphia, writes me that in his 
collection is a specimen from Alexandria, Virginia, labeled as gibba; 
one from Boston, Massachusetts, taken in a warehouse, identical with 
a specimen (European) in the Horn collection of similis; two labeled 
mucronata, supposedly from Canada. 

In the Horn collection (Academy of Natural Science, in Phila- 
delphia) there are three forms—one a broad form taken by Doctor 
Castle at Alexandria, Virginia, and labeled gibba; two forms have 


4See Can. Ent., XVI, p. 37, pl. xxi. 


¥ 


REVISION OF ELEODIINI—RBLAISDELL. 505 


the label s/mi/is—one a broad form, single specimen from Europe, 
and the other an elongate slender form, of which some are labeled 
“ Md.” 

A study of the genitalia of Blaps and their careful comparison 
with typical forms of the genitalia of //eodes, Embaphion, and 
ELleodimorpha \eaves no doubt in my mind that they are not in any 
way directly related and that it is absolutely necessary to ereet a 
new tribe (/Jeodiin/) to receive our species. No species of the Blap- 
tint (Blaptida), except Blaps, has yet been discovered in the United 
States fauna. 

With Blaps eliminated from the /Jeodsini the tribe becomes com- 
posed of homogeneous elements, with possibly the exception of 
Trogloderus, as IT could not study the genitalia in all points of detail 
for scarcity of material. 


APPENDIX. 


In May, 1908, there appeared in the Bulletin of the Department 
of Geology, Vol. 5, No. 12, of the California University in Berkeley, 
a paper on the Quaternary Myriopods and Insects of California, by 
Fordyce Grinnell, jr. 

In this paper the author recorded seven species of Hleodes, three 
being described as new, all having been taken from asphalt beds at 
Rosemary, near Los Angeles, California. They are supposedly the 
fossil remains of recent or older species. These are the fossils 
referred to on page 29 of this monograph. 

I had the pleasure of identifying the species for Mr. Grinnell. 
The following is a list of the species: 

Eleodes acuticauda LeConte, also the punctate form; F/eodes 
acuticauda var. laticollis, Eleodes consobrina LeConte. 

The following were described as new: E'leodes (subgenus Eleodes) 
elongata Grinnell; Eleodes behrii Grinnell and intermedia Grinnell. 
The last two belong to the subgenus Blapylis. 

In order to more fully correlate these new forms with the more 
recent forms considered in the present monograph, I carefully 
reexamined the types and record the following facts: 

All of the specimens are very brittle, deep black in color, and shin- 
ing; they appear as if the asphalt had exerted some chemical effect 
upon them that to a certain extent had modified the structural 
details. 

These specimens are interesting and present some points of struc- 
ture and sculpturing which can not be correctly described until a 
larger or more perfect series shall have been collected. 

The heads, pronota, and appendages are missing from all the speci- 
mens, and the elytra alone can not tell the whole story. 


506 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


ELEODES ELONGATA Grinnell. 
> 


The type is a right elytron, fully exposed except the epipleural 
margin, which is sunken into the asphalt. The humerus and epipleura 
immediately beneath are free. The specimen is too poor and fragile 
to give a clear idea of the sculpturing. 

The elytron is in form like that of #leodes grandicollis ; the punctu- 
ation is similar, very sparse and.small, scarcely at all subasperate, 
and quite sharply defined; general surface glabrous and shining. 
Humerus rounded, and the superior epipleural margin is thin and 
sharp. 

Length, 21.5 mm.; width, 9 mm. 

T'ype-specimen.—Number 10025, University California Collection 
of Invertebrate Paleontology. 

This species really appears to be a form of grandicollis; it differs 
but slightly from some of those of to-day. Gvrandicollis is so distinet 
that an elytron or thorax ought to be readily recognized. 


ELEODES BEHRII Grinnell. 


One elytron (left), apical sixth buried in the asphalt, dorsal sur- 
face and epipleura exposed, and in a fairly good state of preservation. 

The elytral base is truncate, humerus rounded and not in the least 
prominent ; séde evenly arcuate; dish: moderately convex, broadly and 
evenly arcuately declivous laterally, not striate, punctuation diffuse, 
rather dense, subasperate, and with a tendency to coalesce in twos and 
threes, the intervals more or less convex and forming slight trans- 
verse rugule. 

E'pipleura rather broad and concave, superior margin moderately 
prominent and rather sharp, gently curving upward anteriorly to 
the humerus, thence caudad more broadly and arcuately curving 
downward to apex; surface smooth, sparsely and distinctly pune- 
tate, punctures moderately small; there is also a trace of transverse 
rugule. 

This species is without a doubt closely related to consobrina or 
parvicollis; the elytral sculpturing is more like that observed in the 
latter. 

The epipleural characters are unlike anything known to me; the 
marked concavity is rarely observed in sporadic instances in individ- 
uals of to-day; the curve of the superior margin is more. strongly 
downward to apex than in any species. It is possible that these well- 
marked characters are the result of warping in the asphalt. 

This species is the only one of the three under consideration that 
probably deserves a name. It is surely a true Eleodes (Blapylis) 
and not at all worthy of a generic standing, as the author suggest- 
ively wrote. 


 —- 


REVISION OF ELEODIINI—BLAISDELL. 507 


Length, 10 mm.; width, 4.5 mm.; greatest width of epipleura, 
1.5 mm. 

Type-specimen.—Number 10023, University California Collection 
of Invertebrate Palwontology. 

The type, an elytron which is “narrowly oval” with its fellow of 
the opposite side would give a more broadly oval figure, like that 
observed in a small parvicollis, the dorsal surface being somewhat 
flattened. 

ELEODES INTERMEDIA Grinnell. 


A complete abdomen, meso- and metathorax, with elytra, consti- 
tutes the type; the specimen is quite strongly flattened. 

The form of the body is broadly oval and truncate anteriorly, 

Elytra truncate at base; humeri rounded and not prominent; sides 
evenly arcuate; apex not narrowly rounded; disk evidently flattened 
on the dorsum, and moderately convex, more strongly and arcuately 
rounded laterally; surface quite densely and submuricately punc- 
tate, punctures coalescing more or less, the interstices convex, forming 
slight transverse rugule; sculpturing denser and coarser laterally 
and on apex, not in the least striate; surface evidently evenly and 
moderately arcuately declivous posteriorly. 

Epipleure rather broad at base, gradually narrowing to apex; sur- 
face distinctly concave throughout, finely, subasperately, and sparsely 
punctate; superior margin strong, quite sharp and prominent, not 
sinuate beneath the humeri, but straight in the anterior moiety, and 
thence quite evenly and gradually arcuately descending to apex as 
viewed longitudinally. 

Mesosternum evidently feebly arcuately declivous, deeply concave 
at middle between the prominent and rounded acetabular borders; 
very densely, rather finely and confluently punctate. 

Metasternum and episterna densely and rather finely punctate. The 
metasternum laterally between the coxee apparently about two-thirds 
as long as the post-coxal part of the first abdominal segment. 

Abdomen densely and moderately coarsely punctate. First segment 
at middle evidently as long as the second and third segments com- 
bined; intercoxal salient quadrate and distinctly transverse, surface 
concave and evidently with glabrous areas, equal in length to the post- 
coxal part of the same segment; second and third segments evidently 
equal in length and each subequal to the length of the post-coxal part 
of the first. 

Length, 12 mm.; width of elytra, 8.5 mm. 

Ty pe-specimen.—Number 10026, University California collection 
of Invertebrate Paleontology. 

This specimen is related to parvicollis, and I believe specifically 
identical with beArii; the latter is smaller and probably a male, while 
intermedia is the female. 


508 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


There is scarcely any difference at all except in size and the epi- 
pleural curve. ; 

The age of the asphalt beds at Rosemary, near Los Angeles, is 
Quaternary. The remains of the saber-toothed tiger were also taken 
from this deposit. The remains of the Coleoptera were found in the 
neighborhood of the bones. 

My colleague, Dr. E. C. Van Dyke, who has devoted time to the 
study of the origin of the Californian entomological fauna, consid- 
ers that the above data substantiate the statements that have been 
made from time to time that a great deal of the so-called Upper 
Sonoran or Californian fauna found its way into this State, from the 
south, at a very early period, probably during early Tertiary times. 
In this list are to be included most of the typical genera and sub- 
genera of the most southern parts of the State. Already at the 
time at which these deposits were made, most of the species had 
assumed the appearance that we see in their descendants, the forms 
living at the present day. 


EXPLANATION OF PLATES. 
PLATE 1. 


Fig. 1. Dorsal view of the right valve of the genital segment of Hleodes 
nigrina; showing the chitinous, moderately produced apex, evenly 
and broadly rounded at tip, with the valvular membrane rather 
largely exposed within and caudad to the superior pudendal mem- 
brane. 

2. Ventral view of the left valve of Hlcodes nigrina; showing submar- 
ginal groove, concave ventral surface of apex and part of the genital 
fissure. 

3. Dorsal view of the apicale of the edeagophore of Lleodes nigrina; show- 
ing the depressed and broadly membrano-chitinous area at middle. 

4. Dorsal view of the genital segment of Hleodes gigantca; showing the 

fully exposed superior pudendal membrane, falciform dorsal plates, 
chitinous and everted apices, and minute appendages. 

. Ventral view of the left valve of the genital segment of Hleodes gi- 
gantea; showing the broad submarginal groove continuous with the 
concavity of apex, straight internal margin of the valve, and absence 
of the genital fissure. 

6. Dorsal view of the apicale of the edeagophore of Hleodes gigantea; 
showing the broadly and strongly depressed membranous surface 
and slightly recurved apex. 

7. Dorsal view of the left valve of the genital segment of Hleodes longicol- 
lis; showing its resemblance to that of gigantea, but more strongly 
punctate. 

8. Dorsal view of the right valve of the genital segment of Eleodes grandi- 
collis ; note the truncate apical margin of the dorsal plate, rectangular 
angle, truncate appendage, and short pudendal membrane. 

9. Dorsal view of the right valve of the genital segment of Eleodes tene- 
brosa ; note the plane dorsal plate, long, slender, subacute, and mem- 
branous apex; compare with figs. 1 and 2. 

10. Ventral view of the left valve of the genital segment of Eleodes tene- 
brosa; showing evenly convex surface and subapical fissure. 

11. Dorsal view of the right valve of the genital segment of Hleodes pedi- 
noides, typical form; note the explanate and plane dorsal plate which 
is evanescent apically into the well-developed external apical lobe, 
also the small rudimentary internal lobe and long pudendal membrane. 

12. Dorsal view of the right valve of the genital segment of Eleodes ex- 
tricata, typical form; note the oval-oblong dorsal plate, acute and 
small apex, large and flattened appendage. 

13. Dorsal view of the right valve of the genital segment of Eleodes ob- 
scura, typical variety; note the dorsal position of the submarginal 
groove, dorsal plate apparently contracted with sides arcuately re- 
flexed, surface deeply concave, basal prominences strongly developed. 

14. Dorsal view of the right valve of the genital segment of Eleodes sutura- 
lis ; compare with figs. 23, 13, 15, 16, and 17. 


or 


509 


510 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Fig. 15. Dorsal view of the right valve of Hleodes dis peusa ; compare with figs. 
13, 16, 14, and 23. 

16. Dorsal view of the right valve of Hleodes sulcipennis; compare with 
figs. 13, 15, 14, and 23. 

17. Dorsal view of the right valve of Hleodes acuta; compare with figs. 
14 and 23. 

18. Dorsal view of the right valve of Hleodes armata; note the nearly 
plane dorsal plate, reflexed internally, the flattened subconical ap- 
pendage, and that the basal prominence is not well developed. 

19. Lateral view of the apicale of the edeagophore of Hleodes terana; note 
the moderately protruded edeagus between the apex above and the 
abducted clava beneath. 

20. Dorsal view of the right valve of the genital segment of Hleodes mili- 
taris; note that the dorsal plate is concave with sides reflexed, sub- 
marginal groove somewhat visible from above, appendage large and 
flattened, the basal prominences evident. 

21. Dorsal view of the right valve of Hleodes lucw; compare with figs. 18, 
20, and 22. 

22. Dorsal view of the right valve of Hleodes luce, forma inflata; compare 
with figs. 20, 21, 14, 17, and 23. 

23. Dorsal view of the right valve of Hleodes terana; note the resemblance 
between figs. 22, and 21 as compared with 13, 14, 15, 16, and 17. 


PLATE 2. 


Fig. 1. Dorsal view of the edeagophore of Hlecodes omissa; note the membra- 
nous groove of the apicale, the fully abducted clave and the extruded 
edeagus, 

2. Sternite (male) of Hleodes omissa, forma communis, exhibiting varia- 
tion from the typical form; note the sparsely setose interlobar mem- 
brane; a, the produced internal basal angle of the triangular chitinous 
plate of the sternite (see p. 18). 

3. Dorsal view of the genital segment of Hleodes omissa, common form; 
compare with figs. 6 and 9 of this plate and fig. 11, Plate 1. 

4. Ventral view of the genital segment of Hleodes omissa, common form; 
note the surface contour, the direction of the appendages and the 
broad fusiform genital fissure, the visible inferior pudendal mem- 
brane. 

Sternite (male) of Hleodes quadricollis, forma tarsalis; compare with 

fig. 2 


so 


or 


6. Dorsal view of the genital segment of Lleodes quadricollis, forma tar- 
salis; note the differences about apices of the valves. 
. Ventral view of the genital segment of Hleodes quadricollis, forma tar- 
sdlis ; compare with figs. 4 and 10. 

Dorsal view of the edeagophore of Bleodes quadricollis; compare with 
Hevl. . 
9. Dorsal view of the genital segment of Hleodes pygmea; valves slightly 

abducted; note the resemblance to fig. 3. 


=] 


10. Ventral view of the genital segment of Hleodes pygmaa; valves moder- 
ately abducted. 
11. Sternite (male) of Hleodes marginata; note the rather rounded lobes 


and long setze; compare with fig. 2; Plate 3, fig. 2, and Plate 4, fig. 15. 

12. Dorsal view of the genital segment of Hleodes marginata; note the 
nearly fully chitinized valves, the produced and everted apices, and 
punctiform appendages. Compare with fig. 16, Plate 5. 


«1 


Fig. 13. 


Fig. 1. 


ty 


ad 


» 


x 


Fig. 1. 


2. 


REVISION OF ELEODIINI—BLAISDELL. 511 


Ventral view of the genital segment of HKleodes marginata; note the 
contiguous internal margins of the valves and compare with fig. 16, 
Plate 5. 


PLATE 3. 


Dorsal view of the edeagophore of Lleodes obsoleta; see explanation 
of fig. 1, Plate 2. ‘ 

Sternite (male) of Bleodes obsoleta; compare with figs. 2 and 5 of 
Plate 2. 

Dorsal view of the genital segment of Hleodes obsoleta; note the very 
explanate dorsal plates, especially at apex; compare with figs. 3, 9, 
and 6 of Plate 2. 

Ventral view of the genital segment of Lleodes obsoleta; compare with 
figs. 4, 10, and 7, Plate 2; a, paranal plate (see p. 19). 

Dorsal view of the right valve of ELleodes letcheri; note resemblance to 
fig. 12, Plate 1. 

Ventral view of the left valve of Eleodes letcheri. 

Dorsal view of the right valve of Pleodes acuticauda; compare with 
figs. 12, 13, and 9 of this plate, and figs. S and 18 of Plate 1. 

Ventral view of the left valve of Eleodes acuticauda; note the large 
appendage and the concave surface before the apex. 

Dorsal view of the right valve of Eleodes dentipes; compare with figs. 
11 and 13 (type): note the variability of the apical margin of the 
dorsal plate and form of the appendage. 

Ventral view of the left valve of Pleodes dentipes. 

Dorsal view of the right valve of Eleodes dentipes, showing variation. 


. Dorsal view of the valve of Eleodes laticollis ; compare with fig. T. 
. Dorsal view of the right valve of Fleodes dentipes, forma confinis 


(type); compare with figs. 7, 9, and 12. 

Dorsal view of the right valve of Eleodes porcata; note the broadly 
rounded external apical lobe; compare with fig. 3; the explanate mar- 
gin of the dorsal plate is less diaphanous, so that the body of valve 
does not show through it. 

Dorsal view of the right valve of Eleodes obsoleta; variety showing the 
obsolete explanate margin toward base, 

Dorsal view of the right valve of Lleodes carbonaria; compare with 
figs. 3, 14, 17, 18, and 20; also figs. 3, 6, and 9, Plate 2. 

Dorsal view of the right valve of Eleodes humeralis; see for com- 
parison fig. 16. 

Dorsal view of the right valve of Bleodes cuneaticollis; see for com- 
parison fig. 16; note the more membranous external apical lobe. 

Dorsal view of the right valve of Eleodes hispilabris; compare with 
figs. 7, 8, 19, and 23 of Plate 4. 

Dorsal view of the right valve of Hleodes tricostata; note the distinctly 
oblong dorsal plate and compare with fig. 11, Plate 1. 


PLATE 4, 


Dorsal view of the right valve of Hleodes scabricula; compare with 
fig. 12, Plate 2; the notch defining the apex from the dorsal plate 
appears to be constant. 

Dorsal view of the right valve of Fleodes neotome; note the subapical 
position of the appendage and compare figs. 9, 10, and 16, 


512 


Fig. 


= 


oO 


10. 


11. 


BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


. Dorsal view of the right valve of Hleodes (Promus) fusiformis; note 


the broad and setose dorsal plate as compared to the species of the 
other subgenera; compare figs. 4, 5, and 22; also figs. 2 and 3, Plate 5. 


. Dorsal view of the right valve of Eleodes opaca; note the less setose 


condition. See for comparison fig. 3. 


. Dorsal view of the right valve of Eleodes goryi. See for comparison 


figs. 38 and 4. 


. Dorsal tiew of the right valve of the genital segment of Hleodes cor- 


data; note the oblong dorsal plate with apical angle evident; com- 
pare with figs. 9, 10, 11, 12, and 16, also observe the shortness of the 
superior pudendal membrane as compared to figs. 9, 10, 11, 12, and 16. 

Dorsal view of the right valve of Hleodes caudifera; compare with 
figs. 8 and 23. 


. Dorsal view of the right valve of Eleodes sponsa. 
. Dorsal view of the right valve of Hleodes consobrina; compare with 


fig. 10, which is a little less swollen at apex, but the same. 

Dorsal view of the right valve of Hleodes consobrina; compare with 
fig. 9, which is more swollen at apex, but the same. 

Dorsal view of the right valve of Eleodes clavicornis; note the elon- 
gate form and narrow dorsal plate and small membranous apex; 
compare with fig. 22, Plate 5. 


. Dorsal view of the right valve of Hleodes snowii; compare with fig. 9, 


Plate 1, and fig. 21, Plate 5. 
Dorsal view of the right valve of Eleodes granosa; compare with figs. 
14 and 28, noting the short superior pudendal membrane. 


. Dorsal view of the right valve of EHleodes pilosa; compare with figs. 


25 and 28; also with fig. 12, Plate 1; fig. 5, Plate 3. 


. Sternite (male) of Hleodes sulcipennis; compare with fig. 5, Plate 2; 


also fig. 2, Plate 8, and fig. 11, Plate 2. Note the circular membranous 
area at base between the lobes, the latter densely pubescent. 


. Dorsal view of the right valves of Eleodes tibialis; note the elongate 


form, short appendage, and apex; compare with figs. 2, 10, and 11, + 


. Dorsal view of the right valve of Bleodes veterator; note the undiffer- 


entiated apex from dorsal plate, and absence of the appendage. 


. Dorsal view of the right valve of Eleodes femorata; compare with figs. 


17 and 20, Plate 1. 


. Dorsal view of the right valve of Eleodes gracilis; compare with fig. 


23; also figs. 9 and 11, Plate 3. 


. Bleodes innocens.—Dorsal view of the apicale of the edeagophore; note 


resemblance to fig. 6, Plate 1. 


21. Hleodes innocens (female).—Dorsal view of the genital segment; com- 


26. 


pare with figs. 4 and 7, Plate 1. 


. Bleodes insularis (female).—Dorsal view of the right valve; com- 


pare with figs. 8, 4, and 5, Plate 4. 


. Bleodes longipilosa (female).—Dorsal view of the right valve; com- 


pare with figs. 7 and 8, Plate 4. 


. Hleodes knausii (female).—Dorsal view of the right valve; compare 


with figs. 16 and 3, Plate 3. 


. Eleodes arcuata (female).—Dorsal view of the right valve; compare 


with fig. 12, Plate 1. 

Bleodes debilis (female).—Dorsal view of the right valve; note the 
less rounded and more strongly chitinized dorsal plate and short 
membrane; compare with fig. 16, Plate 38. These characters were 
present in all specimens examined. 


REVISION OF ELEODIINI—BLAISDELL. 513 


ig. 27. Bleodes ampla (female).—Dorsal view of the right valve; compare with 
fig. 16, Plate 3. 
28. Eleodes granulata (female).—Dorsal view of the right valve; compare 
with fig. 12, Plate 1, and fig. 5, Plate 3. 


PLATE. 8. 
Fig. 1. Eleodes ventricosa (female).—Dorsal view of the right valve; note the 
oblique apical margin and prominent angle; compare with figs. S, 21, 
and 22 of Plate 1; figs. 12 and 13 of Plate 3. 
2. Bleodes striolata (female).—Dorsal view of the right valve; compare 
with fig. 3; also, figs. 3, 4, 5, and 22 of Plate 4. 
3. Eleodes subnitens (female).—Dorsal view of the right valve; com- 
parisons the same as for fig. 2. 
4, Bleodes parvicollis (female; typieal).—Dorsal view of the right valve; 
compare with fig. 22; also, figs. 2, 6, and 10, Plate 4. 
vt 5. Bleodes pimelioides (female).—Dorsal view of the right valve; compare 
with fig. 6, Plate 4. 


“ 6. Bleodes planipennis (female).—Dorsal view of the genital segment: 
' note resemblance to figs. 7, 8, 9, 14, and 20; also, compare with figs. 
7 6 and 10, Plate 4. 

a : 7. Embaphion muricatum (female).—Dorsal view of the genital segment ; 


comparisons are to be made with figs. 6, 8, 9, 14, and 20, 
Z 8. Embaphion contusum (female).—Dorsal view and comparisons the same 
as for fig. 7. 
9. Embaphion planum (female).—Dorsal view and comparisons the same 
as for fig. T. 

10. Bleodes schwarzii (female).—Dorsal view of the genital segment; com- 
pare with figs. 11 and 12, noting that the valvular apices are quite 
different, but fully chitinized; compare also with fig. 1, Plate 1. 

| Bleodes nevadensis (female).—Dorsal view of the right valve; compare 
L with fig. 12; and also fig. 1, Plate 1; note the produced parallel apices 
with tips broadly rounded, convex above and concave beneath. 
; 12. Bleodes dissimilis (female).—Dorsal view of the right valve. 
18. Eleodes inculta (female).—Dorsal view of the genital segment; com- 
pare with figs. 9 and 10, Plate 4. 
14. Embaphion glabrum (female).—Dorsal view of the right valve; com- 
pare with figs. 6, T, 8, 9, and 20. 
Eleodes hornii (female).—Dorsal view of the genital segment: note the 
tumid apices and smooth membrane; compare with fig. 9, Plate 4. 
16. Blaps similis (female; West Virginia).—Dorsal view of the genital 
segment; note the fully chitinized and coalesced condition of the 
parts; compare with fig. 12, Plate 2. 

17. Blaps similis (female; West Virginia).—Ventral view of the genital 
segment: note the position of the genital fissure and compare with 
fig. 18, Plate 2. 

18. Eleodes subpinguis (femate).—Dorsal view of the right valve; compare 
with fig. 19, Plate 4, and fig. 9, Plate 3. 

19. Bleodes distans (female).—Dorsal view of the right valve; compare 
with fig. 19, Plate 4, 

20. Embaphion contractum (female).—Dorsal view of the apical moiety of 
the right valve; compare with figs. 6, 7, 8, 9, and 14. 

597S80O—Bull. 683—09——33 


a 
= 


Pi — =eat 
mT. 


— 
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514 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Fig. 21. Eleodes lecontei (typical; femmale).—Dorsal view of the right valve; 
compare with figs. 10 and 12, Plate 4. > 
22. Eleodes scabrosa (female).—Dorsal view of the right valve; compare 
with fig. 11, Plate 4; also fig. 4 of this Plate. 


PLATE 6. : 


Fig. 1. Bleodes blanchardii, female.—Dorsal view of the right valve; compare 
with fig. 10, Plate 4. 

2. Eleodes hoppingii, female.—Dorsal view of the genital segment; compare 
with fig. 4, Plate 5. 

8. Bleodes caseyi, female.—Dorsal view of the genital segment; compare 
with fig. 14, Plate 4, and fig. 5, Plate 5. 

4, Dorsal view of the genital segment of Cerenopus concolor. 

5. Edeagophore of Cerenopus concolor, dorsal view. 

6. Sternite of Cerenopus concolor. 

. Dorsal view of the edeagophore of Tregloderus costatus, 

. Sternite of Trogloderus costatus. 

9, Dorsal view of the edeagophore of Vyctoporus aquicollis. 

10. Dorsal view of the genital segment of Nyctoporus oequicollis. 

11. Edeagophore of Eleodimorpha bolcan, dorsal view. 

12. Dorsal view of the genital segment of Bleodimorpha bolean. 

13. Dorsal view of the genital segment of Trogloderus costatus, female. 

14. Dorsal view of the genital segment of Trogloderus tuberculatus, female. 

15. Last two joints of the maxillary palpi of J7'rogloderus tuberculatus. 

16. Mentum of Lleodes nigrina, female. 

17. Labrum of Lleodes nigrina, female. 

18. Inferior interungual process of a metatarsus in Eleodes dentipes, male. 

19. Unguis from metatarsus in #. dentipes, male. 

20. Mentum of Discogenia marginata, male. 


PLATE T. 


Fig. 1. Ventral view of the male genitalia of Fleodes cordata, edeagophore 
partly extruded. 
2. Dorsal view of the same, edeagophore completely extruded. 
3. Ventral view of the genital segments of Hleodes cordata, female, valves 
abducted, 
4. Dorsal view of the same. 
5. Ventral view of the sternite of Lleodes parvicollis, apicale slightly visible. 
6. Ventral view of the apicale of the edeagophore of Bleodes cordata, show- 
ing clay partly abducted and edeagus nearly protruded. 
Ventral view of the apical half of the edeagophore of Embaphion con- 
tusum, Showing parts in place. 
8. Ventral view of the apical half of the edeagophore of Hleodimorpha 
bolcan, Showing relative position of edeagus, cClavee, and subale; com- 
pare with figs. 7 and 1. 


=a 


Beplanation of figures.—a, apieale; b, edeagus; ¢c, clava; d, basale; e, sternite; 
f, apex of right lobe of sternite; g, membranous interlobar region of ster- 
nite; 4, membranous apex of the valve; i, appendage; j, apical angle of the 
dorsal plate; k, submarginal groove; 7, explanate external border of the 
dorsal plate; m, ventro-lateral surface; nm, membranous Sth segment, with 
sclerites of the paranal region and their articulation with base of the ventro- 


- 


REVISION OF ELEODIINI—BLAISDELL. 515 


ateral Jamina showing through; 0, sternite of the 7th segment: o’, median 
basal process of the same; p, sternite of the Gth segment, figs. 1 and 3; tergite 
of the 6th segment, figs. 2 and 4; q, supra-anal plate; r, tergite of the 7th seg- 
“ment; s, fossa for the appendage; ¢, superior pudendal membrane; u, dorsal 
plate; v, valvular membrane; 7, inferior pudendal membrane; «, genital ori- 
fice; y, inflexed alee of the apicale; 2, inflexed sides of the basale, 


ra PLATE 8. , 

Fig. 1. Head of Pleodes dentipes, showing lines of measurements: a, median 

Sy line; b, post-marginal ocular line; ¢c, post-ocular line. 

2. Left mandible of Lleodes quadricollis (female), internal surface show- 
ing: molar (¢), lacinia (b), and post-molar membranous lobe (d). 

3. Labium of Lleodes dentipes: a, the chitinous paraglossa which is connate 


with the body. 

4. Diagram showing the relative lengths of the different segments of the 

legs and abdomen in Lleodes sulcipennis: 

A, femora: a, metafemora; b, mesofemora; ¢c, profemora. 

B, tibite: d, metatibia; ¢e, mesotibia; /, protibia. 

C, tarsi: g, metatarsus; /, mesotarsus; i, protarsus. 

D, abdominal segments: j, first visible; k, second; 1, third; m, fourth; 
n, fifth. 

5. Mentum and labium of ZLleodes scabrosa, 

6. Mentum and labium of F. clavicornis, slightly extended to show relation 

% between labium and lateral lobes of the mentum, the middle lobe 
i free at apex. 

Protibial spurs of Hleodes quadricollis, female: a, posterior; b, an- 

ve terior. 

Side view of the first two joints of the protarsus of Eleodes sulcipennis, 

. female: a, tuft of modified spinules at apex beneath, the latter 

thickened and produced; b, apico-marginal tuft of spinules bounding 

5 groove at apical margin of second joint; c, apical spines. 

5 9. Outline of side view of first two joints of the protarsus of BF. sulci- 

= pennis, male, showing the greatly thickened (@) and produced apices 

; beneath, spines not shown, groove obliterated. 
Ja 10. Inferior surface of right mandible of FP. dentipes, female, * 28. 

i 11. Maxilla of EF. dentipes, female: a, palpus; b, galea; c, lacinia. 

12. Superior surface of right mandible of &. dentipes, female, X 28. 

13. Hypopharynx of FE. dentipes, female, * 28. 

Protibial spurs of 2. sulcipennis, female: a, anterior; posterior without 
letter. 

15. Inferior view of the mouth parts of BF. dentipes, female, showing men- 
tum, submentum, gular sutures, labium partly exposed, maxilla, and 
mandibles. 

16. Ventral view of the first two joints of a protarsus of FE. dentipes, male, 
showing entire plantar groove. 

17. Ventral view of first two protarsal joints of 2. dentipes, female, showing 

plantar groove entire. 

Lateral view of first two protarsal joints of FE. scabricula, male, showing 
first joint moderately produced at tip beneath, groove entire on the 
second. 

Lateral view of first two protarsal joints of B. scabricula, female, show- 
ing tip of first joint more produced with tuft of moderately fine modi- 
fied spinules. 


a 


ad 


_ 
a 


— 
9° 


19 


516 


Fig. 20. 


Zi, 


NOTE 


BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Outline of side view of first two protarsal joints gf BE. quadricollis, male; 
tip of first joint scarcely at all produced, with apico-marginal tufts. 

Outline of side view of first two protarsal joints of #. quadricollis, 
female; first joint distinctly produced at apex beneath, with tuft of 
modified spines, groove interrupted. 

Outline of side view of first two protarsal joints of 2. longicollis, male; 
a, tuft of golden and rather soft pubescence, groove interrupted; b, 
second joint with pointed apico-marginal tufts, groove entire between 
the same. 


. Outline of side view of first two protarsal joints of H#. longicollis, 


female; joints partly tipped to one side showing the entire plantar 
groove. 

Bleodimorpha bolcan.—a, mentum, buccal fissures and produced buceal 
processes; the submentum is subobsolete, gular sutures distinct as in 
Bleodes; b, outline of the ventral apical margin of the prothorax, arcu- 
ate at middle, sinuate laterally. 


PLATE 9. 


. Dorsal view of the edeagophore of Hydrophilus triangularis; a, and d, 


the inner and outer lobes of Packard. 

Ventral view of the same. 

Dorsal view of the edeagophore of Blaps similis, showing the apicale as 
formed by the uniting of the inner lobes of Packard. 

Ventral view of the same, showing the pallium (@) and edeagus (d). 

Dorsal view of the edeagophore of Eleodes dentipes. 

Ventral view of the same. 

Ventral view of edeagophore Hleodes planipennis. 

Dorsal view of the edeagophore of Eleodes seabricula. 

Ventral view of the same. 


. Dorsal view of the genital segment of Hydrophilus triangularis, female. 
. Ventral view of the edeagus (a), basal chitinous processes (¢) and 


clave abducted (d) of EBleodes dentipes. 


. Dorsal view of the same. 


PLATE 10. 


.—The bilateral asymmetry is intentional to illustrate variations which 


are constantly observed, when the pronota are viewed from above. 


ia hea se 


13; 


Pronotum of Eleodes debilis, female. 

Pronotum of Hleodes omissa, male. 

Pronotum of Bleodes omissa, female: variations a and b. 
Pronotum of Bleodes omissa, female; variations @ and b. 
Pronotum of Eleodes pygmea, female type. 

Pronotum of Eleodes pygmaa, female, variation. 


. Pronotum of ELleodes ampla, male. 
. Pronotum of Zleodes ampla, female: a, marginal curve as viewed 


obliquely from the side and from above; b, as viewed vertically from 
above. 


. Pronotum of Hleodes carbonaria, forma glabra, female. 
10. 
. Pronotum of Lleodes carbonaria, var. soror, female. 


Pronotum of Bleodes carbonaria, forma typica, female. 


Pronotum of HLleodes obsoleta, var. porcata, male. 
Pronotum of Llcodes obsoleta, male; a and b, showing variations in form. — 


REVISION OF ELEODIINI—BLAISDELL. 517 


Fig. 14. Pronotum of Bleodes obsoleta, female; a, as seen vertically from above; 
b, as seen obliquely from above and to the side. 


15. 


19. 


Pronotum 
form. 
Pronotum 
form. 
Pronotum 
form. 
Pronotum 


of Lleodes knausii, male; illustrating two variations in 


of Bleodes knausii, female; illustrating two variations in 


of Eleodes quadricollis, male; illustrating two variations In 


: 
of Eleodes quadricollis, female; illustrating two variations in 


form: a, viewed vertically from above; b, as seen obliquely trom 
above and to the side, 


Pronotum 
Pronotum 
Pronotum 
form. 
Pronotum 
Pronotum 
Pronotum 
Pronotum 


. Pronotum 


Pronotum 
Pronotum 
Pronotum 
Pronotum 
Pronotum 
Pronotum 
Pronotum 
form. 
Pronotum 


. Pronotum 


of Bleodes humeralis, female. 
of Bleodes humeralis forma granulata-muricata, 
of Eleodes cuneaticollis, male; illustrating two variations in 


of Eleades tricostata, female. 

of Bleodes neomexricana, female. 

of Eleodes pedinoides, male. 

of Bleodes obscura, female. 

of Eleodes hispilabris, female. 

of Eleodes sponsa, female. 

of Eleodes sponsa, female; a variation. 
of Fleodes caudifera, female. 

of Eleodes caudifera, male; a variation. 
of Licodes longipilosa, female. 

of Dleodes gracilis, female. 

of Eleodes dentipes, female; illustrating two variations in 


of Eleodes dentipes, female; taken from an aberration, 
of Eleodes dentipes, female; drawn from an aberration in my 


own collection; the right side is that of confinis, and probably a 
reversion to the ancestral armata. 


Pronotum 


of Eleodes dentipes forma confinis, female; drawn from the 


type in my own collection; probably a reversional form. 
Pronotum of Bleodes armata var. impotens, female; drawn from the type. 


. Pronotum 
. Pronotum 


Pronotum 


. Pronotum 


Pronotum 
Pronotum 


. Pronotum 
. Pronotum 


Pronotum 
Pronotum 


. Pronotum 
. Pronotum 
. Pronotum 


Pronotum 


. Pronotum 
. Pronotum 


of Bleodes armata, female: a variation. 
of Eleodes armata, male. 
of Bleodes militaris, female. 


PLATE 11. 


of Eleodes femorata, male. 

of Bleodes laticollis, male. 

of Eleodes dcuticada, male; typical. 

of Bleodes luca, caudate form. 

of Eleodes eschscholtzi, female. 

of Bleodes tenuipes, male. 

of Bleodes wickhami, male. 

of EBleodes ventricosa, male. 

of Eleodes_ventricosa, female; variation. 
of Eleodes falli, male; drawn from type. 
of Eleodes grandicollis, male. 

of Eleodes insularis, female. 

of Bleodes subnitens, female. 


518 


Fig. 


14. 
15. 
16. 
a7 
18. 
aie} 
20. 
21. 
22. 


9° 


mt De 


24. 


Cs at 


=. 
26. 
21. 
28. 
29. 
30. 
31. 
82. 


29 
De 


bo oe 


ot go 


OOD 


— 
> . 


BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 


Pronotum 
Pronotum 
Pronotum 
Pronotum 
Pronotum 
Pronotum 
Pronotum 
Prenotum 
Pronotum 
Pronotum 
Pronotum 
Pronotum 
Pronotum 
Pronotum 
Pronotum 
Pronotum 
Pronotum 
Pronotum 
Pronotum 
Pronotum 
Pronotum 
Pronotum 
Pronotum 
Pronotum 
Pronotum 


Pronotum 
Pronotum 
Pronotum 
Pronotum 
Pronotum 
Pronotum 
Pronotum 
Pronotum 
Pronotum 


. Pronotum 


Pronotum 
Pronotum 
Pronotum 
Pronotum 

lating h 
Pronotum 
Pronotum 
Pronotum 
Pronotum 
Pronotum 
Pronotum 
Pronotun 
Pronotum 
Pronotum 
Pronotum 
Pronotum 


of Eleodes fusiformis, female. 
of Eleodes striolata, male. 

of Eleodes goryi, female. 

of Eleodes opaca, female, 

of Lleodes veterator, female. 
of Eleodes arcuata, female. 

of Eleodes exrtricata, male; variation. 
of Eleodes extricata, female. 
of Eleodes granulata, male, forma tuberculata. 
of Lleodes vandykei, male. 

of Eleodes vandykei, female. 

of Bleodes letcheri, male. 

of Eleodes letcheri, female. 

of ELleodes pilosa, male. 

of EBleodes pilosa, female, variation. 

of Eleodes pilosa, female. 

of Bleodes hirsuta, female. 

of Bleodes granosa, male. 

of Bleodes granosa, female. 

of Eleodes tibialis, male. 

of Hleodes inculta, female. 

of Eleodes snowii, female, New Mexico. 
of Fleodes snowii, female, Arizona. 

of Eleodes tenebrosa, female. 

of Eleodes clavicornis, female. 


PLATE 12. 


of Hleodes hornii, female. 

of Eleodes neotome, female. 

of Eleodes consobrina, female, typical form. 
of Bleodes scabrosa, male. 

of Eleodes rotundipennis, male. 

of Eleodes rotundipennis, female. 

of Bleodes parvicollis, male, typical form. 

of Bleodes producta, male. 

of Bleodes fuchsii, female. 

of Eleodes cordata, female. 

of Eleodes pimelioides, male, two variations. 
of Eleodes caseyi, female. é 

of Bleodes nigrina, male, showing two slight variations. 


of Eleodes nigrina, female, drawn from an aberration simu- 


umeralis. 

of Bleodes perlonga, male. 

of PRleodes nevadensis, male. 

of Eleodes dissimilis, male. 

of Bleodes schiwarzii, female. 

of Pleodes gigantea, male. 

of Bleodes gigantea, female. 

of Rleodes estriata, female. 

of Dleodes longicollis, male. 

of Bleodes longicollis, male, elongate cylindrical form. 
of Rleodes innocens, female. . 
of Eleodes marginata, male. 


je 
R 


go go 1 


ad 


ba 


. Pronotum of Embaphion clongatum, male. 


REVISION OF ELEODIINI—BLAISDELL. 519 


. Pronotum of Eleades scabricula, male. 


Pronotum of Lleodes planipennis, female. 


. Pronotum of Embaphion muricatum, male. 


Pronotum of Lmbaphion contusum, female. 
Pronotum of Embaphion planum, wale. 
Pronotum of Embaphion glabrum, female. 


Pronotum of Lleodes hornii, male. 
Pronotum of ELleodes distans, male. 


. Pronotum of Eleodes subpinguis, female. 


Pronotum of Lleodes gentilis, male, drawn from a LeConte type. 
Pronotum of Pmbaphion contractum. 


. Pronotum of Trogloderus tuberculatus, female. 


Pronotum of Eleodimorpha bolcan, mate. 


PLATE 183. 


Pupa of Eleodes clavicornis, ventral view. 

Pupa of Bleodes clavicornis, dorsal view. 

Pupa of ELleodes clavicornis, lateral view. 

Larva of FLleodes dentipes, dorsal view, 10 mm.; x 10. 

Dorsal view of the pygidium of Pleodes dentipes. 

Ventral view of the last five abdominal segments of the larva of Bleodes 
dentipes, showing position of stigmata on the pleurites. 

Ventral surface of the right prothoracic leg of the larva of Eleodes 
dentipes. 

Ventral view of the maxilla of the larva of Eleodes dentipes; lettering 
same as on fig. 11. 


. Dorsal view of the head of the larva of EB. dentipes. 
. Antenne of the same, X 28. 
. Gula (g), mentum (7m), submentum (s), &@ membranous lobe (a), and 


labium of larva of FE. dentipes. 
Dorsal surface of the left mandible of the larva of Eleodes dentipes. 


. Ventral surface of the right mandible of the larva of Eleodes dentipes. 
. Internal or oral surface of the left mandible of the larva of Eleodes 


dentipes, showing molar. 


. Pygidium of Lieodes larva shortly after hatching, x 60. After Gissler. 
. Antenna of Blaps larva. After Gissler. 

. Pygidium of Pieodes larva. After Gissler. 

. Pygidium of Blaps larva. After Gissler. 


- 
INDEX. 
The black-faced numbers indicate generic or specific descriptions. 
A. Page 
‘ Page. | contractum, Embaphion...............-.- 453,460 
- acuta, Blaps...............-. feeeeeaeeeeeesens 195 | contusum, Embaphion..................- 453,467 
Eleodes........-..---++-++-e++002++ 185,195 | convexa, Eleodes=connexa............ 217,220, 222 
acuticauda, Eleodes.............-.---- 234,276,505 | cordata, Eleodes............... 19,310,311,313,879 
7 Akis(?) NE ig in os sch aae racket nant aal 473 costatus Trogloderus 486 
alternata, Pimelia..........-------++000++++ 104 | cuneaticollis, Eleodes.-.........c.ccececa 38, 40,91 
od. enn ncauwanwentpan 3 SR 8 a et ad ae Sas 
PRIOR Son cos oo osc ccennesomcenes 38, 40,53 D. 
anthracina, Eleodes.............-.---++-++-- 87 | debits, Eleodes......-...-.----ccoccceoss 38, 40,41 
peters, Blcodes.....-.............-.---- aaa hacdoialh ee a ene 185, 187 
arcuata, Eleodes......-.......-.-+--+++-++++. 116 | dentipes, Eleodes........-...--- 8,234,251, 496,497 
,@ Euassomensis, Eleodes......-...........2.--- 116,125 ideneriicn, Flaodes....2.-<--..ca-ssseeseeetee 453 
_ armata, Eleodes...-......-....-.--+-+-+--- 234,259 | depressum, Embaphion................----- 453 
_ aspera, Eleodes....................-.- 127,180,131 | Discopenia...........<<--+s-eee0-caeeeeae 33,35, 484 
CE ESE ea Ne ee 109, 111 Group , Pa ee ee re ely 434 
BR. NSPOUIEE decline kaa tear ates 434 
MN es ca cacacwapaee 505, 506 dispersa, MANUS ii, my, aan Satire so 185, 187, 190,191 
 blanchardii, Eleodes .............. 310, 311,312,339 | dissimilis, Eleodes................-.------ 393,398 
OR ee 3, 25,29, 30,502 | distans, Eleodes.............-..--+++++-++ 235,242 
te ne BS ae a 105 | | OGlOSN, TELGQUGO saan dwnSiacs on ennusnd al 38, 40,57 
I oo Son t's es xn cos ae ee se 45 E. 
TINGS, oC cc pct ocn dat cache nnnbananns 121 
ee eRe OF ee a 217 | Early stages, Eleodfini ................-.... 495 
. mucronata....- ED EAL OR RE ae 504 EET, SNOORES sea si ccy nds os du avhad oncsneby 495, 496 
Oe ee eee 187} elegans, Eleodes........... equa nee in = uke 251,255 
Ce 2 Pek he Ba | OOM CS ce san whan cenaaw 3,29,30,31, 34,35, 183 
Noe. ons ou ty inwant ae kcaunon 175 QOUCR. son nies cacavnccnenntucnccsucnnsaceps 185, 195 
Rt elo 504 ROUUCKUGE .. - 5 ces eecneccevensesees 234,276,505 
MMR Poon. on oR dcu cece. 199 var. laticollis.... 234, 276,280,505 
ta ee ee re | 104 MTN ea xa 5 A edie dna danwaewnacureee 38, 40,53 
EL ae ea a 505 CRT MOMOEB own oc ineies -nscccieecscones 38,40,57 
I rte 58 tice cade vk Sasdcw 501 SUTRA. « «5 -< 5. snccneeesvawees-a2ae 38, 40,87 
OS ee 34, 35,306, 505 SY ek eee Sn ee ee 185, 187, 194 
bolean, Eleodimorpha....................... 47 ATCUALB. .....-- +222 e-e seer seen eeeeereveee 116 
brunnipes, Eleodes.................... 313, 384, 386 GERRI s sin sae payasn ana dgsancbapadhnns 116,125 
oe i Se eee ey ee 234,259 
c. Pr ROUT iis Wow dvin aadsbus eens 234, 259,263 
NS RUMI Ds 355 cerca Sens dane 45 BRL Ga Tata sb dinates cs kbancurt an Oe 127,130 
1 et ee 38, 40,45 MMREMR Cen eal a dasienda~srasascectk ans 109,111 
MS MMMOANG nn ccsactnecae>stnanct AE O08 Ts a aa eng Nu ends cenpcacsare 505, 506 
Gauidifera, Eleodes. .. 2. ...2.c..scccncacces 212, 225 Eas ons cc ssedureescasd 310, 311,312, 
clavicornis, Eleodes........... 310,311,312, 371,500 INES oh bis on dui sanckiceuewnneaws 38, 40,45 
OCs EOMODOES. 3. so cnc esass=duasdnhwa 121,123 DMN fc dan Sncinesanveesiuseeed 38, 40,50 
compositus, Eleodes. .. 2... ....s.sncsssensws 217 Pe Pi ahtarreekaeidaper seiusarinets 3, 13,388 
concavum, Embaphion.............. Se Pe 473 i Ee ee eae ab potate abe 212,225 
CO ROOM, Suid ann sane shawveelent dey 251,255 ES ae ae 310,311, 312,371,500 
COME IO, cco ddemidevinn heracedassace 217 St nha seetticn dkians ord oceuknp ab wan 121 
consobrina, Eleodes............... 310,311,333, 505 SN nach Sekthiuhs 4udu exp ou cnbouske 217,221 
comstricta, Eleodes. .................+. SOO (1 oe IRS o bccn dhaBa ere scaknacsccdevstes 251,255 
521 
9 


~s 


522 INDEX. 

Page Page. 
Bleodes connexd. a2 --=.car-abexeeceraese seer 217 | Eleodes nevadensis PE 393,402 
CONSODTING) = eee set oeee 310,311,312,333, 505 NIGTING oss 5 sno od omens oe ee 393 
COMsiFichal: 2222-5 - fs oecce eee steer oee 312, 355, 365 var: perlonga. 2.252225 s5.25 6 sen eee 393,398 
cordatasc-23i 2523. kee eee - 19,310,311, 213,379. Hitida.. sion cosa nane ae eee 450, 493 
var. rotundipennis.....2. .<)sssceseters 379, 384 NUP TA. 22 55 aos a ae ee Spee 217,220 
cuneaticollis....... <2. ss. cosncwasasuee 38,40, 91 Obseura). ....-235 5. 0i ee 185, 187, 193 
(10) U1 {Sa i RE EF 28,40,41 par. dispersa... 23-2228 552 eee 185, 187,190,191 
deletae o-oo. been ee ee 185, 187, 194 var. suleipennis -\.../.002- eee 185, 187,190, 194 
dentipesss- tek seas oe 8,234,251, 496, 497 var. typica.... 0.2. oe eee 190, 193 
CG Presses - 322. sec ne cece Sell) eee 458 obsoleta.<. . .:..00=2-.4- 02-05 38, 40,58 
GISMOTER”—2--c0. 2: cs op c= oe eee 185, 187,190,191 var. porcata..:_). 2.2. 2s-.2e eee 38, 40, 58, 63 
dissimilis:..<«.. 2. 5. ucae des eee 393, 398 obtusac 2: 2 «2 oes ss: oe 127,130 
par. Devadens!s..< << 5,.<2sssseeaoeeeeee 393, 402 OMISSA... 5 ns 0.ds so enolase 38, 40, 72 
Cistans 255 - 9... 0s.) pane ate ane eee 242 var. peninsularis... . ... 5 i245 38, 40,79 
GOlOsa: 525.5. BA ata ee eee 38, 40,47 VOM. DY PINGS... «62s etek ee 38, 40,77 
CIO PANS 20-2 he oe ee ee ees 251, 255 OPSCS «. ..-o. 5 = ooo Se 156,175 
elongata. 20-0222 es < case eee eee 505, 506 parvicollis:.. 7.3. scss.0e- eee 310, 311,312, 354 
eschscholtz - 2.0%. 2n...2seeeesesse eee 235, 285 var. constricta:: 2< =~ 22-5. 312, 355,365 
par NIG. _.< 5-4. Se Se oe 235. 285,289 planata.-. {..... +. a eee 312, 355,359 
esthidtas 2. cs .225 eek seseeeeeee = 412,421 produeta... -- 3.22545 312,355,362 
extelcata. 2: oee 6a os 21 pier krone 116,121 pedinoides..... wisiies 38, 41,109 
var: avizonensis .. -.- ..<esces tena seare= 125 var. neomexicana..-..--.---).-c-aees 38, 41,113 
fallié - os. (eS. eee ee se ree 235,305 peninsularis:. .....:-.5:2.:-5- eee 38, 40,79 
famoOtata.s cose... ois ae ee ote 272 perlongs.-\2<.. 62-26. op eee 393,398 
Achar 7 Jo. sc- eek a eee =o eee ae 436 | < pilosa Js2s6cs2 322 J3-2222 oe ee 138, 142 
PBA SE 32 a aka = ma ec 310, 311, 312,343 / pirnelioides= = ..-52:50 os ee 310, 311,313, 384 
cr Ree en a gn ae 156,172 | var: bruminipes .- ....-222.2a-5eee 313, 384, 386 
PON MIS. 5. anemic apa one Pear eae 412,416 planata. <2 <52..2.02255--22 40s soe eee 104 
Siganteas $2 os. . 2:2 SS oe ese see 411,412,496 Planipennis ....05-.. 5... eae 435,445 
WAT OSETIA. Sos ena ao a ees eee 412.421 POTCAIAS 22:0. sos foes Soe ue ee 38, 40, 63 
SBN ss oe Ae eee co oe eee 412;416. | ‘producta......20<. 1... 7-.22sp=ss eee 312, 362 
DOTY ES oece Cece ® ote oer eee 156, 164 PrOmMIMeNS - ..--.5 52.52—en ean 251,254 
PTACUIS oe es op Mca ee eee 235,237 PYGMIES oe Se ore esse nee = eee 38, 40,77, 418 
fay CIstans: <2 5.7 Ad2 2... a> eeugeet eames 235, 242 quadricollis. 2.3. jeans oes eee 38, 40,81 
STANGICOMISE.. =. eevee soc c ac ee eee 207 | 0ar..anthracina..... =... =... .ssseseeeee 38, 40,87 
MEAMOSA 1a cance set ent pean ew eee 147 hustrans- 52352 cee. Ep 38, 40,89 
Bra Ais sa cSas ar lek ceo e = Soe le eee 116,127 reflexicollis. .. 222s <.0<. <+2+55+5200s eee 450,494 
Hay deni <= sect see sie cans Soe + aerate anes 425 TIGYI=-~ foes oh eee oS ao 38, 41, 100 
DIES thie chaser os nee $6 eee 138,139 TobUStAls 203s 2- Se Se 104,106 
ISHsrisse=. s,- - asa3s enone sone 212,217 seabricula...... ee ‘a 435,440 
LOG 50) 0) 0 24 1 0 a aoe Spee ae Soe 310,311, 312,368 seabripennis <°-....- -.- S5ck= ee 312,338 
hornilea oe. coe eS eee ee 310, 311,312,350 BCADIOSe = fo Sos eae ow ee 310, 311,312,375 
uUMmMeras 7) oe a5. obs an ene eee 38, 41,95 var. rotundipennis.-.....°3..aceesuee 312, 384 
ANON. oe ak ie ee 45 SChwarvil. <=. 205. Jase ses ~ sees ¢ oeweee 393,406 
MI pRLENS Ss aoe6s oa en Ren eee 259, 263 SOMSta.. 2.22 50.5c0cnn he 4s 36s e ee 164 
Smpressicollis: ...<.Sae —..cteemn cee sas oer 371 | sinuata......----.--------.---------- +2222 266 
imoriitea 92 5c 5 eo. ee ena 310.311 8294) BSnOWil . < -....cc. se -eve~ eae ce aeee ones 310,311,317 
mriocens: =.= 8 ee se eee 5 Sci eee 412,429 BOVORS 4c 24. ond ote oche eee ee 38, 40, 45, 50 
STISUIATIS Oo tle Ans sees cee ee ee 156 | SPOMSO...4+- 2-02-22 ee eee es ceec er eeeeees 212,213 
intermedia. |... 225< 0. sosssanaeeeeeee aes 505,507 |< strieta.... 22. i0ce.5s0e.> 20d ee 379 
AN GOITUp ta. ek. ee te eee eee 72 strolatac..s<c>...5-se- 6st ee 156, 168 
BIE CATE aca. Se cei a te pean eames 379 SUbaSPCFrA: .....52-2 5 -<92-- 022 se be ee 127 
RTIANSI eae | ae. ox ae Geta wean ew kone 38, 40,67 SUDASPOTSs é ss au cease oa ee 321 
MAE ei Se ste ees dee itateiess ae 496 subcylindties. . ..25..5.2tccce-s~ eee 235, 246 
leeanibel- sce tote ses - ce taewa ten cee 217 subligats.-......s2- 2. 2. -c.<- 2c 384 
lengntel- seep. obs en ot soe eee 310, 311,321 Supnitens.....5. do. 202.22. Se 156,161 
Letcher 3222. 5 2S... 5. oe ee 116, 133 Subpingiis.<.....22.5--. . <=... - Soe 234,247 
var. vandy kel. ... Sere eenaebee eves shnees 136 BUMCAIG % . ocean con es 2 se one 217 
JON CICOTIS: =. 5 <s.ccuise cle ptters < teem orn cama 411,425 Siicipennis... 2.5.2.2 seca as tae 185, 187, 190, 194 
lenpipilosa.< 25 4:5 tence we ewe eae 212,230 Suthralis;.c.. 5... Ane sepeeeeee 185, 199, 203 
PURGERIIN 5 So Poe che on cee ee ater 38, 40,89 raf. texene.* 58S 185, 199, 202, 203 
minrpingta® 4:3 25 205-52 od caeeoccue ee 435,436 typita....220/ dei aoe 202 
OLE PATIS oo sin ctu errins ote wc ane wee 234, 267 tarsslis 4... 2. Sacccescws ts sl neem 81 
Par MGMOLsthe. «uaa (feo. ca Mes cee 234,272 FENG WHOA. 2. Sages «ts sae wand Se 310,311,325 
WIBOTNORICATBo os < 2. cents an seen neeete 38, 41,113 Oar nana ..- 5: so <5. ee ee 311,328 
MOOLOINGG sos waweseces tossed 310, 311,312, 347 PANU PSS 255-5 Sos cease sneer 235, 294 


R % INDEX. 523 
3 Page. I. 

Bones TOXONA ............ ce ceene 185, 199, 202, 203 Page. 
a I OR eae 310,311,313 ramus, HUGS os a sion ae cow cevinevine 45 
RR C2 Pyrite ae $8, 41,104 | impotens, Eleodes..................++. 234, 259,263 
ETE Sto 8.48. Sh ld - 379 | impressicollis, Eleodes...................see- 371 
i Mca agecks wh Jonteget antsy 207,208 | inculta, Fleodes............-.+--se00+: 310,311,329 
tien si fCak pena date 116,186 | innocens, Eleodes ..............20sssce0es 412,429 
oS 935,300 | insularis, Eleodes............0..0ssccceeeeees 156 

A ap a ee 235, 305 intermedia, Eleodes......:.........-..e+-- $05,607 
ke, Sa eee 933 | Imterrupta, Eleodes.........22..ssccccccoost 72 
I ere ee oe sa 179 mrereoete, WlegGeics 2 sot cackcdctbacecnadcve. 379 
a a RIES Ee EE 5 ke a eree ee 384 K. 
ns gapggg | veal Eleoden nnn enescepcecnee 98, 40,63 

RE Bina web nog caine cpeawe 27,30, 495 Ls 
_ Eleodimorpha...............-..-+-+++++- 29, 30,477 | jaminatum, Embaphion.................. 467,472 
bolean . ....---..- 2+ essere renee Ore h Dares, MUNOMOR.. 5 nals .ccecsuateussceceus 496 
) Glongata, Eleodes............-......-.---. 505, 506 ORIN 3s vs as-cast 496, 497 
_ elongatum, Embaphion................... 453, 454 a CAE EEE Peer 496 
acnteenceecnescereneresecees 29, 30,450 | jaticollis, Eleodes................. 234,276,280, 505 
etna eteese<csenncescasaeneccoeces SIS | ecsmaitel, MleONG. 352), oo2.5. liz. sn ecdenes enon = SAT 
teste eeseeeceeeeseeeeseeeeees 453,460 | jecontei, Eleodes...................... 310,311,821 
PASE eS he TSR Re hase wld oes 453,467 MOUGHIGRA; SOMBOGER S 2c cacsnccccaes s+ osssuceet kia eee 
Rance earencceeereneveres 407,472 | “Litheleodes.....'.........002.<02++s---+- 94 35,04 
Bsr en own ses +onveenereananeasnene 453 | longicollis, Eleodes........................ 411.4% 
sees estes eec eens ee ereeeeeees 453,454 | jongipilosa, Eleodes.....................-. 212, 
oats teense ee ec eee e rene rece eens 453,457 | luca, Eleodes.....................-... 235,285,289 
tee c nett ener eect eee neers 453,473 | justrans, Eleodes................-2..-.---.-. 89 
ek. oS A ree ee 453, 464 
tt Te SRS 235, 285 M. 
LAS > ee ee 412,421 | marginata, Discogenia....................... 486 
SE FRE PO a 121 VT Sn ame | 
‘et AST Se ae oe Seer 116,121, |~Melaneleodes~..: 5.2... 2-2 .-5..52-2.~ 8B; 84,35, 86 
WOMB eons ane nbs ws gn uce meade 34, 35,391 
SURES, NOMS 5 So ke wea ee one 234, 267 
wtttncceccscccseccence 235,305 TAUTORRUR, TADS... ce... cis.ctcsuc~wceseas ODS 
weet ete e eet eeeeeeeeeeeee 202) Wsntrionts (8) Akis.c% <.255.55 oop cod ae MGB 
tone e teen eee ee eee eeeeee eee eee 436 | muricatum, Embaphion.................. 453,478 
310, 311, 312, 343 
eN LUR eect a 156, 172 N. 
TIME, TORN S Oa carats dara aim in eae 311,328 
neomexicana Eleodes................... 38, 41,113 
sence tee ee ee teeeeeeeeeeeeee 412,416 | neotomm, Eleodes................ 310,311, 312,347 
wn ee css eee een eeeeeeee 411,412,496 | nevadensis, Eleodes....................... 393,402 
Ake iisoernadeaswesae 453,407 beltog ir OBS Ss 2 ee ee re yee 
woe ee eee nee e tenet eee ee eens 156,164 | nitida, Eleodes.......................-..-- 450,498 
cect ese tate eaten eee eee e eee 235,237 | nupta, Eleodes....................... 217,220,221 
whe cil shat hada ae ti 207 RENO rand os ope a chile: Wein +n etaes steele 3 
(Se Ee ae ae M47 
OE Se I es 116,127 0. 
BERETOUND BL, DISCOPEINA,. . oc... oo ccccsecsacees SIRE ODMOTER PENN nn a cae hiine ne na esas turk ivetdes 187 
ee as ee 434 MIE 3 so Sapas as paint os 185, 187, 190, 193 
eden aie.2a RES ale 41 RN Ga hc cha cawaies ies ae cuCie nt” 100 
Lushan diaswdicaie wires URS NN ERIN ae tian wren dcuvaavevacccasdy 58 
Sats dw okt Wie eer kc ephale im iitndecn Doe atk jo eee 
die Ee 184,205 | obtusa, Eleodes....................... 127,180,131 
Se ee ee ON eS ee 186 |) Oman, MONON s yo e soc. c oneness ss... 340,98 
Po | a a Pe ere eee 156,175 
SS ee en ae 156, 175 
bcbg aw itn adhe pktah pas ane eee 425 
Miticn dsih, Ohad eobaaker see 33,35, 179 P. 
is Ue ae RAN WRN RH Eee aeeeer eh 138,189 | parvicollis, Eleodes ............... 310,311,312, 354 
bk wed Gu LOR melas 212,217 | pedinoides, Eleodes..................... 38,41,109 
310,311,312,368 | peninsularis, Eleodes..................... 38, 40,79 
310,311,312,350 ) ORI IGOGOD Ss cute dw and <sdacnscccusecs 393,398 
OOS SESS 52° TEL Oa ee ene 
LV ncn nals nestitditetekseieibes ee ee ee | 


524 INDEX. 
Page. Page. 
pimelioides, Eleodes..........----- 310,311,313,384 | subcylindrica, Eleod@S...............-.0+- 235, 246 
pilosa, Eleodess<2322-== 45. 5:hssdoacceewaee 138,142 | subligata, Eleodes. ....2..-..2-<<--seneea osse “2884 
planata, Eleodes...........--.---- 104, 312,355,359 | subnitens, Eleodes......----......-----.-- 156,161 
planipennis, Eleodes..............-....-- 435,445 | subpinguis, Eleodes................-...--- 234,247 
planum;-Hmibaphion.-.....-.-~-- aeeeeeees 453,464 | sulcata, Eleodes.............----cse--e--e0ee 217 
porcata, Wleodes...25.3535.522-222soe 38,40,58,63 | sulcipennis, Eleodes..............- 185, 187, 190, 194 
producta, Eleodes..........-...---.2.- 312,355,362 | suturalis, Blaps.....-........25-.seeeeeeeee 199 
prominens, Eleodes-.......-..2-5+.---s.-- 251,254 Eleodés:....:.=-- 2. ose 185, 199, 202 
PrOUIhes 3. be a eess- «wel os cede 34,35, 152 T 
Paandeleodes.. jai 2~-:.2: 02.2.0 34,35, 146 | ; 
Pupa, Eleodes clavicornis. ...........-..--- 500 | Tagoma.....-.-+-.-+-+-+-+-0e0e eee ee sees eres 3 
pygmeea, Eleodes............-------- 38,40,77,418 | tarsalis, Eleodes.......-.-----------+----+++- 81,85 
a pace St i aie on utes eee 310,311,325 
F tenuipes; Eleodes. ........-s¢s>esse==eeeeee 235, 294 
quadricollis, Eleodes: ..........-+-.--2-5- 38,40,81 | texana, Eleodes...... ee yo? 185, 199, 202, 203 
R. | tibialis, Eleodes. .....:.=. 5+: ===sseeee 310, 311,313 
Tricheleodes: .... -.2...<=.-»ene= eee 34, 35,138 
reflexicollis, Eleodes sous vinnie anna a an ae oS 450,494 | tricostata, Blaps ncecdwe abner 104 
THEY1, Eleodes. 52225 Senet ane eles onsen 38,41,100 | Eileodés........ 2 38, 41, 104 
robusta, PAGQGES 65 ochp cad -porseeaceeeess 104, 106 Trogloderus i ae © *. ecw ecnece toe 29 483 
rotundipennis, Eleodes............--.- 312,379,384 COStatUS ...<c ac. men ee 486 
s. tuberculatus. 2. - 2.06 s.0< neater 486, 490 
; y - |. tuberculata, Eleodes. - --- .-.23)ceag ee 37S 
seabricula, a. oe ae eae 4 se ee tuberculatus, Trogloderus.............-.-- 486,490 
scabripennis, Eleodes..........-.-.--.- 312,313,338 | Vv. 
scabrosa, Eleodes.........--------- 310,311,312,375 | yandykei, Eleodes.............2-.-------- 116,136 
zig zetia ee ree 393,406 | ventricosa, Eleodes-... <<. 2202. eee 235,300 
Sariath, HlG0deS< 3.5. s<coae pete cccaen eons 164 i 
BR, lope... 08d a anced bana 004 | Vetelator, Bloodes.... 
snowii, Eleodes.....----..-.----------- 310,311,817 | viator, Eleodes..........--------------+-e--- 384 
soror, Eleodes.....---------+--++++++-- 38,40,45,50 | vicina, Eleodes............-.----ceeeeeeeee 450,493 
Sponss. PlGOdeS. 2<- -~ S15 ac. nn nano eee 212,213 
Phentlnaden oc <5. ake 33, 34, 35,409 W. 
stricta, Eleodes..........------.-----2---<--- 379 | wickhami, Eleodes...........------------ 235,297 
Striolata, .wieodes;. 122... = 22s. eke eee 156, 168 
subaspersd,|(2) Mlepdes. 2-5 =. saceee- sees 127 x. 
subaspera, mleodes.... «2 ssos55 sce. eee 321: | “Kysta......-2 5... - 0... --s-2e 


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